The long-billed corella typically digs for roots, seeds, corms, and bulbs, especially from the weed onion grass. Native plants eaten include murnong Microseris lanceolata, but a substantial portion of the bird's diet now includes introduced plants.


The yellow-rumped cacique (Cacicus cela) is a passerine bird in the New World family Icteridae. It breeds in much of northern South America from Panama and Trinidad south to Peru, Bolivia and central Brazil. However, they have been sighted as far north as Nayarit state in Mexico.


The male is on average 28 centimetres (11 in) long and weighs about 104 grams (3.7 oz), with the female 23 centimetres (9.1 in) long and weighing approximately 60 grams (2.1 oz). The yellow-rumped cacique is a slim bird, with a long tail, blue eyes, and a pale yellow pointed bill. It has mainly black plumage, apart from a bright yellow rump, tail base, lower belly and wing "epaulets". The female is duller black than the male, and the juvenile bird resembles the female, but has dark eyes and a brown bill base.
The song of the male yellow-rumped cacique is a brilliant mixture of fluting notes with cackles, wheezes and sometimes mimicry.  There are also many varied calls, and an active colony can be heard from a considerable distance.


The fan-tailed cuckoo (Cacomantis flabelliformis) is a species of cuckoo in the family Cuculidae.
It is found in Australia, Fiji, New Caledonia, New Zealand, New Guinea, Solomon Islands, and Vanuatu.


Six subspecies have been recognised:
Cacomantis flabelliformis excitus (Rothschild & Hartert, 1907) – Mountains of New Guinea.
Cacomantis flabelliformis flabelliformis (Latham, 1802) – Eastern Australia from Cape York through to southern South Australia; south-western Western Australia.
Cacomantis flabelliformis meeki (Rothschild & Hartert, 1902) – Solomon Islands including Santa Isabel, Kolombangara, Guadalcanal and Bellona. Originally designated Cacomantis meeki.
Cacomantis flabelliformis pyrrhophanus (Vieillot, 1817) – New Caledonia including the Loyalty Islands.
Cacomantis flabelliformis schistaceigularis (Sharpe, 1900) – Vanuatu.
Cacomantis flabelliformis simus (Peale, 1848) – Fiji.


The fan-tailed cuckoo is about 25–27 cm (9.8–10.6 in) long. The fan-tailed cuckoo has a slate-grey head, back and wings, rufous underparts and barred black and white tail. Its eye is surrounded by a yellow orbital eye ring which helps to distinguish it from the smaller and paler brush cuckoo (C. variolosus) and the also smaller chestnut-breasted cuckoo (C. castaneiventris).


Caenurgina erechtea, the forage looper or common grass moth, is a moth of the family Erebidae. The species was first described by Pieter Cramer in 1780. It is found from coast to coast in the United States and adjacent parts of Canada. It is not found in Newfoundland, New Brunswick, Prince Edward Island, Yukon, or the Northwest Territories.
The wingspan is 30–42 mm. Adults are on wing from March to November depending on the location.
The larvae feed on Ambrosia trifida and various species of clover, grass, and alfalfa.


The Muscovy duck (Cairina moschata) is a large duck native to the Americas, from the Rio Grande Valley of Texas and Mexico south to Argentina and Uruguay. Small wild and feral breeding populations have established themselves in the United States, particularly in Florida, Louisiana, Massachusetts, the Big Island of Hawaii, as well as in many other parts of North America, including southern Canada. Feral Muscovy ducks are found in New Zealand, Australia, and in parts of Europe.
It is a large duck, with the males about 76 cm (30 in) long, and weighing up to 7 kg (15 lb). Females are considerably smaller, and only grow to 3 kg (6.6 lb), roughly half the males' size. The bird is predominantly black and white, with the back feathers being iridescent and glossy in males, while the females are more drab. The amount of white on the neck and head is variable, as well as the bill, which can be yellow, pink, black, or any mixture of these colors. It may have white patches or bars on the wings, which become more noticeable during flight. Both sexes have pink or red wattles around the bill, those of the male being larger and more brightly colored.
Although the Muscovy duck is a tropical bird, it adapts well to cooler climates, thriving in weather as cold as −12 °C (10 °F) and able to survive even colder conditions. In general, Barbary duck is the term used for C. moschata in a culinary context.
The domestic subspecies, Cairina moschata domestica, is commonly known in Spanish as the pato criollo. They have been bred since pre-Columbian times by Native Americans and are heavier and less able to fly long distances than the wild subspecies. Their plumage color is also more variable. Other names for the domestic breed in Spanish are pato casero ("backyard duck") and pato mudo ("mute duck").


All Muscovy ducks have long claws on their feet and a wide, flat tail. In the domestic drake (male), length is about 86 cm (34 in) and weight is 4.6–6.8 kg (10–15 lb), while the domestic hen (female) is much smaller, at 64 cm (25 in) in length and 2.7–3.6 kg (6.0–7.9 lb) in weight. Large domesticated males often weigh up to 8 kg (18 lb), and large domesticated females up to 5 kg (11 lb).
The true wild Muscovy duck, from which all domestic Muscovies originated, is blackish, with large white wing patches. Length can range from 66 to 84 cm (26 to 33 in), wingspan from 137 to 152 cm (54 to 60 in) and weight from 1.1 to 4.1 kg (2.4 to 9.0 lb). On the head, the wild male has a short crest on the nape. The bill is black with a speckling of pale pink. A blackish or dark red knob can be seen at the bill base, which is similar in colour to the bare skin of the face. The eyes are yellowish-brown. The legs and webbed feet are blackish. The wild female is similar in plumage, but much smaller, with a feathered face and lacking the prominent knob. The juvenile is duller overall, with little or no white on the upperwing.
Domesticated birds may look similar; most are dark brown or black mixed with white, particularly on the head. Other colors, such as lavender or all-white, are also seen. Both sexes have a nude black-and-red or all-red face; the drake also has pronounced caruncles at the base of the bill and a low erectile crest of feathers. C. moschata ducklings are mostly yellow with buff-brown markings on the tail and wings. For a while after hatching, juveniles lack the distinctive wattles associated with adult individuals, and resemble the offspring of various other ducks, such as mallards. Some domesticated ducklings have a dark head and blue eyes, others a light brown crown and dark markings on their nape. They are agile and speedy precocial birds.
The drake has a low breathy call, and the hen a quiet trilling coo.
The karyotype of the Muscovy duck is 2n=80, consisting of three pairs of macrochromosomes, 36 pairs of microchromosomes, and a pair of sex chromosomes. The two largest macrochromosome pairs are submetacentric, while all other chromosomes are acrocentric or probably telocentric for the smallest microchromosomes. The submetacentric chromosomes and the Z (female) chromosome show rather little constitutive heterochromatin (C bands), while the W chromosomes are at least two-thirds heterochromatin.
Male Muscovy ducks have helical penises that become erect to 19 cm (7 in) in 0.3 s. Females have vaginas that coil in the opposite direction that appear to have evolved to limit forced copulation by males.


This non-migratory species normally inhabits forested swamps, lakes, streams and nearby grassland and farm crops, and often roosts in trees at night. The Muscovy duck's diet consists of plant material (such as the roots, stems, leaves, and seeds of aquatic plants and grasses, as well as terrestrial plants, including agricultural crops) obtained by grazing or dabbling in shallow water, and small fish, amphibians, reptiles, crustaceans, insects, millipedes, and worms. This is an aggressive duck; males often fight over food, territory or mates. The females fight with each other less often. Some adults will peck at the ducklings if they are eating at the same food source.
The Muscovy duck has benefited from nest boxes in Mexico, but is somewhat uncommon in much of the eastern part of its range due to excessive hunting. It is not considered a globally threatened species by the IUCN, however, as it is widely distributed.


This species, like the mallard, does not form stable pairs. They will mate on land or in water (note the submerged female in the image below). Domestic Muscovy ducks can breed up to three times each year.
The hen lays a clutch of 8–16 white eggs, usually in a tree hole or hollow, which are incubated for 35 days. The sitting hen will leave the nest once a day from 20 minutes to one and a half hours, and will then defecate, drink water, eat and sometimes bathe. Once the eggs begin to hatch, it may take 24 hours for all the chicks to break through their shells. When feral chicks are born, they usually stay with their mother for about 10–12 weeks. Their bodies cannot produce all the heat they need, especially in temperate regions, so they will stay close to the mother, especially at night.
Often, the drake will stay in close contact with the brood for several weeks. The male will walk with the young during their normal travels in search for food, providing protection. Anecdotal evidence from East Anglia, U.K. suggests that, in response to different environmental conditions, other adults assist in protecting chicks and providing warmth at night. It has been suggested that this is in response to local efforts to cull the eggs, which has led to an atypical distribution of males and females, as well as young and mature birds.
For the first few weeks of their lives, Muscovy chicks feed on grains, corn, grass, insects, and almost anything that moves. Their mother instructs them at an early age how to feed.


Feral Muscovy ducks can breed near urban and suburban lakes and on farms, nesting in tree cavities or on the ground, under shrubs in yards, on apartment balconies, or under roof overhangs. Some feral populations, such as that in southern Florida, have a reputation of becoming pests on occasion. At night they often sleep at water, if there is a water source available, to flee quickly from predators if awakened. Small populations of Muscovy ducks can also be found in Ely, Cambridgeshire, Calstock, Cornwall, and Lincoln, Lincolnshire, U.K. Muscovy ducks have also been spotted in the Walsall Arboretum. There has been a small population in the Pavilion Gardens public park in Buxton, Derbyshire for many years.
In the U.S., Muscovy ducks are considered an invasive species. An owner may raise them for food production only (not for hunting). Similarly, if the ducks have no owner, 50CFR Part 21 (Migratory Bird Permits) allows the removal or destruction of the ducks, their eggs and their nests anywhere in the United States outside of Hidalgo, Starr and Zapata Counties in Texas, where they are considered indigenous. The population in southern Florida is considered, with numbers in the several thousands, to be established enough to be considered "countable" for bird watchers.
Legal methods to restrict breeding include not feeding these ducks, deterring them with noise or chasing them away.
Although legislation passed in the U.S. prohibiting trade of Muscovy ducks, Fish and Wildlife Services intend to revise the regulations. They are not currently implementing them, though release of Muscovy ducks to the wild outside their natural range is prohibited.


Cakile edentula  is most commonly found on the beaches and dunes of North America. More commonly, it is found on the East coast of the United States. American sea-rocket is most likely to be found in areas along the coastline. This plant is not a wetland plant, but can occasionally be found in wetland environments. It is also a non-native, invasive species in other parts of the world, especially Japan and Australia.


Flowers are separate, radially symmetrical, and contain four petals. These flowers can be pink to red, blue to purple, or white. Fruit are small and green, about 1–2 centimeters in diameter. These fruit are dry and do not split open when ripe. Fruit are released at the end of each growing season.


Caladenia carnea was first formally described by Robert Brown in 1810 and the description was published in Prodromus Florae Novae Hollandiae. In 2001, David Jones and Mark Clements proposed changing the name to Petalochilus carneus but the change has not been widely accepted.


Calamagrostis epigejos has a broad distribution in temperate Eurasia, from France and Great Britain to Japan. A distinct variety is found in southern and eastern Africa.


The lark bunting is the most prevalent of the passerine species found in the grasslands of North America. Their breeding habitat is prairie regions in central Canada and the mid-western United States. These birds migrate in flocks to winter southern Texas, Arizona and the high plateau of northern Mexico in the fall.


The Lapland longspur is a robust bird, with a thick yellow seed-eater's bill. The summer male has a black head and throat, white eyestripe, chestnut nape, white underparts, and a heavily streaked black-grey back. Other plumages have a plainer orange-brown head, a browner back and chestnut nape and wing panels.
Measurements:
Length: 5.9-6.3 in (15-16 cm)
Weight: 0.8-1.2 oz (22.3-33.1 g)
Wingspan: 8.7-11.4 in (22-29 cm)


The food habits of the Lapland longspur are quite simple: mostly seeds in winter and arthropods in the summer, when they are in activity.
During the winter, the longspur feeds on seeds. They pick them on the ground, rarely feeding directly on plants. They will forage around the same area for a period varying between a few minutes and an hour, then fly away looking for a new foraging area. Their seed diet is composed mainly of seeds from grass, foxtail, cultivated millet, crabgrass and wheat. During the breeding season, the birds migrate to the north, where their diet switches to arthropods. Nestlings are only fed arthropods, which also constitute the diet of the parents at that time of the year (June to July). The birds often catch insects in mid-air, but do forage through vegetation when climatic conditions prevent the insects from flying. Longspurs can consume between 3000 and 10,000 prey items (insects or seeds) per day, depending on their energy needs. Dipteran larvae and adults form the major part of their insectivorous diet.


The sanderling is a small plump sandpiper, 18–20 cm (7.1–7.9 in) in length. Its weight ranges from 40–100 g (1.4–3.5 oz). The winter bird is very pale, almost white apart from a dark shoulder patch. This is the source of the specific name, alba, which is the Latin for "white". Later in the summer, the face and throat become brick-red. The juvenile bird is spangled black and white, and shows much more contrast than the adult.
If its size is misjudged, a sanderling in breeding plumage can be mistaken for some varieties of stint, or a sanderling in winter plumage can be mistaken for a dunlin or red knot. It can be told from other small wading birds, given good views, by its lack of a hind toe. Its behavior is also distinctive.


The sanderling breeds in the High Arctic areas of North America, Europe and Asia. In North America, it breeds in the Canadian Arctic Archipelago, Nunavut, Greenland (and to a lesser extent Alaska). In Eurasia, it breeds in Spitsbergen and areas of northern Russia from the Taymyr Peninsula to the New Siberian Islands. In the northern winter, it has a nearly cosmopolitan distribution across the world's marine coasts. It is a complete migrant, travelling between 3,000 to 10,000 km (1,900 to 6,200 mi) from its breeding grounds to its wintering sites. Birds that travel further also arrive later and leave sooner. Most adults leave the breeding grounds in July and early August, whereas juvenile birds leave in late August and early September. The northward migration begins in March at the southern end of their winter distribution.
The breeding habitat of the sanderling is coastal tundra north of 5 °C (41 °F) July isotherm. The species typically chooses nesting sites on dry stony areas near wet areas, from 60 m (200 ft) above sea level to 800 m (2,600 ft). During the winter and its migration, it is most commonly found on coastal sandy beaches, but also occurs on tidal sand flats, mud flats and the shores of lakes and rivers. More infrequently, it may occur on rocky shores.


Sanderlings feed on invertebrate prey buried in the sand in the upper intertidal zone. In North America, this diet largely consists of the isopods Excirolana linguifrons, Excirolana kincaidii, and the mole crab, Emerita analoga. When the tide is out, these crustaceans live in burrows some way beneath the surface. When the tide comes in, they move into the upper layers of sand and feed on the plankton and detritus that washes over them with each wave. They then burrow rapidly down again as the water retreats. They leave no marks on the surface, so the sanderlings hunt for them by plunging their beaks into the sand at random, consuming whatever they find. Their bills can penetrate only 2 or 3 cm (0.79 or 1.18 in) and as the water swirls around and retreats, the sand is softer; this makes it easier for the birds' beaks to penetrate further. In the spring, when much breeding activity is taking place in the benthic community, there may be as many as 4000 invertebrates per square metre, but their average size is smaller than later in the year. The birds appear to rush madly around at the edge of the surf, but in reality they are maximising their chances of catching as many prey animals as possible when they are at their most vulnerable near the surface.


The dunlin (Calidris alpina) is a small wader, formerly sometimes separated with the other "stints" in the genus Erolia. The English name is a dialect form of "dunling", first recorded in 1531–1532. It derives from dun, "dull brown", with the suffix -ling, meaning a person or thing with the given quality.
It is a circumpolar breeder in Arctic or subarctic regions. Birds that breed in northern Europe and Asia are long-distance migrants, wintering south to Africa, southeast Asia and the Middle East. Birds that breed in Alaska and the Canadian Arctic migrate short distances to the Pacific and Atlantic coasts of North America, although those nesting in northern Alaska overwinter in Asia. Many dunlins winter along the Iberian south coast.


Dunlin are small migratory waders, however they show strong philopatry with individuals of the Southern Dunlin (Calidris alpina schinzii) in Sweden and Finland returning to, or very close to, their natal patches. Habitat fragmentation has reduced the availability of habitat patches to these birds through reducing patch size and increasing patch isolation. This reduced connectivity between patches has reduced the movements of Dunlin leaving them more susceptible to inbreeding in these locations. Future management for the conservation of Southern Dunlin should include increasing the connectivity between habitat patches.


The dunlin is highly gregarious in winter, sometimes forming large flocks on coastal mudflats or sandy beaches. Large numbers can often be seen swirling in synchronized flight on stop-overs during migration or on their winter habitat.
This bird is one of the most common waders throughout its breeding and wintering ranges, and it is the species with which other waders tend to be compared. At 17–21 cm (6.7–8.3 in) length and with a 32–36 cm (13–14 in) wingspan, it is similar in size to a common starling, but stouter, with a thicker bill.
The dunlin moves along the coastal mudflat beaches it prefers with a characteristic "sewing machine" feeding action, methodically picking small food items. Insects form the main part of the dunlin's diet on the nesting grounds; it eats molluscs, worms and crustaceans in coastal areas.


The red knot (Calidris canutus) (just knot in English-speaking Europe) is a medium-sized shorebird which breeds in tundra and the Arctic Cordillera in the far north of Canada, Europe, and Russia. It is a large member of the Calidris sandpipers, second only to the great knot. Six subspecies are recognised.
Their diet varies according to season; arthropods and larvae are the preferred food items at the breeding grounds, while various hard-shelled molluscs are consumed at other feeding sites at other times. North American breeders migrate to coastal areas in Europe and South America, while the Eurasian populations winter in Africa, Papua New Guinea, Australia, and New Zealand. This species forms enormous flocks when not breeding.


In the breeding season, the red knot has a circumpolar distribution in the high Arctic, then migrates to coasts around the world from 50° N to 58° S. The red knot has one of the longest migrations of any bird. Every year it travels more than 9,000 mi (14,000 km) from the Arctic to the southern tip of South America and repeats the trip in reverse. The exact migration routes and wintering grounds of individual subspecies are still somewhat uncertain. The nominate race C. c. canutus breeds in the Taymyr Peninsula and possibly Yakutia and migrates to Western Europe and then down to western and southern Africa. C. c. rogersi breeds in the Chukchi Peninsula in eastern Siberia, and winters in eastern Australia and New Zealand. Small and declining numbers of rogersi (but possibly of the later described piersmai) winter in the mudflats in the Gulf of Mannar and on the eastern coast of India. The recently split race C. c. piersmai breeds in the New Siberian Islands and winters in north-western Australia. C. c. roselaari breeds in Wrangel Island in Siberia and north-western Alaska, and it apparently winters in Florida, Panama and Venezuela. C. c. rufa breeds in the Canadian low Arctic, and winters South America, and C. c. islandica breeds in the Canadian high Arctic as well as Greenland, and winters in Western Europe.
Birds wintering in west Africa were found to restrict their daily foraging to a range of just 2–16 km² (0.77–6.18 sq mi) of intertidal area and roosted a single site for several months. In temperate regions such as the Wadden Sea they have been found to change roost sites each week and their feeding range may be as much as 800 km² (310 sq mi) during the course of a week.
B95, also known as Moonbird, is a noted individual of the subspecies C. c. rufa. A male, he has become famous amongst conservationists for his extreme longevity — he was aged at least 20 as of his last sighting in May 2014.


An adult red knot is the second largest Calidris sandpiper, measuring 23–26 cm (9.1–10.2 in) long with a 47–53 cm (19–21 in) wingspan. The body shape is typical for the genus, with a small head and eyes, a short neck and a slightly tapering bill that is no longer than its head. It has short dark legs and a medium thin dark bill. The winter, or basic, plumage becomes uniformly pale grey, and is similar between the sexes. The alternate, or breeding, plumage is mottled grey on top with a cinnamon face, throat and breast and light-coloured rear belly. The alternate plumage of females is similar to that of the male except it is slightly lighter and the eye-line is less distinct. Canutus, islandica and piersmai are the "darker" subspecies. Subspecies rogersi has a lighter belly than either roselaari or piersmai, and rufa is the lightest in overall plumage. The transition from alternate to basic plumages begins at the breeding site but is most pronounced during the southwards migration. The molt to alternate plumage begins just prior to the northwards migration to the breeding grounds, but is mostly during the migration period.
The large size, white wing bar and grey rump and tail make it easy to identify in flight. When feeding the short dark green legs give it a characteristic 'low-slung' appearance. When foraging singly, they rarely call, but when flying in a flock they make a low monosyllabic knutt and when migrating they utter a disyllabic knuup-knuup. They breed in the moist tundra during June to August. The display song of the male is a fluty poor-me. The display includes circling high with quivering wing beats and tumbling to the ground with the wings held upward. Both sexes incubate the eggs, but the female leaves parental care to the male once the eggs have hatched.
Juvenile birds have distinctive submarginal lines and brown coverts during the first year. In the breeding season the males can be separated with difficulty (<80% accuracy in comparison to molecular methods) based on the more even shade of the red underparts that extend towards the rear of the belly.
The weight varies with subspecies, but ranges between 100 and 200 g (3.5 and 7.1 oz). Red knots can double their weight prior to migration. Like many migratory birds they also reduce the size of their digestive organs prior to migration. The extent of the atrophy is not as pronounced as species like the bar-tailed godwit, probably because there are more opportunities to feed during migration for the red knot. Red knots are also able to change the size of their digestive organs seasonally. The size of the gizzard increases in thickness when feeding on harder foods on the wintering ground and decreases in size while feeding on softer foods in the breeding grounds. These changes can be very rapid, occurring in as little as six days.


As one of  the “longest distance migrants in the animal kingdom,” the red knot relies heavily on the same stopping sites each year along their migratory routes to refuel their bodies in order to complete their migrations to and from breeding sites. Red knots travel “in larger flocks than do most shorebirds" flying “9300 miles from south to north every [northern hemisphere] spring and repeat the trip in reverse every autumn”. Northern hemisphere winters are spent in Tierra del Fuego South America and migration routes lead to breeding locations on islands and mainland above the Arctic circle during the short arctic summer. These long expeditions are broken into various segments about 1500 miles each ending at staging areas that are visited yearly. Specifically, the Delaware Bay is the most vital migratory rest stop for the red knot, as much of their physiological demands are met by consuming the abundance of horseshoe crab eggs as their main food source during migration. The relationship between red knot and horseshoe crab is evolutionarily intertwined as “their arrival coincides with the annual horseshoe crab spawning in the Delaware Bay”. Data supports the hypothesis that abundance of horseshoe crab eggs on beaches such as in the Delaware Bay drive movement and distribution of red knots, and the number of horseshoe crabs in the Delaware Bay characterize its importance/ relevance to their migration route. Other stopover spots in the US include islands off the coast of Massachusetts, Virginia, South Carolina, and Georgia. Fewer red knots undergo overland migration routes and winter on the Gulf Coast. Stopover areas on this route are found in the Mississippi river drainage, Northern U.S. saline lakes, and plains in Southern Canada.


Red knots undergo various physiological changes before their migration to account for the physical demand of the long expedition: “flight muscle mass increases, while leg muscle mass decreases. Stomach and gizzard masses decrease, while fat mass increases by more than 50 percent”. They arrive at stopover sites extremely thin. Since the gizzard is shrunken for their travel, fewer hard foods are eaten and instead soft and nutritious horseshoe crab eggs are the desired food source. As the migration is timed with the release of eggs, they are widely available in these specific stop over locations, making the resource easy to locate and digest, saving the birds energy. Thus, the abundance and accessibility of horseshoe crab eggs at these specific locations justify their intertwined relationship. Body weight may be up to doubled during stopover stays lasting around 10–14 days from constant consumption of food to increase their body fat enough to continue the remainder of their trip. The abundant horseshoe crab population inhabiting the Delaware bay deems it the most important stopover habitat in the red knots migration “supporting an estimated 50 to 80 percent of all migrating rufa red knots each year”.
Due to the interrelatedness between horseshoe crab egg abundance and red knot viability, the health of the horseshoe crab population is increasingly relevant in the discussion of red knot population fluctuation and success. Horseshoe crabs were harvested for fertilizer and to feed animals in the early 20th century, and currently are harvested for bait usage by U.S. fishing companies. Horseshoe crab  populations took a toll, and red knot population “numbers in Tierra del Fuego (winter) and Delaware bay (spring) declined about 75 percent from 1980s to 2000s”. Over harvesting of the crabs in the 1990s provoked the action for population management by the Atlantic States Marine Fisheries Commission to establish “mandatory state-by state harvest quotas and [create] the 1,500-square-mile Carl N. Shuster Jr. Horseshoe Crab Sanctuary off the mouth of Delaware bay.”  Subsequently, commercial horseshoe crab bait use has been reduced by population management and innovative bait conservation techniques, and a correlated stabilization in knot populations has been recorded as well.


Red knot populations are greatly affected by climate change since middle and high arctic habitats are necessary for breeding. Higher latitudes and coastal areas where red knots breed and winter are most affected by climate change. Nesting sites are generally on open ground in the tundra near water. Male knots prepare 3-5 sites for nests on “normally dry, stony areas of tundra in upland areas, often near ridges and not far from wetlands”. Thus, sea level rise, coastal erosion, and general warming temperatures due to global warming destroy the ideal breeding habitats for the red knot in these arctic coastal regions.
Migratory animals by nature have adapted to various environments. Subsequently, rapid and complex responses to climate change are prominent and in the case of the red knot in the recent changing climate, evolutionarily damaging. As arctic breeding grounds continue to warm, red knot body size has decreased, and less success for survival of birds born in warmer years is reported. Even more significantly, their wintering areas in the tropics have become more stabilized, resulting in shorter bill birds (likely due to the fact that stable conditions breed greater ecological success and less variability within species). Consequently, knots struggle to reach their main food sources which include deeply buried mollusks, adding to lesser access to food as well as more expenditure of energy in attempting to attain food.
Thus, indirect threats including horseshoe crab over harvesting and climate change greatly threaten red knot populations. Issues such as climate change may be harder to immediately/directly alter, yet limiting horseshoe crab harvest and human disturbance to protect knot food sources and migratory/breeding sites are effective methods of conservation for the red knot.


In 2003, scientists projected that at its current rate of decline the American subspecies, rufa, might become extinct as early as 2010, but as of April 2011 the subspecies is still extant.  In New Jersey, state and local agencies are taking steps to protect these birds by limiting horseshoe crab harvesting and restricting beach access. In Delaware, a two-year ban on the harvesting of horseshoe crabs was enacted but struck down by a judge who cited insufficient evidence to justify the potential disruption to the fishing industry but a male-only harvest has been in place in recent years.
Late in the fall of 2014, the red knot rufa was listed as a federally threatened species under the United States Endangered Species Act – the second most critical status that can be awarded to a subspecies. This followed a decade of intensive petitioning by environmental groups and a lawsuit against the Department of the Interior for alleged negligence in the protection of endangered species through failure to evaluate and list them. The reasons for the red knot rufa's listing were varied; habitat degradation, loss of key food supplies, and threats posed by climate change and sea level rise were all listed as factors that were considered when the red knot rufa was listed.


The white-rumped sandpiper (Calidris fuscicollis) is a small shorebird that breeds in the northern tundra of Canada and Alaska. This bird can be difficult to distinguish from other similar tiny shorebirds; these are known collectively as "peeps" or "stints".
These birds are not often spotted. In the summer, they are rarely seen because they are in such an obscure breeding location. Similarly, in the winter they are rarely seen because they travel too far south for many birdwatchers. Therefore, the majority of sightings occur during the spring or fall in temperate regions and are generally in small numbers around water.


The white-rumped sandpiper is placed in the order Charadriiformes along with gulls, alcids, plovers and oystercatchers. Its family, Scolopacidae, encompasses all sandpipers, and as a stint it is classified in the genus Calidris. It is classified as monotypic species, meaning that no population differentiation or subspecies have been described.
The genus name is from Ancient Greek kalidris or skalidris, a term used by Aristotle for some grey-coloured waterside birds. The specific fuscicollis is from Latin fuscus, "dusky" and New Latin collis, "necked ".


The white-rumped sandpiper inhabits relatively vegetated patches of the tundra. More specifically, they live in the marshy, heavily vegetated, hummocky arctic tundra of Alaska and Canada during the breeding season. They can be found in various types of wetlands while migrating. During the winter months they inhabit a variety of freshwater and saltwater habitats such as lagoons, estuaries and marshes. In general, they tend to avoid sandy beaches and fast moving water.


They are a long distance migrant, wintering in southern South America and the Caribbean. They are considered one of the most extreme long distance migrants in the world, traversing the entire continent of North America in the span of one month.
The white-rumped sandpiper is a nearctic migrant. After breeding in northern Canada and Alaska, it flies over the Atlantic Ocean to spend the majority of its non-breeding period in South America, particularly along the Patagonian coast in both Chile and Argentina. It also frequently visits Fracasso Beach, Argentina. Many shorebirds concentrate in this area due to the abundance of intertidal invertebrates, especially clams. The white-rumped sandpiper has also been spotted in Venezuela, Suriname, Brazil and Paraguay during its migration. They are rare but regular vagrants to western Europe and a rare vagrant to Australia.
The birds migrate with both nonstop and short-distance multiple-stop flights. During their migration from north to south, they fly over the Atlantic Ocean, gradually moving along the northeastern coast of South America before heading inland towards the islands. This travel generally takes about one month. On their migration from south to north, the white-rumped sandpiper follows a similar path, but does it much more quickly. This migration is done in a fast series of long flights without stopping. One nonstop flight can be as long as 4,200 km (2,600 mi).
Weather patterns play a crucial role in determining the migration route. Birds like to travel so that the temperature, pressure and humidity work with the tailwinds. Strong winds can blow birds off of their regular migration route. For example, the effects of a large storm lead to a higher presence of the white-rumped sandpiper on the King George Islands.


The warming of the climate has led to changes in the number of individuals and the length of their stay in the South Shetland Islands. White-rumped sandpipers are now observed more frequently in this area as a result of both long and short term climate variations. The higher air temperatures, which occur as a result of the northern winds, bring warm, moist air which creates more open habitats and better food resources that allow these birds to persist and survive. Migration routes, as well as both winter and summer foraging grounds, may also be altered.


Their diet consists primarily of small invertebrates including: molluscs, crustaceans, annelids and both adult and larval insects. They are mainly reported to eat aquatic invertebrates. Although it was previously thought they only consumed invertebrates, it has been shown that seeds also make up a portion of the white-rumped sandpiper diet. This discovery lead to the idea that they might be opportunistic feeders depending on time, season and habitat. Stones and algae have also been found in their stomachs but these are most likely ingested by accident. The stones are of the same general shape and size as the seeds so it could be a case of mistaken identity. Algae could also be accidentally ingested through aquatic feeding.
While on mudflats, the white-rumped sandpiper forages by probing in shallow waters and in mud but can also pick up some items from the surface. When they are in the tundra, they must probe deeply in the moss and other vegetation.


The purple sandpiper (Calidris maritima) is a small shorebird in the sandpiper family Scolopacidae. This is a hardy sandpit  that breeds in the arctic and subarctic regions of Eurasia and North America and winters further south on the Atlantic coast.


The purple sandpiper's breeding range extends from the arctic islands of northern Canada, eastwards to Greenland, Iceland, Svalbard and northern Scandinavia across to Western Siberia and the Taymyr Peninsula. In the high arctic the sandpiper breeds at low altitude on the tundra, sometimes far from the coastline, but in the subarctic regions of Sweden and Norway it breeds on barren mountain sides near the limit of the frozen ground. Birds breeding at high latitudes migrate south and spend the winter on rocky shores on both sides of the north Atlantic. They winter along the North America coast as far south as South Carolina and on the eastern Atlantic coast as far south as France and northern Iberia.
Birds wintering in northern Scotland and southwest Ireland migrate to Canada (Baffin Island and Devon Island) to breed.
In Britain, these birds occur in winter in good numbers principally along the east and south coasts, where they favour rocky shorelines adjacent to the sea. It is much rarer as a breeding bird, found only in a localised area of the Cairngorms National Park, where 1–3 pairs have bred since 1978. Records of breeding by this species in the UK are monitored and archived by the Rare Breeding Birds Panel.
They are late migrants and move to rocky, ice-free Atlantic coasts in winter. Most go no further south than North Carolina and northern Portugal. They are fairly gregarious, forming small flocks, often with Ruddy Turnstones. This species is tame and approachable.


It is a very long-distance migrant, and about half of the species breeds in the boggy tundra of northeast Asia, the rest nesting in a range from Alaska to central Canada. The American and most of the Asian birds winter in South America, but some Asian breeders winter in southern and Australia and New Zealand. On migration and in winter, the pectoral sandpiper is typically found in freshwater habitats.
This species also occurs as a regular migrant to western Europe, and is seen most years in Ireland or Great Britain. While the pectoral sandpiper has not been recorded as breeding species in Europe, vagrant individuals were found in Scotland in suitable breeding habitat during summer. Many of the birds occurring in Western Europe may be on a regular migration from Asian breeding grounds to winter in Southern Africa. September 2003 saw a record influx to those two countries, with 40 found in Ireland and 150 in Great Britain. On the US Pacific coast, such stagings of migrant flocks appear to be rarer. Vagrant individuals are sometimes seen elsewhere off the usual migration routes, e.g. on the Marianas, the Marshall Islands and Palau in Micronesia; they are somewhat more frequently encountered on the Hawaiian Islands.
The pectoral sandpiper's migration might be affected by global warming, as is suspected for many Arctic-breeding birds: 100 years ago, migrating pectoral sandpipers were observed to pass through northern Ohio in early-mid May and again in late August; today, the bulk of the northward migration takes place in April already, and most birds do not return until mid-September.
These birds forage on grasslands and mudflats, picking up food by sight, sometimes by probing. They mainly eat arthropods and other invertebrates. The male has a courtship display which involves puffing up his breast, which has a fat sac in the breeding season to enhance his performance.
The pectoral sandpiper builds a steep-sided scrape nest with a considerable volume of lining material. The nest is deep enough that the eggs sit about 3 cm (1.2 in) below ground level, which helps to minimize heat loss from the cool breezes which occur at the latitudes where the species nests. The female lays four eggs.
Pectoral sandpipers have decreased in number 50% since 1974.


The little stint (Calidris minuta) (or Erolia minuta), is a very small wader. It breeds in arctic Europe and Asia, and is a long-distance migrant, wintering south to Africa and south Asia. It occasionally is a vagrant to North America and to Australia. The genus name is from Ancient Greek kalidris or skalidris, a term used by Aristotle for some grey-coloured waterside birds. The specific minuta is Latin for "small.


This species has greenish legs and a short, thin, dark bill. Breeding adults are brown with dark brown streaks on top and white underneath. They have a light line above the eye and a dark crown. In winter, Least sandpipers are grey above. The juveniles are brightly patterned above with rufous coloration and white mantle stripes.
This bird can be difficult to distinguish from other similar tiny shorebirds; these are known collectively as "peeps" or "stints". In particular, least sandpiper is very similar to its Asian counterpart, long-toed stint. It differs from that species in its more compact, shorter-necked appearance, shorter toes, somewhat duller colors, and stronger wingbar.
Measurements:
Length: 5.1-5.9 in (13-15 cm)
Weight: 0.7-1.1 oz (19-30 g)
Wingspan: 10.6-11.0 in (27-28 cm)


The ruff has a distinctive gravy boat appearance, with a small head, medium-length bill, longish neck and pot-bellied body. It has long legs that are variable in colour, being dark greenish in juveniles, pink to orange in adults with some males having reddish orange legs only during the breeding season. In flight, it has a deeper, slower wing stroke than other waders of a similar size, and displays a thin, indistinct white bar on the wing, and white ovals on the sides of the tail. The species shows sexual dimorphism. Although a small percentage of males resemble females, the typical male is much larger than the female and has an elaborate breeding plumage. The males are 29–32 cm (11–13 in) long with a 54–60 cm (21–24 in) wingspan, and weighs about 180 g (6.3 oz). In the May-to-June breeding season, the male's legs, bill and warty bare facial skin are orange, and they have distinctive head tufts and a neck ruff. These ornaments vary on individual birds, being black, chestnut or white, with the colouring solid, barred or irregular. The grey-brown back has a scale-like pattern, often with black or chestnut feathers, and the underparts are white with extensive black on the breast. The extreme variability of the main breeding plumage is thought to have developed to aid individual recognition in a species that has communal breeding displays, but is usually mute.
Outside the breeding season, the male's head and neck decorations and the bare facial skin are lost and the legs and bill become duller. The upperparts are grey-brown, and the underparts are white with grey mottling on the breast and flanks.
The females, or "reeve", is 22–26 cm (8.7–10.2 in) long with a 46–49 cm (18–19 in) wingspan, and weighs about 110 g (3.9 oz). In breeding plumage, they have grey-brown upperparts with white-fringed, dark-centred feathers. The breast and flanks are variably blotched with black. In winter, the females' plumage is similar to that of the male, but the sexes are distinguishable by their size. The plumage of the juvenile ruff resembles the non-breeding adult, but has upperparts with a neat, scale-like pattern with dark feather centres, and a strong buff tinge to the underparts.
Adult male ruffs start to moult into the main display plumage before their return to the breeding areas, and the proportion of birds with head and neck decorations gradually increases through the spring. Second-year birds lag behind full adults in developing breeding plumage. They have a lower body mass and a slower weight increase than full adults, and perhaps the demands made on their energy reserves during the migration flight are the main reason of the delayed moult.
Ruffs of both sexes have an additional moult stage between the winter and final summer plumages, a phenomenon also seen in the bar-tailed godwit. Before developing the full display finery with coloured ruff and tufts, the males replace part of their winter plumage with striped feathers. Females also develop a mix of winter and striped feathers before reaching their summer appearance. The final male breeding plumage results from the replacement of both winter and striped feathers, but the female retains the striped feathers and replaces only the winter feathers to reach her summer plumage. The striped prenuptial plumages may represent the original breeding appearance of this species, the male's showy nuptial feathers evolving later under strong sexual selection pressures.
Adult males and most adult females start their pre-winter moult before returning south, but complete most feather replacement on the wintering grounds. In Kenya, males moult 3–4 weeks ahead of the females, finishing before December, whereas females typically complete feather replacement during December and early January. Juveniles moult from their first summer body plumage into winter plumage during late September to November, and later undergo a pre-breeding moult similar in timing and duration to that of the adults, and often producing as brightly coloured an appearance.
Two other waders can be confused with the ruff. The juvenile sharp-tailed sandpiper is a little smaller than a juvenile female ruff and has a similar rich orange-buff breast, but the ruff is slimmer with a longer neck and legs, a rounder head, and a much plainer face. The buff-breasted sandpiper also resembles a small juvenile ruff, but even the female ruff is noticeably larger than the sandpiper, with a longer bill, more rotund body and scaly-patterned upperparts.


The ruff is a migratory species, breeding in wetlands in colder regions of northern Eurasia, and spends the northern winter in the tropics, mainly in Africa. Some Siberian breeders undertake an annual round trip of up to 30,000 km (19,000 mi) to the West African wintering grounds. There is a limited overlap of the summer and winter ranges in western Europe. The ruff breeds in extensive lowland freshwater marshes and damp grasslands. It avoids barren tundra and areas badly affected by severe weather, preferring hummocky marshes and deltas with shallow water. The wetter areas provide a source of food, the mounds and slopes may be used for leks, and dry areas with sedge or low scrub offer nesting sites. A Hungarian study showed that moderately intensive grazing of grassland, with more than one cow per hectare (2.5 acres), was found to attract more nesting pairs. When not breeding, the birds use a wider range of shallow wetlands, such as irrigated fields, lake margins, and mining subsidence and other floodlands. Dry grassland, tidal mudflats and the seashore are less frequently used. The density can reach 129 individuals per square kilometre (334 per square mile), but is usually much lower.
The ruff breeds in Europe and Asia from Scandinavia and Great Britain almost to the Pacific. In Europe it is found in cool temperate areas, but over its Russian range it is an Arctic species, occurring mainly north of about 65°N. The largest numbers breed in Russia (more than 1 million pairs), Sweden (61,000 pairs), Finland (39,000 pairs) and Norway (14,000 pairs). Although it also breeds from Britain east through the Low Countries to Poland, Germany and Denmark, there are fewer than 2,000 pairs in these more southerly areas.
It is highly gregarious on migration, travelling in large flocks that can contain hundreds or thousands of individuals. Huge dense groups form on the wintering grounds; one flock in Senegal contained a million birds. A minority winter further east to Burma, south China, New Guinea and scattered parts of southern Australia, or on the Atlantic and Mediterranean coasts of Europe. In Great Britain and parts of coastal western Europe, where the breeding and wintering ranges overlap, birds may be present throughout the year. Non-breeding birds may also remain year round in the tropical wintering quarters. The Ruff is an uncommon visitor to Alaska (where it has occasionally bred), Canada and the contiguous states of the US, and has wandered to Iceland, Middle America, northern South America, Madagascar and New Zealand. It has been recorded as breeding well south of its main range in northern Kazakhstan, a major migration stopover area.
The male, which plays no part in nesting or chick care, leaves the breeding grounds in late June or early July, followed later in July by the female and juveniles. Males typically make shorter flights and winter further north than females; for example, virtually all wintering ruffs in Britain are males, whereas in Kenya most are females. Many migratory species use this differential wintering strategy, since it reduces feeding competition between the sexes and enables territorial males to reach the breeding grounds as early as possible, improving their chances of successful mating. Male ruffs may also be able to tolerate colder winter conditions because they are larger than females.
Birds returning north in spring across the central Mediterranean appear to follow a well-defined route. Large concentrations of ruffs form every year at particular stopover sites to feed, and individuals marked with rings or dye reappear in subsequent years. The refuelling sites are closer together than the theoretical maximum travel distance calculated from the mean body mass, and provide evidence of a migration strategy using favoured intermediate sites. The ruff stores fat as a fuel, but unlike mammals, uses lipids as the main energy source for exercise (including migration) and, when necessary, keeps warm by shivering; however, little research has been conducted on the mechanisms by which they oxidise lipids.


Males display during the breeding season at a lek in a traditional open grassy arena. The ruff is one of the few lekking species in which the display is primarily directed at other males rather than to the females, and it is among the small percentage of birds in which the males have well-marked and inherited variations in plumage and mating behaviour. There are three male forms: the typical territorial males, satellite males which have a white neck ruff, and a very rare variant with female-like plumage. The behaviour and appearance for an individual male remain constant through its adult life, and are determined by its genes (see §Biology of variation among males).
The territorial males, about 84% of the total, have strongly coloured black or chestnut ruffs and stake out and occupy small mating territories in the lek. They actively court females and display a high degree of aggression towards other resident males; 5–20 territorial males each hold an area of the lek about 1 m (1.1 yd) across, usually with bare soil in the centre. They perform an elaborate display that includes wing fluttering, jumping, standing upright, crouching with ruff erect, or lunging at rivals. They are typically silent even when displaying, although a soft gue-gue-gue may occasionally be given.
Territorial males are very site-faithful; 90% return to the same lekking sites in subsequent seasons, the most dominant males being the most likely to reappear. Site-faithful males can acquire accurate information about the competitive abilities of other males, leading to well-developed dominance relationships. Such stable relationships reduce conflict and the risk of injury, and the consequent lower levels of male aggression are less likely to scare off females. Lower-ranked territorial males also benefit from site fidelity since they can remain on the leks while waiting for the top males eventually to drop out.
Satellite males, about 16% of the total number, have white or mottled ruffs and do not occupy territories; they enter leks and attempt to mate with the females visiting the territories occupied by the resident males. Resident males tolerate the satellite birds because, although they are competitors for mating with the females, the presence of both types of male on a territory attracts additional females. Females also prefer larger leks, and leks surrounded by taller plants, which give better nesting habitat.
Although satellite males are on average slightly smaller and lighter than residents, the nutrition of the chicks does not, as previously thought, influence mating strategy; rather, the inherited mating strategy influences body size. Resident-type chicks will, if provided with the same amount of food, grow heavier than satellite-type chicks. Satellite males do not have to expend energy to defend a territory, and can spend more time foraging, so they do not need to be as bulky as the residents; indeed, since they fly more, there would be a physiological cost to additional weight.
A third type of male was first described in 2006; this is a permanent female mimic, the first such reported for a bird. About 1% of males are small, intermediate in size between males and females, and do not grow the elaborate breeding plumage of the territorial and satellite males, although they have much larger internal testes than the ruffed males. Although the males of most lekking bird species have relatively small testes for their size, male ruffs have the most disproportionately large testes of any bird.
This cryptic male, or "faeder" (Old English "father") obtains access to mating territories together with the females, and "steals" matings when the females crouch to solicit copulation. The faeder moults into the prenuptial male plumage with striped feathers, but does not go on to develop the ornamental feathers of the normal male. As described above, this stage is thought to show the original male breeding plumage, before other male types evolved. A faeder can be distinguished in the hand by its wing length, which is intermediate between those of displaying males and females. Despite their feminine appearance, the faeders migrate with the larger 'normal' lekking males and spend the winter with them.
The faeders are sometimes mounted by independent or satellite males, but are as often "on top" in homosexual mountings as the ruffed males, suggesting that their true identity is known by the other males. Females never mount males. Females often seem to prefer mating with faeders to copulation with normal males, and normal males also copulate with faeders (and vice versa) relatively more often than with females. The homosexual copulations may attract females to the lek, like the presence of satellite males.
Not all mating takes place at the lek, since only a minority of the males attend an active lek. As alternative strategies, males can also directly pursue females ("following") or wait for them as they approached good feeding sites ("intercepting"). Males switched between the three tactics, being more likely to attend a lek when the copulation rate the previous day was high or when fewer females were available after nesting had started. Lekking rates were low in cold weather early in the season when off-lek males spent most of their time feeding.
The level of polyandry in the ruff is the highest known for any avian lekking species and for any shorebird. More than half of female ruffs mate with, and have clutches fertilised by, more than one male, and individual females mate with males of both main behavioural morphs more often than expected by chance. In lekking species, females can choose mates without risking the loss of support from males in nesting and rearing chicks, since the males take no part in raising the brood anyway. In the absence of this cost, if polyandry is advantageous, it would be expected to occur at a higher rate in lekking than among pair-bonded species.


The nest is a shallow ground scrape lined with grass leaves and stems, and concealed in marsh plants or tall grass up to 400 m (440 yd) from the lek. Nesting is solitary, although several females may lay in the general vicinity of a lek. The eggs are slightly glossy, green or olive, and marked with dark blotches; they are laid from mid-March to early June depending on latitude.
The typical clutch is four eggs, each egg measuring 44 mm × 31 mm (1.7 in × 1.2 in) in size and weighing 21.0 g (0.74 oz) of which 5% is shell. Incubation is by the female alone, and the time to hatching is 20–23 days, with a further 25–28 days to fledging. The precocial chicks have buff and chestnut down, streaked and barred with black, and frosted with white; they feed themselves on a variety of small invertebrates, but are brooded by the female. One brood is raised each year.
Ruffs often show a pronounced inequality in the numbers of each sex. A study of juveniles in Finland found that only 34% were males and 1% were faeders. It appears that females produce a larger proportion of males at the egg stage when they are in poor physical condition. When females are in better condition, any bias in the sex ratio is smaller or absent.
Predators of waders breeding in wet grasslands include birds such as large gulls, common raven, carrion and hooded crows, and great and Arctic skuas; foxes occasionally take waders, and the impact of feral cats and stoats is unknown. Overgrazing can increase predation by making nests easier to find. In captivity, the main causes of chick mortality were stress-related sudden death and twisted neck syndrome. Adults seem to show little evidence of external parasites, but may have significant levels of disease on their tropical wintering grounds, including avian malaria in their inland freshwater habitats, and so they might be expected to invest strongly in their immune systems; however, a 2006 study that analysed the blood of migrating ruffs intercepted in Friesland showed that this bird actually has unexplained low levels of immune responses on at least one measure of resistance. The ruff can breed from its second year, and the average lifespan for birds that have passed the chick stage is about 4.4 years, although a Finnish bird lived to a record 13 years and 11 months.


The ruff normally feeds using a steady walk and pecking action, selecting food items by sight, but it will also wade deeply and submerge its head. On saline lakes in East Africa it often swims like a phalarope, picking items off the surface. It will feed at night as well as during the day. It is thought that Ruff use both visual and auditory cues to find prey. When feeding, the ruff frequently raises its back feathers, producing a loose pointed peak on the back; this habit is shared only by the black-tailed godwit.
During the breeding season, the ruff’s diet consists almost exclusively of the adults and larva of terrestrial and aquatic insects such as beetles and flies. On migration and during the winter, the ruff eats insects (including caddis flies, water-beetles, mayflies and grasshoppers), crustaceans, spiders, molluscs, worms, frogs, small fish, and also the seeds of rice and other cereals, sedges, grasses and aquatic plants. Migrating birds in Italy varied their diet according to what was available at each stopover site. Green aquatic plant material, spilt rice and maize, flies and beetles were found, along with varying amounts of grit. On the main wintering grounds in West Africa, rice is a favoured food during the later part of the season as the ricefields dry out.
Just before migration, the ruff increases its body mass at a rate of about 1% a day, much slower than the bar-tailed godwits breeding in Alaska, which fatten at four times that rate. This is thought to be because the godwit cannot use refuelling areas to feed on its trans-Pacific flight, whereas the ruff is able to make regular stops and take in food during overland migration. For the same reason, the ruff does not physiologically shrink its digestive organs to reduce bodyweight before migrating, unlike the godwit.


Ruffs were formerly trapped for food in England in large numbers; on one occasion, 2,400 were served at Archbishop Neville's enthronement banquet in 1465. The birds were netted while lekking, sometimes being fattened with bread, milk and seed in holding pens before preparation for the table.
...if expedition is required, sugar is added, which will make them in a fortnight's
time a lump of fat: they then sell for two Shillings or half-a-crown a piece… The method of killing them is by cutting off their head with a pair of scissars [sic], the quantity of blood that issues is very great, considering the size of the bird. They are dressed like the Woodcock, with their intestines; and, when killed at the critical time, say the Epicures, are reckoned the most delicious of all morsels.
The heavy toll on breeding birds, together with loss of habitat through drainage and collection by nineteenth-century trophy hunters and egg collectors, meant that the species became almost extinct in England by the 1880s, although recolonisation in small numbers has occurred since 1963. The draining of wetlands from the 1800s onwards in southern Sweden has resulted in the ruff's disappearance from many areas there, although it remains common in the north of the country. The use of insecticides and draining of wetlands has led to a decrease in the number of ruff in Denmark since the early 1900s. There are still areas where the ruff and other wetland birds are hunted legally or otherwise for food. A large-scale example is the capture of more than one million waterbirds (including ruffs) in a single year from Lake Chilwa in Malawi.
Although this bird eats rice on the wintering grounds, where it can make up nearly 40% of its diet, it takes mainly waste and residues from cropping and threshing, not harvestable grain. It has sometimes been viewed as a pest, but the deeper water and presence of invertebrate prey in the economically important early winter period means that the wader has little effect on crop yield.


The ruff has a large range, estimated at 1–10 million square kilometres (0.38–3.8 million square miles) and a population of at least 2,000,000 birds. The European population of 200,000–510,000 pairs, occupying more than half of the total breeding range, seems to have declined by up to 30% over ten years, but this may reflect geographical changes in breeding populations. Numbers in Asia do not appear to be declining, and more ruffs are wintering in Africa. The species as a whole is therefore not believed to approach the thresholds for the population decline criterion of the IUCN Red List (that is, declining more than 30 percent in ten years or three generations). For these reasons, the ruff is classified as "least concern".
The most important breeding populations in Europe, in Russia and Sweden are stable, and the breeding range in Norway has expanded to the south, but populations have more than halved in Finland, Estonia, Poland, Latvia and The Netherlands. Although the small populations in these countries are of limited overall significance, the decline is a continuation of trend towards range contraction that has occurred over the last two centuries. The drop in numbers in Europe has been attributed to drainage, increased fertiliser use, the loss of formerly mown or grazed breeding sites and over-hunting.
Fossils from the Pleistocene suggest that this species bred further south in Europe in the cool periods between glaciations than it does now. Its sensitivity to changing climate as well as to water table levels and the speed of vegetation growth has led to suggestions that its range is affected by global warming, and the ruff might act as an indicator species for monitoring climate change. Potential threats to this species may also include outbreaks of diseases to which it is susceptible such as influenza, botulism and avian malaria.
The ruff is one of the species to which the Agreement on the Conservation of African-Eurasian Migratory Waterbirds (AEWA) applies, where it is allocated to category 2c; that is, the populations in need of special attention as they are showing "significant long-term decline" in much of its range. This commits signatories to regulate the taking of listed species or their eggs, to establish protected areas to conserve habitats for the listed species, to regulate hunting and to monitor the populations of the birds concerned.


Their breeding habitat is the southern tundra in Canada and Alaska near water. They nest on the ground. The male makes several shallow scrapes; the female chooses one and adds grass and other material to line the nest. The female lays 4 eggs; the male assists in incubation. After a few days, the female leaves the young with the male; the young feed themselves.
These birds forage on mudflats, picking up food by sight and feel (bill). They mainly eat aquatic insects and crustaceans.


They are long distance migrants and winter in coastal South America, with some going to the southern United States and the Caribbean. They migrate in flocks which can number in the hundreds of thousands, particularly in favoured feeding locations such as the Bay of Fundy and Delaware Bay. This species is a rare but regular vagrant to western Europe.
Although very numerous, these birds are highly dependent on a few key stopover habitats during their migration, notably Mary's Point and Johnson's Mills along Shepody Bay, an arm of the Bay of Fundy. During the months of July and August, the Nature Conservancy of Canada runs an information center about these shorebirds in Johnson's Mills, New Brunswick.


The red-necked stint (Calidris ruficollis) is a small migratory wader. The genus name is from Ancient Greek kalidris or skalidris, a term used by Aristotle for some grey-coloured waterside birds. The specific ruficollis is from Latin rufus, "red" and collum, "neck".


Red-necked stints are highly gregarious and will form flocks with other small Calidris waders, such as sharp-tailed sandpipers and curlew sandpipers in their non-breeding areas.


C. subruficollis breeds in the open arctic tundra of North America and is a very long-distance migrant, spending the non-breeding season mainly in South America, especially Argentina.
It migrates mainly through central North America, and is uncommon on the coasts. It occurs as a regular wanderer to western Europe, and is not classed as rare in Great Britain or Ireland, where small flocks have occurred. Only the pectoral sandpiper is a more common American shorebird visitor to Europe.
This species nests as far north as Canada including Alaska on the ground, laying four eggs. The male has a display which includes raising the wings to display the white undersides, which is also given on migration, sometimes when no other buff-breasted sandpipers are present. Outside the breeding season, this bird is normally found on short-grass habitats such as airfields or golf-courses, rather than near water.
These birds pick up food by sight, mainly eating insects and other invertebrates. The buff-breasted sandpipers are known to prey on Bombus polaris, a species of bumblebee found within the Arctic Circle. They will either eat the bees or feed them to their young. They are often very tame.
Buff-breasted sandpipers are suspected to have hybridized with the white-rumped or Baird's sandpiper.


This species has been sighted in South Asia on at least three occasions. It is believed that instead of going to Argentina, this bird might have been wind-blown from the Great Plains Flyway of North America and landed up in South Asia. In November 2011, this species was sighted by Dr.Jayan Thomas and P.C. Rajeevan near Kannur, Kerala in South India. and photographed by Dr. Jayan Thomas. The buff-breasted sandpiper has also been recorded in Australia on at least eight occasions.
In 1978, Phillips recorded the species from Sri Lanka, and after that very few sightings were recorded. In 2001 a single bird was also recorded in Marievale, South Africa.


The surfbird (Calidris virgata) is a small stocky wader in the family Scolopacidae. It was once considered to be allied to the turnstones, and placed in the monotypic genus Aphriza, but is now placed in the genus Calidris.
This bird has a short dark bill, yellow legs and a black band at the end of the white rump. In breeding plumage, it has dark streaks on the brownish head and breast with dark spots on its white underparts; the upperparts are dark with rust colouring on the wings. Birds in winter plumage and immature birds are mainly grey on the upperparts and breast and white on the underparts with streaking.
Their breeding habitat is rocky tundra areas in Alaska and the Yukon. The female lays 4 eggs in a depression on the ground lined with vegetation. Both parents look after the young birds, who feed themselves.
These birds migrate to the Pacific coasts of North and South America, from southern Alaska to Tierra del Fuego.
On the nesting grounds, these birds mainly eat insects and some seeds. At other times of year, they eat mollusks and crustaceans found along the surf line on rocky coasts and are usually found in small flocks, often with turnstones.
The song is a whistled tee tee tee.


The surfbird is a large sandpiper, reminiscent of the turnstones or the great knot. It measures 23 to 26 cm (9.1–10.2 in) and weighs 133–251 g (4.7–8.9 oz). The sexes look the same, but the females are slightly larger than males on average.
Surfbirds their breeding plumage from March through August, and have white heads, necks breast and belly which are streaked with black, except for the crown which are rusty. The streaking becomes less prominent lower in the body.


The surfbird has the longest and narrowest non-breeding distribution of any North American bird, being found from Kodiak Island in Alaska to the Straits of Magellan at the southern tip of South America. Along that range it is rarely found more than a few meters from the shore. During the breeding season it is found on the mountains of Alaska and the western Yukon, from 150–1,800 m (490–5,910 ft).
The surfbird starts leaving its breeding grounds from July (the last leaving in October), and begins reaching its main wintering grounds in Chile and Peru in mid August. On its return migration it begins to leave South America in early March. Some birds remain on the wintering grounds year-round. Vagrant birds have been seen in the Falkland Islands, Florida, and Texas. The migration routes are almost entirely coastal, a very small number of birds have been seen inland during migration.
Its breeding habitat is alpine tundra, preferably rocky ridges dominated by scree, rock fields, lichens, dwarf shrubs and Dryas (mountain avens), and less commonly in tundra with mosses and sedges. It is generally found away from suitable habitat that is close to forest. In the non breeding season, it is a rocky shore specialist, feeding on rocky shores, reefs, and ledges on the coast. It will feed from the spray zone on the water's edge to just above the tide line. In some circumstances, it will feed on sandy beaches and mudflats near rocky areas.


The diet of breeding surfbirds is dominated by insects. On one study examining stomach contents, flies (Diptera), including eggs, pupae and adults, composed 55% of the food taken by surfbirds, and beetles Coleoptera 36%. Other insects taken include moths and butterflies (Lepidoptera) and bees and wasps (Hymenoptera). Seeds and snails are also taken, albeit rarely.


The breeding behavior of the surfbird is poorly known. The surfbird arrives in its breeding habitat in early May; due to high winds in their exposed nesting areas the snow has usually cleared by then. Nesting sites are typically on north or west facing slopes. Nests are exposed, and a re simple depressions lined with lichens and, sometimes, Dryas. The species is assumed to be territorial and monogamous. Clutch size is usually four eggs, although smaller clutches have been found.
The eggs are buff-colored with gray or purple markings, and measure 43 mm × 31 mm (1.7 in × 1.2 in). Both sexes incubate the eggs and the incubation period is estimated to be between 22 and 24 days.


The yellow-faced honeyeater (Caligavis chrysops) is a small to medium-sized bird in the honeyeater family, Meliphagidae. It takes its common and scientific names from the distinctive yellow stripes on the sides of its head. Its loud, clear call often begins twenty or thirty minutes before dawn. It is widespread across eastern and southeastern Australia, in open sclerophyll forests from coastal dunes to high-altitude subalpine areas, and woodlands along creeks and rivers. Comparatively short-billed for a honeyeater, it is thought to have adapted to a diet of flies, spiders, and beetles, as well as nectar and pollen from the flowers of plants, such as Banksia and Grevillea, and soft fruits. It catches insects in flight as well as gleaning them from the foliage of trees and shrubs.
Some yellow-faced honeyeaters are sedentary, but hundreds of thousands migrate northwards between March and May to spend the winter in southern Queensland, and return in July and August to breed in southern New South Wales and Victoria. They form socially monogamous pairs and lay two or three eggs in a delicate cup-shaped nest. The success rate can be low, and the pairs nest several times during the breeding season.
Honeyeaters' preferred woodland habitat is vulnerable to the effects of land-clearing, grazing, and weeds. As it is common and widespread, the yellow-faced honeyeater is considered by the International Union for Conservation of Nature (IUCN) to be of least concern for conservation. It is considered a pest in orchards in some areas.


Across its range, the yellow-faced honeyeater is found in a variety of habitats—in open sclerophyll forests from coastal dunes to high-altitude subalpine areas, and often in riparian woodlands. It most commonly dwells in open forests dominated by spotted gum (Corymbia maculata) with ironbarks and stringybarks, such as narrow-leaved ironbark (Eucalyptus crebra) and silver-leaved ironbark (E. melanophloia), with a light, shrubby understorey, and less often in dry, open forests and woodlands dominated by Angophora, Acacia, Banksia, Casuarina or Callitris, or in high-altitude, tall, open forests of alpine ash (Eucalyptus delegatensis) and woodlands dominated by snow gum (E. pauciflora) or white box (E. albens). It has been recorded in coastal heath when banksias are blooming, and among flowering mangroves. It occupies areas infested with weeds, such as Scotch broom (Cytisus scoparius) and blackberries, and in developed areas including orchards, parks and gardens, where it feeds on cultivated fruit and flowers. It can be found in forests regenerating after fire or logging, though it is more common in mature forests. Where it is found in woodland, it is usually woodland near forest or with an understory of sclerophyll plants.


There are resident populations of the yellow-faced honeyeater throughout its range, but it is for the most part a seasonal, latitudinal, daytime migrant. During the autumn (March to May), it migrates north along the highlands and coastal fringe of eastern Australia to southern Queensland, to return in the spring (August to October) of the same year. The birds commonly move in flocks of 10 to 100 birds, but occasionally in larger groups of 1,000 or more. The groups can include other species, such as the white-naped honeyeater, fuscous honeyeater, noisy friarbird, and silvereye. They move in successive flocks at a rate of up to several thousand birds an hour. Over 100,000 birds were recorded passing Hastings Point in New South Wales over the course of a single day in May 1965. The species is able to detect geomagnetic fields, and uses them to navigate while migrating. Experiments, where the vertical component of the magnetic field was reversed, indicate that the magnetic compass of the yellow-faced honeyeater is based on the inclination of the field lines and not on polarity, meaning they distinguish between the direction of the equator and the South Pole, rather than north and south. Their flight is in one general direction, but is not in a straight line, as the flocks stay in vegetated areas, negotiate gaps in the mountain ranges, and detour around cities.
The migration of many birds in Australia, including honeyeaters, has generally been described as occurring mainly in response to external environmental stimuli, such as food availability or an influx of water. The yellow-faced honeyeater has been found to have a broad range of characteristics that are more often associated with Northern Hemisphere migrants. These are an annual cycle of migratory restlessness, seasonally appropriate orientation based on magnetic, solar and polarised light cues, and a migration program based on the magnetic inclination compass.


Comparatively short-billed for a honeyeater, the yellow-faced honeyeater is thought to have adapted to a mixed diet. Its diet consists of nectar, pollen, fruit, seeds, honeydew, and insects. It is arboreal, foraging primarily among the foliage and flowers of trees, shrubs, and mistletoes, less often on branches and tree-trunk, and rarely on the ground. Yellow-faced honeyeaters feed on nectar around 40% of the time, and on insects around 60% of the time. The yellow-faced honeyeater feeds on insects by gleaning, sallying, catching in flight, or probing in bark crevices. The insects eaten are primarily Diptera (flies, mosquitoes, maggots, gnats, and midges), beetles, and spiders. A study of the pollen on the bills and foreheads of captured birds found that 70% carried pollen from silver banksia (Banksia marginata), 61% from heath-leaved banksia (Banksia ericifolia), and 22% carried pollen from other plants in the area including fern-leaved banksia (Banksia oblongifolia), mountain devil (Lambertia formosa), and green spider-flower (Grevillea mucronulata).
In April and May, before the autumn migration, the yellow-faced honeyeater increases its nectar consumption, which increases its body mass. The average body mass in late autumn of 17.5 grams (0.62 oz) is 13% higher than the average recorded between January and April, and the yellow-faced honeyeater begins the migration with healthy fat reserves.


The yellow-faced honeyeater breeds in monogamous pairs in a breeding season that extends from July to March, with migrating birds nesting later than sedentary birds. They nest solitarily in all-purpose territories that both parents defend against conspecifics and other species including thornbills, spinebills and silvereyes, although the male is involved in more aggressive interactions than the female. Within a breeding season, females lay two or three clutches of eggs, re-nesting with the same partner in the same territory. Banded birds have been identified in the same territory for periods of up to five years.
The nest is built in the understorey shrubs, relatively close to the ground. Nests have been recorded in prickly coprosma (Coprosma quadrifida), Cassinia, tea-trees (Melaleuca), eucalypts and acacias, as well as in garden shrubs. The nest is a fragile, cup-shaped structure, swollen at the sides and narrower at the rim. The female builds the nest, but is often accompanied by the male as she gathers nesting material. Most nests are built of greenish material, which varies with the location; in coastal areas, grass is the primary material; in mountain forests, the nest is often covered with moss. One bird was recorded repeatedly flying between the nest and a koala 36 metres (118 ft) away and plucking the long hair near its ears to incorporate in the nest. The nests are very fine, with the eggs visible through the gauze-like walls, and they sometimes fall apart. They have been known to disintegrate with eggs and nestlings falling through the bottom.
The female undertakes the incubation alone. Eggs are oval, approximately 21 millimetres (0.83 in) long and 14 millimetres (0.55 in) wide, and pinkish white in colour with spots and blotches of dark reddish-brown. The clutch size varies from one to three eggs, and eggs take around two weeks to hatch. Upon hatching, both parents feed the nestlings and remove faecal pellets. The chicks fledge after thirteen days, and leave the parental territory after a further two weeks. The success rate can be as low as 16% of eggs developing into fledged young, with nest failure, hot weather, heavy rain, human activity (including fungicide spraying and nest damage), egg destruction by brood parasites, and predation by brown snakes, cats, and currawongs, all recorded as contributing to brood failure. Among the species that parasitize the nests of yellow-faced honeyeaters are fan-tailed cuckoos, brush cuckoos, pallid cuckoos, shining bronze-cuckoos, and Horsfield's bronze-cuckoos. The yellow-faced honeyeater promptly nests again after both successful and failed breeding attempts.
A paternity analysis of yellow-faced honeyeater nestlings found that 10 of 18 nestlings were fathered by the male of the nesting pair, with clear evidence for extra-pair paternity in the case of the remaining 44%. This conflicts with the usual pattern, where genetic monogamy is linked to the characteristics of strong social pairing, essential paternal contributions to brood-rearing, and to sexual monomorphism; characteristics that are exhibited by the yellow honeyeater, for example.


It is a rhizomatous herbaceous perennial plant growing in bogs and ponds. The leaves are rounded to heart-shaped, 6–12 cm (2+1⁄4–4+3⁄4 in) long on a 10–20 cm (4–8 in) petiole, and 4–12 cm (1+1⁄2–4+3⁄4 in) broad. The greenish-yellow inflorescence is produced on a spadix about 4–6 cm (1+1⁄2–2+1⁄4 in) long, enclosed in a white spathe. The fruit is a cluster of red berries, each berry containing several seeds.
The plant is very poisonous when fresh due to its high oxalic acid content, but the rhizome (like that of Caladium, Colocasia, and Arum) is edible after drying, grinding, leaching and boiling.


It is native to cool temperate regions of the Northern Hemisphere, in central, eastern and northern Europe (France and Norway eastward), northern Asia and northern North America (Alaska, Canada, and northeastern contiguous United States).


Callimorpha dominula has a wingspan of 45–55 millimetres (1.8–2.2 in). Adults of this species are quite variable in color. The forewings usually have a metallic-green sheen on the blackish areas, with white and yellow or orange markings. Hindwings are red with three large and irregular black markings. These moths may also occur in rare color forms, one with yellow hindwings and body and one with extended black on hindwings. The thorax is black glossed with green and shows two longitudinal short yellow stripes. The abdomen is black. The scarlet tiger moth has developed mouthparts, that allow it to feed on nectar. The caterpillars can reach a length of about 40 millimetres (1.6 in). They are dark gray with yellow stripes and small white dots.


The California quail (Callipepla californica), also known as the California valley quail or Valley quail, is a small ground-dwelling bird in the New World quail family. These birds have a curving crest or plume, made of six feathers, that droops forward: black in males and brown in females; the flanks are brown with white streaks. Males have a dark brown cap and a black face with a brown back, a grey-blue chest and a light brown belly. Females and immature birds are mainly grey-brown with a light-colored belly.
Their closest relative is Gambel's quail which has a more southerly distribution and a longer crest at 2.5 in (6.4 cm), a brighter head and lacks the scaly appearance of the California Quail. The two species separated about 1–2 million years ago, during the Late Pliocene or Early Pleistocene. It was selected as the state bird of California in 1931.


The California quail is a highly sociable bird that often gathers in small flocks known as "coveys". One of their daily communal activities is a dust bath. A group of quail will select an area where the ground has been newly turned or is soft, and using their underbellies, will burrow downward into the soil some one to two inches. They then wriggle about in the indentations they have created, flapping their wings and ruffling their feathers, causing dust to rise in the air. They seem to prefer sunny places in which to create these dust baths. An ornithologist is able to detect the presence of quail in an area by spotting the circular indentations left behind in the soft dirt, some 7–15 cm (2.8–5.9 in) in diameter.
They are year-round residents. Although this bird coexists well at the edges of urban areas, it is declining in some areas as human populations increase. They were originally found mainly in the southwestern United States but they have been introduced into other areas including British Columbia, Hawaii, Chile, Uruguay, Brazil, Argentina, Peru, South Africa, New Zealand, and to Norfolk Island and King Island in Australia.
These birds forage on the ground, often scratching at the soil. They can sometimes be seen feeding at the sides of roads. Their diet consists mainly of seeds and leaves, but they also eat some berries and insects; for example, Toyon berries are a common food source. If startled, these birds explode into short rapid flight, called "flushing". Given a choice, they will normally escape on foot.


The scaled quail (Callipepla squamata), also commonly called blue quail or cottontop, is a species of the New World quail family. It is a bluish gray bird found in the arid regions of the Southwestern United States to Central Mexico. This species is an early offshoot of the genus Callipepla, diverging in the Pliocene.
This bird is named for the scaly appearance of its breast and back feathers. Along with its scaly markings, the bird is easily identified by its white crest that resembles a tuft of cotton.
The nest is typically a grass-lined hollow containing 9–16 speckled eggs. When disturbed, it prefers to run rather than fly.
Widespread and common throughout its range, the scaled quail is evaluated as Least Concern on the IUCN Red List of Threatened Species.


Scaled quail occur from south-central Arizona, northern New Mexico, east-central Colorado, and southwestern Kansas south through western Oklahoma and western and central Texas into Mexico to northeastern Jalisco, Guanajuato, Queretaro, Hidalgo, and western Tamaulipas. It has been introduced to Cuba, Jamaica, Haiti, Dominican Republic, Puerto Rico and to the United States in Hawaii, central Washington, eastern Nevada, and Nebraska, but is only considered established in central Washington and eastern Nevada.
Scaled quail has formed several subspecies, 3 of which range into the USA:
Callipepla squamata squamata Vigors, 1830 (Altiplano scaled quail). The nominate subspecies; it is only found on the Central Plateau (altiplano) of Mexico.
Callipepla squamata pallida Brewster, 1881 (northern scaled quail). The most common subspecies, it occurs from Arizona and New Mexico to Colorado and just into Oklahoma, and western Texas, northern Chihuahua, and Sonora. It is paler than the nominate subspecies.
Callipepla squamata hargravei Rea, 1973 (Upper Sonoran scaled quail). A form of arid habitat, it is only found in the area where the states of Colorado, Kansas and Oklahoma meet, and in northwestern New Mexico. It is the palest subspecies, adapted to dry and sandy habitat.
Callipepla squamata castanogastris Brewster, 1883 (chestnut-bellied scaled quail). Found in southern Texas from Eagle Pass and San Antonio south to adjacent northwestern Mexico (Coahuila, Nuevo León, and Tamaulipas). Similar individuals are sometimes found in the extreme northeast and west of the species' range. The chestnut brown belly distinguishes it from all other subspecies; it is also darker than the other two found in the USA.


According to Ligon, the distribution of scaled quail is largely coextensive with mesquite (Prosopis spp.), condalia (Condalia spp.), and
cholla (Cylindropuntia spp.).
In Oklahoma, scaled quail occur in sand sagebrush (Artemisia filifolia)-grassland, pinyon-juniper (Pinus spp.-Juniperus spp.), and shortgrass High Plains. Sand sagebrush-grasslands include sand sagebrush, soapweed yucca (Yucca glauca), skunkbush sumac (Rhus trilobata), and sand plum (Prunus watsonii). Scaled Quail in Oklahoma inhabit rough or rolling land, especially where sagebrush (Artemisia spp.), mesquite, cactus (Opuntia spp. and others), yucca (Yucca spp.), juniper, sand shinnery oak (Quercus havardii), and rocks furnish cover.
In Colorado, scaled quail occupy sand sagebrush and/or yucca stands on sandy soils. The cover types used by scaled quail in Colorado are, in descending order, sand sagebrush-grassland, pinyon-juniper, dense cholla-grassland, dryland farmland, irrigated farmland, and greasewood (Sarcobatus spp.)-saltbush (Atriplex spp.) washes. Scaled Quail made little or no use of sparse cholla-grassland, riparian areas, reseeded grasslands, or shortgrass prairie disclimax.


Scaled quail inhabit dry, open valleys, plains, foothills, rocky slopes, draws, gullies, and canyons that have a mixture of bare ground, low herbaceous growth, and scattered brushy cover. Good scaled quail habitat is characterized by low-growing grasses with forbs and shrubs. Overall ground cover is between 10 and 50%. Trees and shrubs should be less than 6.6 feet (2.0 m) tall. Scaled quail avoid the dense growth associated with streamsides. Transmitter-fitted scaled quail had individual home range sizes of 52 and 60 acres (21 and 24 ha).
An absolute requirement by scaled quail for a source of open water has not been established; there is some debate in the literature whether there is such a requirement. Scaled quail have been reported as inhabiting an area 7 or 8 miles (11 or 13 kilometres) from the nearest water in Arizona. In New Mexico, it was not unusual to find Scaled Quail 10 to 15 miles (16 to 24 kilometres) from water. Wallmo  observed winter coveys 3 and 7 miles (4.8 and 11.3 kilometres) from water in Big Bend National Park in southwestern Texas.
In Arizona, scaled quail summer habitat is seldom within 660 feet (200 m) of water. Scaled quail were observed drinking at stock tanks from April to June (which was a dry period during the course of the study) every 2 to 3 days. In Oklahoma, scaled quail often migrate to farms and ranches in winter and are thus closer to a source of water in winter than in summer. DeGraaf and others  reported that in winter, scaled quail are usually found within 1.25 miles (2.01 km) of a source of water.


Feeding cover: scaled quail use grass clumps and shrubs for cover while feeding. In one study they were frequently seen crossing 82 to 165 feet (25 to 50 metres) of bare ground. When disturbed, scaled quail hid in snakeweed (Gutierrezia spp.) or in grass clumps. In June and July foraging occurs on open grasslands which are not used at other times.
Loafing cover: scaled quail coveys occupy loafing or resting cover after early morning feeding periods. Scaled quail occupy desert grassland or desert scrub with a minimum of one loafing covert per approximately 70 acres (28 hectares). In northwestern Texas, loafing coverts were characterized by: (1) overhead woody cover, (2) lateral screening cover, (3) a central area with bare soil, and (4) one or more paths through the lateral cover. Covert heights ranged from 1.6 to 5.9 feet (0.49 to 1.80 m) high and 2.6 to 6.9 feet (0.79 to 2.10 m) in diameter. Cholla formed all or part of the overhead cover of 85% of coverts, even though they were dominant at only 12% of the study locations. In areas where scaled quail occur without cholla, woody species such as wolfberry (Lycium spp.) and mesquite are important for overhead cover. In Oklahoma pinyon-juniper habitats, scaled quail use the shade of tree cholla (Cylindropuntia imbricata) and human-made structures. In Arizona, scaled quail occupied wolfberry and mesquite 1.7 to 5 feet (0.52 to 1.52 m) tall for loafing cover. This overhead cover provides midday shade, but is open at the base to allow easy escape from predators. In Oklahoma, winter home ranges always contained skunkbush sumac, tree cholla, or human-made structures providing overhead cover.
Night-roosting cover: scaled quail roosts were observed in yucca (Yucca angustifolia), tree cholla, and true mountain-mahogany (Cercocarpus montanus)-yucca-fragrant sumac (Rhus aromatica) vegetation types. The height of vegetation used for night roosts was less than 1.6 feet (0.49 m).
Nesting cover: In March or April winter coveys spread out into areas with less cover. This use of areas with less cover coincides with a seasonal decrease in the number of raptors in the same area. Scaled quail nests are constructed under tufts of grasses, and are sheltered by sagebrush (Artemisia spp.), creosotebush (Larrea tridentata), mesquite, catclaw acacia (Acacia greggii), cactus, or yucca; under dead Russian-thistle (Salsola kali), mixed forbs, or soapweed yucca; or sheltered in old machinery or other human-made debris. In Oklahoma, 66% of nests were in one of four situations: (1) dead Russian-thistle, (2) machinery and junk, (3) mixed forbs, and (4) soapweed yucca. In New Mexico, ordination of breeding birds and vegetative microhabitats indicated that scaled quail were associated with increased levels of patchiness and increased cover of mesquite and cactus.


Scaled quail are opportunistic eaters. Seeds are consumed year-round. Large seeds (such as those of mesquite and snakeweed) are important in Scaled quail diets. Other seeds include those of elbowbush (Forestiera angustifolia), catclaw acacia, mesquite, hackberry (Celtis spp.), Russian-thistle, rough pigweed (Amaranthus retroflexus), and sunflowers, ragweeds (Ambrosia spp.), and other Asteraceous plants. Scaled quail consume more grass seeds than do other quail species. Other dietary components include leaves, fruits, and insects. Summer diets are high in green vegetation and insects, which are also important sources of moisture.
In Oklahoma, small groups of scaled quail feed among soapweed yucca and in soapweed yucca-sand sagebrush ranges, weed patches, and grain stubble. Also in Oklahoma, early winter foods apparently eaten when other foods are not available included snow-on-the-mountain (Euphorbia marginata), sand paspalum (Paspalum stramineum), field sandbur (Cenchrus pauciflorus), purslane (Portulaca spp.), skunkbush sumac, Fendler spurge (Euphorbia fendleri), and leaf bugs. Jimsonweed (Datura stramonium) and juniper berries were always avoided. Winter foods of the scaled quail in Oklahoma include Russian-thistle and sunflower (Helianthus spp.) seeds.
In northwestern Texas, selection of foods by scaled quail was dependent on foraging techniques, availability, and seed size. Small seeds were selected when they were still on the plant and could be easily stripped, but were not eaten once they had fallen, presumably because they were too small and/or too hard to find. Broom snakeweed (Gutierrezia sarothrae) was a staple in winter diets; it was not highly selected but was consumed in proportion to its availability (and lack of availability of choice items). Generally, in Texas grass seeds (mainly tall dropseed [Sporobolus asper] and rough tridens [Tridens muticus]) were major constituents of scaled quail diets. This was attributed to a precipitation pattern that resulted in a relatively higher amount of grass seed available, and a lower amount of available forbs. In the same study green vegetation formed a higher proportion of the diet than reported for other areas.
In southwestern Texas, chestnut-bellied scaled quail consumed woody plant seeds and green vegetation. The seeds of brush species comprised 68% of the contents of 32 scaled quail crops. Green food, chiefly wild carrot (Daucus carota) and clover (Trifolium spp.) made up 7.17%. Elbowbush was the single most important source, followed by Roemer acacia (Acacia roemeriana), desert-yaupon (Schaefferia cuneifolia), and spiny hackberry (Celtis pallida).
In southeastern New Mexico, staples (comprising at least 5% of scaled quail diet in both summer and winter) were mesquite and croton (Croton spp.) seeds, green vegetation, and snout beetles. Nonpreferred foods eaten in winter and available but not consumed in summer included broom snakeweed (the main winter food), crown-beard (Verbesina encelioides), cycloloma (Cycloloma atriplicifolium), and lace bugs. Mesquite seeds and broom snakeweed seeds together made up 75% of the winter diet. Grasshoppers were a summer staple. Insect galls, cicadas, scarab beetles, spurge (Euphorbia spp.), plains bristlegrass (Setaria macrostachya) seeds, and white ratany (Krameria grayi) were consumed in a less pronounced seasonal pattern. Another study reported substantial amounts of prairie sunflower seeds (Helianthus petiolaris) and pigweed (Amaranthus spp.) seeds in the diet of scaled quail.
Scaled quail feed in alfalfa (Medicago spp.) fields.


In Arizona, potential scaled quail predators include mammals, birds, and reptiles. Most scaled quail kills are made by avian predators including northern harrier (Circus cyaneus), red-tailed hawk (Buteo jamaicensis), American kestrel (Falco sparverius), prairie falcon (Falco mexicanus), and great horned owl (Bubo virginianus). In New Mexico, predators on scaled quail include hawks, owls, coyote (Canis latrans), and snakes. In Colorado, potential predators of scaled quail include coyote, gray fox (Urocyon cinereoargenteus), red fox (Vulpes vulpes), kit fox (V. velox), bobcat (Lynx rufus), northern harrier, rough-legged hawk (Buteo lagopus), prairie falcon, peregrine falcon (Falco peregrinus), American kestrel, golden eagle (Aquila chrysaetos), and bald eagle (Haliaeetus leucocephalus).
Scaled quail are popular gamebirds.