Coyotes occasionally mate with domestic dogs, sometimes producing crosses colloquially known as "coydogs". Such matings are rare in the wild, as the mating cycles of dogs and coyotes do not coincide, and coyotes are usually antagonistic towards dogs. Hybridization usually only occurs when coyotes are expanding into areas where conspecifics are few, and dogs are the only alternatives. Even then, pup survival rates are lower than normal, as dogs do not form pair bonds with coyotes, thus making the rearing of pups more difficult. In captivity, F₁ hybrids (first generation) tend to be more mischievous and less manageable as pups than dogs, and are less trustworthy on maturity than wolf-dog hybrids. Hybrids vary in appearance, but generally retain the coyote's usual characteristics. F₁ hybrids tend to be intermediate in form between dogs and coyotes, while F₂ hybrids (second generation) are more varied. Both F₁ and F₂ hybrids resemble their coyote parents in terms of shyness and intrasexual aggression. Hybrids are fertile and can be successfully bred through four generations. Melanistic coyotes owe their black pelts to a mutation that first arose in domestic dogs. A population of nonalbino white coyotes in Newfoundland owe their coloration to a melanocortin 1 receptor mutation inherited from Golden Retrievers.
Coyotes have hybridized with wolves to varying degrees, particularly in eastern North America. The so-called "eastern coyote" of northeastern North America probably originated in the aftermath of the extermination of gray and eastern wolves in the northeast, thus allowing coyotes to colonize former wolf ranges and mix with the remnant wolf populations. This hybrid is smaller than either the gray or eastern wolf, and holds smaller territories, but is in turn larger and holds more extensive home ranges than the typical western coyote. As of 2010, the eastern coyote's genetic makeup is fairly uniform, with minimal influence from eastern wolves or western coyotes. Adult eastern coyotes are larger than western coyotes, with female eastern coyotes weighing 21% more than male western coyotes. Physical differences become more apparent by the age of 35 days, with eastern coyote pups having longer legs than their western counterparts. Differences in dental development also occurs, with tooth eruption being later, and in a different order in the eastern coyote. Aside from its size, the eastern coyote is physically similar to the western coyote. The four color phases range from dark brown to blond or reddish blond, though the most common phase is gray-brown, with reddish legs, ears, and flanks. No significant differences exist between eastern and western coyotes in aggression and fighting, though eastern coyotes tend to fight less, and are more playful. Unlike western coyote pups, in which fighting precedes play behavior, fighting among eastern coyote pups occurs after the onset of play. Eastern coyotes tend to reach sexual maturity at two years of age, much later than in western coyotes.
Eastern and red wolves are also products of varying degrees of wolf-coyote hybridization. The eastern wolf probably was a result of a wolf-coyote admixture, combined with extensive backcrossing with parent gray wolf populations. The red wolf may have originated during a time of declining wolf populations in the Southeastern Woodlands, forcing a wolf-coyote hybridization, as well as backcrossing with local parent coyote populations to the extent that about 75–80% of the modern red wolf's genome is of coyote derivation.


Like the Eurasian golden jackal, the coyote is gregarious, but not as dependent on conspecifics as more social canid species like wolves are. This is likely because the coyote is not a specialized hunter of large prey as the latter species is. The basic social unit of a coyote pack is a family containing a reproductive female. However, unrelated coyotes may join forces for companionship, or to bring down prey too large to attack singly. Such "nonfamily" packs are only temporary, and may consist of bachelor males, nonreproductive females and subadult young. Families are formed in midwinter, when females enter estrus. Pair bonding can occur 2–3 months before actual copulation takes place. The copulatory tie can last 5–45 minutes. A female entering estrus attracts males by scent marking and howling with increasing frequency. A single female in heat can attract up to seven reproductive males, which can follow her for as long as a month. Although some squabbling may occur among the males, once the female has selected a mate and copulates, the rejected males do not intervene, and move on once they detect other estrous females. Unlike the wolf, which has been known to practice both monogamous and bigamous matings, the coyote is strictly monogamous, even in areas with high coyote densities and abundant food. Females that fail to mate sometimes assist their sisters or mothers in raising their pups, or join their siblings until the next time they can mate. The newly mated pair then establishes a territory and either constructs their own den or cleans out abandoned badger, marmot, or skunk earths. During the pregnancy, the male frequently hunts alone and brings back food for the female. The female may line the den with dried grass or with fur pulled from her belly. The gestation period is 63 days, with an average litter size of six, though the number fluctuates depending on coyote population density and the abundance of food.
Coyote pups are born in dens, hollow trees, or under ledges, and weigh 200 to 500 g (0.44 to 1.10 lb) at birth. They are altricial, and are completely dependent on milk for their first 10 days. The incisors erupt at about 12 days, the canines at 16, and the second premolars at 21. Their eyes open after 10 days, by which point the pups become increasingly more mobile, walking by 20 days, and running at the age of six weeks. The parents begin supplementing the pup's diet with regurgitated solid food after 12–15 days. By the age of four to six weeks, when their milk teeth are fully functional, the pups are given small food items such as mice, rabbits, or pieces of ungulate carcasses, with lactation steadily decreasing after two months. Unlike wolf pups, coyote pups begin seriously fighting (as opposed to play fighting) prior to engaging in play behavior. A common play behavior includes the coyote "hip-slam". By three weeks of age, coyote pups bite each other with less inhibition than wolf pups. By the age of four to five weeks, pups have established dominance hierarchies, and are by then more likely to play rather than fight. The male plays an active role in feeding, grooming, and guarding the pups, but abandons them if the female goes missing before the pups are completely weaned. The den is abandoned by June to July, and the pups follow their parents in patrolling their territory and hunting. Pups may leave their families in August, though can remain for much longer. The pups attain adult dimensions at eight months and gain adult weight a month later.


While the popular consensus is that olfaction is very important for hunting, two studies that experimentally investigated the role of olfactory, auditory, and visual cues found that visual cues are the most important ones for hunting in red foxes and coyotes.
When hunting large prey, the coyote often works in pairs or small groups. Success in killing large ungulates depends on factors such as snow depth and crust density. Younger animals usually avoid participating in such hunts, with the breeding pair typically doing most of the work. Unlike the wolf, which attacks large prey from the rear, the coyote approaches from the front, lacerating its prey's head and throat. Like other canids, the coyote caches excess food. Coyotes catch mouse-sized rodents by pouncing, whereas ground squirrels are chased. Although coyotes can live in large groups, small prey is typically caught singly. Coyotes have been observed to kill porcupines in pairs, using their paws to flip the rodents on their backs, then attacking the soft underbelly. Only old and experienced coyotes can successfully prey on porcupines, with many predation attempts by young coyotes resulting in them being injured by their prey's quills. Coyotes sometimes urinate on their food, possibly to claim ownership over it. Recent evidence demonstrates that at least some coyotes have become more nocturnal in hunting, presumably to avoid humans.
Coyotes may occasionally form mutualistic hunting relationships with American badgers, assisting each other in digging up rodent prey. The relationship between the two species may occasionally border on apparent "friendship", as some coyotes have been observed laying their heads on their badger companions or licking their faces without protest. The amicable interactions between coyotes and badgers were known to pre-Columbian civilizations, as shown on a Mexican jar dated to 1250–1300 CE depicting the relationship between the two.
Food scraps, pet food, and animal feces may attract a coyote to a trash can.


Being both a gregarious and solitary animal, the variability of the coyote's visual and vocal repertoire is intermediate between that of the solitary foxes and the highly social wolf. The aggressive behavior of the coyote bears more similarities to that of foxes than it does that of wolves and dogs. An aggressive coyote arches its back and lowers its tail. Unlike dogs, which solicit playful behavior by performing a "play-bow" followed by a "play-leap", play in coyotes consists of a bow, followed by side-to-side head flexions and a series of "spins" and "dives". Although coyotes will sometimes bite their playmates' scruff as dogs do, they typically approach low, and make upward-directed bites. Pups fight each other regardless of sex, while among adults, aggression is typically reserved for members of the same sex. Combatants approach each other waving their tails and snarling with their jaws open, though fights are typically silent. Males tend to fight in a vertical stance, while females fight on all four paws. Fights among females tend to be more serious than ones among males, as females seize their opponents' forelegs, throat, and shoulders.


The coyote has been described as "the most vocal of all [wild] North American mammals". Its loudness and range of vocalizations was the cause for its binomial name Canis latrans, meaning "barking dog". At least 11 different vocalizations are known in adult coyotes. These sounds are divided into three categories: agonistic and alarm, greeting, and contact. Vocalizations of the first category include woofs, growls, huffs, barks, bark howls, yelps, and high-frequency whines. Woofs are used as low-intensity threats or alarms and are usually heard near den sites, prompting the pups to immediately retreat into their burrows. Growls are used as threats at short distances but have also been heard among pups playing and copulating males. Huffs are high-intensity threat vocalizations produced by rapid expiration of air. Barks can be classed as both long-distance threat vocalizations and alarm calls. Bark howls may serve similar functions. Yelps are emitted as a sign of submission, while high-frequency whines are produced by dominant animals acknowledging the submission of subordinates. Greeting vocalizations include low-frequency whines, 'wow-oo-wows', and group yip howls. Low-frequency whines are emitted by submissive animals and are usually accompanied by tail wagging and muzzle nibbling. The sound known as 'wow-oo-wow' has been described as a "greeting song". The group yip howl is emitted when two or more pack members reunite and may be the final act of a complex greeting ceremony. Contact calls include lone howls and group howls, as well as the previously mentioned group yip howls. The lone howl is the most iconic sound of the coyote and may serve the purpose of announcing the presence of a lone individual separated from its pack. Group howls are used as both substitute group yip howls and as responses to either lone howls, group howls, or group yip howls.


Prior to the near extermination of wolves and cougars, the coyote was most numerous in grasslands inhabited by bison, pronghorn, elk, and other deer, doing particularly well in short-grass areas with prairie dogs, though it was just as much at home in semiarid areas with sagebrush and jackrabbits or in deserts inhabited by cactus, kangaroo rats, and rattlesnakes. As long as it was not in direct competition with the wolf, the coyote ranged from the Sonoran Desert to the alpine regions of adjoining mountains or the plains and mountainous areas of Alberta. With the extermination of the wolf, the coyote's range expanded to encompass broken forests from the tropics of Guatemala and the northern slope of Alaska.
Coyotes walk around 5–16 kilometres (3–10 mi) per day, often along trails such as logging roads and paths; they may use iced-over rivers as travel routes in winter. They are often crepuscular, being more active around evening and the beginning of the night than during the day. Like many canids, coyotes are competent swimmers, reported to be able to travel at least 0.8 kilometres (0.5 mi) across water.


The coyote is ecologically the North American equivalent of the Eurasian golden jackal. Likewise, the coyote is highly versatile in its choice of food, but is primarily carnivorous, with 90% of its diet consisting of meat. Prey species include bison (largely as carrion), white-tailed deer, mule deer, moose, elk, bighorn sheep, pronghorn, rabbits, hares, rodents, birds (especially galliformes, roadrunners, young water birds and pigeons and doves), amphibians (except toads), lizards, snakes, turtles and tortoises, fish, crustaceans, and insects. Coyotes may be picky over the prey they target, as animals such as shrews, moles, and brown rats do not occur in their diet in proportion to their numbers. However, terrestrial and/or burrowing small mammals such as ground squirrels and associated species (marmots, prairie dogs, chipmunks) as well as voles, pocket gophers, kangaroo rats and other ground-favoring rodents may be quite common foods, especially for lone coyotes. More unusual prey include fishers, young black bear cubs, harp seals and rattlesnakes. Coyotes kill rattlesnakes mostly for food (but also to protect their pups at their dens) by teasing the snakes until they stretch out and then biting their heads and snapping and shaking the snakes. Birds taken by coyotes may range in size from thrashers, larks and sparrows to adult wild turkeys and, rarely, brooding adult swans and pelicans. If working in packs or pairs, coyotes may have access to larger prey than lone individuals normally take, such as various prey weighing more than 10 kg (22 lb). In some cases, packs of coyotes have dispatched much larger prey such as adult Odocoileus deer, cow elk, pronghorns and wild sheep, although the young fawn, calves and lambs of these animals are considerably more often taken even by packs, as well as domestic sheep and domestic cattle. In some cases, coyotes can bring down prey weighing up to 100 to 200 kg (220 to 440 lb) or more. When it comes to adult ungulates such as wild deer, they often exploit them when vulnerable such as those that are infirm, stuck in snow or ice, otherwise winter-weakened or heavily pregnant, whereas less wary domestic ungulates may be more easily exploited.
Although coyotes prefer fresh meat, they will scavenge when the opportunity presents itself. Excluding the insects, fruit, and grass eaten, the coyote requires an estimated 600 g (1.3 lb) of food daily, or 250 kg (550 lb) annually. The coyote readily cannibalizes the carcasses of conspecifics, with coyote fat having been successfully used by coyote hunters as a lure or poisoned bait. The coyote's winter diet consists mainly of large ungulate carcasses, with very little plant matter. Rodent prey increases in importance during the spring, summer, and fall.
The coyote feeds on a variety of different produce, including blackberries, blueberries, peaches, pears, apples, prickly pears, chapotes, persimmons, peanuts, watermelons, cantaloupes, and carrots. During the winter and early spring, the coyote eats large quantities of grass, such as green wheat blades. It sometimes eats unusual items such as cotton cake, soybean meal, domestic animal droppings, beans, and cultivated grain such as maize, wheat, and sorghum.
In coastal California, coyotes now consume a higher percentage of marine-based food than their ancestors, which is thought to be due to the extirpation of the grizzly bear from this region. In Death Valley, coyotes may consume great quantities of hawkmoth caterpillars or beetles in the spring flowering months.


In areas where the ranges of coyotes and gray wolves overlap, interference competition and predation by wolves has been hypothesized to limit local coyote densities. Coyote ranges expanded during the 19th and 20th centuries following the extirpation of wolves, while coyotes were driven to extinction on Isle Royale after wolves colonized the island in the 1940s. One study conducted in Yellowstone National Park, where both species coexist, concluded that the coyote population in the Lamar River Valley declined by 39% following the reintroduction of wolves in the 1990s, while coyote populations in wolf inhabited areas of the Grand Teton National Park are 33% lower than in areas where they are absent. Wolves have been observed to not tolerate coyotes in their vicinity, though coyotes have been known to trail wolves to feed on their kills.
Coyotes may compete with cougars in some areas. In the eastern Sierra Nevada, coyotes compete with cougars over mule deer. Cougars normally outcompete and dominate coyotes, and may kill them occasionally, thus reducing coyote predation pressure on smaller carnivores such as foxes and bobcats. Coyotes that are killed are sometimes not eaten, perhaps indicating that these comprise competitive interspecies interactions, however there are multiple confirmed cases of cougars also eating coyotes. In northeastern Mexico, cougar predation on coyotes continues apace but coyotes were absent from the prey spectrum of sympatric jaguars, apparently due to differing habitat usages.
Other than by gray wolves and cougars, predation on adult coyotes is relatively rare but multiple other predators can be occasional threats. In some cases, adult coyotes have been preyed upon by both American black and grizzly bears, American alligators, large Canada lynx and golden eagles. At kill sites and carrion, coyotes, especially if working alone, tend to be dominated by wolves, cougars, bears, wolverines and, usually but not always, eagles (i.e., bald and golden). When such larger, more powerful and/or more aggressive predators such as these come to a shared feeding site, a coyote may either try to fight, wait until the other predator is done or occasionally share a kill, but if a major danger such as wolves or an adult cougar is present, the coyote will tend to flee.
Coyotes rarely kill healthy adult red foxes, and have been observed to feed or den alongside them, though they often kill foxes caught in traps. Coyotes may kill fox kits, but this is not a major source of mortality. In southern California, coyotes frequently kill gray foxes, and these smaller canids tend to avoid areas with high coyote densities.
In some areas, coyotes share their ranges with bobcats. These two similarly-sized species rarely physically confront one another, though bobcat populations tend to diminish in areas with high coyote densities. However, several studies have demonstrated interference competition between coyotes and bobcats, and in all cases coyotes dominated the interaction. Multiple researchers reported instances of coyotes killing bobcats, whereas bobcats killing coyotes is more rare. Coyotes attack bobcats using a bite-and-shake method similar to what is used on medium-sized prey. Coyotes (both single individuals and groups) have been known to occasionally kill bobcats – in most cases, the bobcats were relatively small specimens, such as adult females and juveniles. However, coyote attacks (by an unknown number of coyotes) on adult male bobcats have occurred. In California, coyote and bobcat populations are not negatively correlated across different habitat types, but predation by coyotes is an important source of mortality in bobcats. Biologist Stanley Paul Young noted that in his entire trapping career, he had never successfully saved a captured bobcat from being killed by coyotes, and wrote of two incidents wherein coyotes chased bobcats up trees. Coyotes have been documented to directly kill Canada lynx on occasion, and compete with them for prey, especially snowshoe hares. In some areas, including central Alberta, lynx are more abundant where coyotes are few, thus interactions with coyotes appears to influence lynx populations more than the availability of snowshoe hares.


Due to the coyote's wide range and abundance throughout North America, it is listed as Least Concern by the International Union for Conservation of Nature (IUCN). The coyote's pre-Columbian range was limited to the Southwest and Plains regions of North America, and northern and central Mexico. By the 19th century, the species expanded north and east, expanding further after 1900, coinciding with land conversion and the extirpation of wolves. By this time, its range encompassed the entire North American continent, including all of the contiguous United States and Mexico, southward into Central America, and northward into most of Canada and Alaska. This expansion is ongoing, and the species now occupies the majority of areas between 8°N (Panama) and 70°N (northern Alaska).
Although it was once widely believed that coyotes are recent immigrants to southern Mexico and Central America, aided in their expansion by deforestation, Pleistocene and Early Holocene records, as well as records from the pre-Columbian period and early European colonization show that the animal was present in the area long before modern times. Nevertheless, range expansion did occur south of Costa Rica during the late 1970s and northern Panama in the early 1980s, following the expansion of cattle-grazing lands into tropical rain forests. The coyote is predicted to appear in northern Belize in the near future, as the habitat there is favorable to the species. Concerns have been raised of a possible expansion into South America through the Panamanian Isthmus, should the Darién Gap ever be closed by the Pan-American Highway. This fear was partially confirmed in January 2013, when the species was recorded in eastern Panama's Chepo District, beyond the Panama Canal.
A 2017 genetic study proposes that coyotes were originally not found in the area of the eastern United States. From the 1890s, dense forests were transformed into agricultural land and wolf control implemented on a large scale, leaving a niche for coyotes to disperse into. There were two major dispersals from two populations of genetically distinct coyotes. The first major dispersal to the northeast came in the early 20th century from those coyotes living in the northern Great Plains. These came to New England via the northern Great Lakes region and southern Canada, and to Pennsylvania via the southern Great Lakes region, meeting together in the 1940s in New York and Pennsylvania. These coyotes have hybridized with the remnant gray wolf and eastern wolf populations, which has added to coyote genetic diversity and may have assisted adaptation to the new niche. The second major dispersal to the southeast came in the mid-20th century from Texas and reached the Carolinas in the 1980s. These coyotes have hybridized with the remnant red wolf populations before the 1970s when the red wolf was extirpated in the wild, which has also added to coyote genetic diversity and may have assisted adaptation to this new niche as well. Both of these two major coyote dispersals have experienced rapid population growth and are forecast to meet along the mid-Atlantic coast. The study concludes that for coyotes the long range dispersal, gene flow from local populations, and rapid population growth may be inter-related.


Among large North American carnivores, the coyote probably carries the largest number of diseases and parasites, likely due to its wide range and varied diet. Viral diseases known to infect coyotes include rabies, canine distemper, infectious canine hepatitis, four strains of equine encephalitis, and oral papillomatosis. By the late 1970s, serious rabies outbreaks in coyotes had ceased to be a problem for over 60 years, though sporadic cases every 1–5 years did occur. Distemper causes the deaths of many pups in the wild, though some specimens can survive infection. Tularemia, a bacterial disease, infects coyotes from tick bites and through their rodent and lagomorph prey, and can be deadly for pups.
Coyotes can be infected by both demodectic and sarcoptic mange, the latter being the most common. Mite infestations are rare and incidental in coyotes, while tick infestations are more common, with seasonal peaks depending on locality (May–August in the Northwest, March–November in Arkansas). Coyotes are only rarely infested with lice, while fleas infest coyotes from puphood, though they may be more a source of irritation than serious illness. Pulex simulans is the most common species to infest coyotes, while Ctenocephalides canis tends to occur only in places where coyotes and dogs (its primary host) inhabit the same area. Although coyotes are rarely host to flukes, they can nevertheless have serious effects on coyotes, particularly Nanophyetus salmincola, which can infect them with salmon poisoning disease, a disease with a 90% mortality rate. Trematode Metorchis conjunctus can also infect coyotes. Tapeworms have been recorded to infest 60–95% of all coyotes examined. The most common species to infest coyotes are Taenia pisiformis and Taenia crassiceps, which uses cottontail rabbits as intermediate hosts. The largest species known in coyotes is T. hydatigena, which enters coyotes through infected ungulates, and can grow to lengths of 80 to 400 cm (31 to 157 in). Although once largely limited to wolves, Echinococcus granulosus has expanded to coyotes since the latter began colonizing former wolf ranges. The most frequent ascaroid roundworm in coyotes is Toxascaris leonina, which dwells in the coyote's small intestine and has no ill effects, except for causing the host to eat more frequently. Hookworms of the genus Ancylostoma infest coyotes throughout their range, being particularly prevalent in humid areas. In areas of high moisture, such as coastal Texas, coyotes can carry up to 250 hookworms each. The blood-drinking A. caninum is particularly dangerous, as it damages the coyote through blood loss and lung congestion. A 10-day-old pup can die from being host to as few as 25 A. caninum worms.


Coyote features as a trickster figure and skin-walker in the folktales of some Native Americans, notably several nations in the Southwestern and Plains regions, where he alternately assumes the form of an actual coyote or that of a man. As with other trickster figures, Coyote acts as a picaresque hero who rebels against social convention through deception and humor. Folklorists such as Harris believe coyotes came to be seen as tricksters due to the animal's intelligence and adaptability. After the European colonization of the Americas, Anglo-American depictions of Coyote are of a cowardly and untrustworthy animal. Unlike the gray wolf, which has undergone a radical improvement of its public image, Anglo-American cultural attitudes towards the coyote remain largely negative.
In the Maidu creation story, Coyote introduces work, suffering, and death to the world. Zuni lore has Coyote bringing winter into the world by stealing light from the kachinas. The Chinook, Maidu, Pawnee, Tohono O'odham, and Ute portray the coyote as the companion of The Creator. A Tohono O'odham flood story has Coyote helping Montezuma survive a global deluge that destroys humanity. After The Creator creates humanity, Coyote and Montezuma teach people how to live. The Crow creation story portrays Old Man Coyote as The Creator. In The Dineh creation story, Coyote was present in the First World with First Man and First Woman, though a different version has it being created in the Fourth World. The Navajo Coyote brings death into the world, explaining that without death, too many people would exist, thus no room to plant corn.
Prior to the Spanish conquest of the Aztec Empire, Coyote played a significant role in Mesoamerican cosmology. The coyote symbolized military might in Classic era Teotihuacan, with warriors dressing up in coyote costumes to call upon its predatory power. The species continued to be linked to Central Mexican warrior cults in the centuries leading up to the post-Classic Aztec rule. In Aztec mythology, Huehuecóyotl (meaning "old coyote"), the god of dance, music and carnality, is depicted in several codices as a man with a coyote's head. He is sometimes depicted as a womanizer, responsible for bringing war into the world by seducing Xochiquetzal, the goddess of love. Epigrapher David H. Kelley argued that the god Quetzalcoatl owed its origins to pre-Aztec Uto-Aztecan mythological depictions of the coyote, which is portrayed as mankind's "Elder Brother", a creator, seducer, trickster, and culture hero linked to the morning star.


Coyote attacks on humans are uncommon and rarely cause serious injuries, due to the relatively small size of the coyote, but have been increasingly frequent, especially in California. There have been only two confirmed fatal attacks: one on a three-year-old named Kelly Keen in Glendale, California and another on a nineteen-year-old named Taylor Mitchell in Nova Scotia, Canada. In the 30 years leading up to March 2006, at least 160 attacks occurred in the United States, mostly in the Los Angeles County area. Data from United States Department of Agriculture (USDA) Wildlife Services, the California Department of Fish and Game, and other sources show that while 41 attacks occurred during the period of 1988–1997, 48 attacks were verified from 1998 through 2003. The majority of these incidents occurred in Southern California near the suburban-wildland interface.
In the absence of the harassment of coyotes practiced by rural people, urban coyotes are losing their fear of humans, which is further worsened by people intentionally or unintentionally feeding coyotes. In such situations, some coyotes have begun to act aggressively toward humans, chasing joggers and bicyclists, confronting people walking their dogs, and stalking small children. Non-rabid coyotes in these areas sometimes target small children, mostly under the age of 10, though some adults have been bitten.
Although media reports of such attacks generally identify the animals in question as simply "coyotes", research into the genetics of the eastern coyote indicates those involved in attacks in northeast North America, including Pennsylvania, New York, New England, and eastern Canada, may have actually been coywolves, hybrids of Canis latrans and C. lupus, not fully coyotes.


As of 2007, coyotes were the most abundant livestock predators in western North America, causing the majority of sheep, goat, and cattle losses. For example, according to the National Agricultural Statistics Service, coyotes were responsible for 60.5% of the 224,000 sheep deaths attributed to predation in 2004. The total number of sheep deaths in 2004 comprised 2.22% of the total sheep and lamb population in the United States, which, according to the National Agricultural Statistics Service USDA report, totaled 4.66 million and 7.80 million heads respectively as of July 1, 2005. Because coyote populations are typically many times greater and more widely distributed than those of wolves, coyotes cause more overall predation losses. United States government agents routinely shoot, poison, trap, and kill about 90,000 coyotes each year to protect livestock. An Idaho census taken in 2005 showed that individual coyotes were 5% as likely to attack livestock as individual wolves. In Utah, more than 11,000 coyotes were killed for bounties totaling over $500,000 in the fiscal year ending June 30, 2017.
Livestock guardian dogs are commonly used to aggressively repel predators and have worked well in both fenced pasture and range operations. A 1986 survey of sheep producers in the USA found that 82% reported the use of dogs represented an economic asset.
Re-wilding cattle, which involves increasing the natural protective tendencies of cattle, is a method for controlling coyotes discussed by Temple Grandin of Colorado State University. This method is gaining popularity among producers who allow their herds to calve on the range and whose cattle graze open pastures throughout the year.
Coyotes typically bite the throat just behind the jaw and below the ear when attacking adult sheep or goats, with death commonly resulting from suffocation. Blood loss is usually a secondary cause of death. Calves and heavily fleeced sheep are killed by attacking the flanks or hindquarters, causing shock and blood loss. When attacking smaller prey, such as young lambs, the kill is made by biting the skull and spinal regions, causing massive tissue and bone damage. Small or young prey may be completely carried off, leaving only blood as evidence of a kill. Coyotes usually leave the hide and most of the skeleton of larger animals relatively intact, unless food is scarce, in which case they may leave only the largest bones. Scattered bits of wool, skin, and other parts are characteristic where coyotes feed extensively on larger carcasses.
Tracks are an important factor in distinguishing coyote from dog predation. Coyote tracks tend to be more oval-shaped and compact than those of domestic dogs, and their claw marks are less prominent and the tracks tend to follow a straight line more closely than those of dogs. With the exception of sighthounds, most dogs of similar weight to coyotes have a slightly shorter stride. Coyote kills can be distinguished from wolf kills by less damage to the underlying tissues in the former. Also, coyote scat tends to be smaller than wolf scat.
Coyotes are often attracted to dog food and animals that are small enough to appear as prey. Items such as garbage, pet food, and sometimes feeding stations for birds and squirrels attract coyotes into backyards. About three to five pets attacked by coyotes are brought into the Animal Urgent Care hospital of South Orange County (California) each week, the majority of which are dogs, since cats typically do not survive the attacks. Scat analysis collected near Claremont, California, revealed that coyotes relied heavily on pets as a food source in winter and spring. At one location in Southern California, coyotes began relying on a colony of feral cats as a food source. Over time, the coyotes killed most of the cats and then continued to eat the cat food placed daily at the colony site by people who were maintaining the cat colony.
Coyotes usually attack smaller-sized dogs, but they have been known to attack even large, powerful breeds such as the Rottweiler in exceptional cases. Dogs larger than coyotes, such as greyhounds, are generally able to drive them off and have been known to kill coyotes. Smaller breeds are more likely to suffer injury or death.


Coyote hunting is one of the most common forms of predator hunting that humans partake in. There are not many regulations with regard to the taking of the coyote which means there are many different methods that can be used to hunt the animal. The most common forms are trapping, calling, and hound hunting. Since coyotes are colorblind, seeing only in shades of gray and subtle blues, open camouflages, and plain patterns can be used. The average male coyote weighs 8 to 20 kg (18 to 44 lbs) and the average female coyote 7 to 18 kg (15 to 40 lbs). A universal projectile that can perform between those weights is the .223 Remington so that the projectile expands in the target after the entry but before the exit thus delivering the most energy. Coyotes being the light and agile animals they are, they often leave a very light impression on terrain. The coyote's footprint is oblong, approximately 6.35 cm (2.5-inches) long and 5.08 cm (2-inches) wide. There are 4 claws in both their front and hind paws. The coyote's center pad is relatively shaped like that of a rounded triangle. Like the domestic dog the coyote's front paw is slightly larger than the hind paw. The coyote's paw is most similar to that of the domestic dog.


Prior to the mid-19th century, coyote fur was considered worthless. This changed with the diminution of beavers, and by 1860, the hunting of coyotes for their fur became a great source of income (75 cents to $1.50 per skin) for wolfers in the Great Plains. Coyote pelts were of significant economic importance during the early 1950s, ranging in price from $5 to $25 per pelt, depending on locality. The coyote's fur is not durable enough to make rugs, but can be used for coats and jackets, scarves, or muffs. The majority of pelts are used for making trimmings, such as coat collars and sleeves for women's clothing. Coyote fur is sometimes dyed black as imitation silver fox.
Coyotes were occasionally eaten by trappers and mountain men during the western expansion. Coyotes sometimes featured in the feasts of the Plains Indians, and coyote pups were eaten by the indigenous people of San Gabriel, California. The taste of coyote meat has been likened to that of the wolf and is more tender than pork when boiled. Coyote fat, when taken in the fall, has been used on occasion to grease leather or eaten as a spread.


Coyotes were likely semidomesticated by various pre-Columbian cultures. Some 19th-century writers wrote of coyotes being kept in native villages in the Great Plains. The coyote is easily tamed as a pup, but can become destructive as an adult. Both full-blooded and hybrid coyotes can be playful and confiding with their owners, but are suspicious and shy of strangers, though coyotes being tractable enough to be used for practical purposes like retrieving and pointing have been recorded. A tame coyote named "Butch", caught in the summer of 1945, had a short-lived career in cinema, appearing in Smoky and Ramrod before being shot while raiding a henhouse.


The black-backed jackal (Lupulella mesomelas) is a medium-sized canine native to eastern and southern Africa. These regions are separated by roughly 900 kilometers.
One region includes the southernmost tip of the continent, including South Africa, Namibia, Botswana, and Zimbabwe. The other area is along the eastern coastline, including Kenya, Somalia, Djibouti, Eritrea, and Ethiopia. It is listed by the IUCN as least concern, due to its widespread range and adaptability, although it is still persecuted as a livestock predator and rabies vector.
Compared to other members of the genus Canis, the black-backed jackal is a very ancient species, and has changed little since the Pleistocene, being the most basal wolf-like canine, alongside the closely related side-striped jackal. It is a fox-like animal with a reddish brown to tan coat and a black saddle that extends from the shoulders to the base of the tail. It is a monogamous animal, whose young may remain with the family to help raise new generations of pups. The black-backed jackal has a wide array of food sources, feeding on small to medium-sized animals, as well as plant matter and human refuse.


Johann Christian Daniel von Schreber named Canis mesomelas in 1775. It was later proposed as the genus Lupulella Hilzheimer 1906.
The black-backed jackal has occupied eastern and southern Africa for at least 2–3 million years, as shown by fossil deposits in Kenya, Tanzania, and South Africa. Specimens from fossil sites in Transvaal are almost identical to their modern counterparts, but have slightly different nasal bones. As no fossils have been found north of Ethiopia, the species likely has always been sub-Saharan in distribution. The black-backed jackal is relatively unspecialised, and can thrive in a wide variety of habitats, including deserts, as its kidneys are well adapted for water deprivation. It is, however, more adapted to a carnivorous diet than the other jackals, as shown by its well-developed carnassial shear and the longer cutting blade of the premolars.
Juliet Clutton-Brock classed the black-backed jackal as being closely related to the side-striped jackal, based on cranial and dental characters. Studies on allozyme divergence within the Canidae indicate that the black-backed jackal and other members of the genus Canis are separated by a considerable degree of genetic distance. Further studies show a large difference in mitochondrial DNA sequences between black-backed jackals and other sympatric "jackal" species, consistent with divergence 2.3–4.5 million years ago.
A mitochondrial DNA (mDNA) sequence alignment for the wolf-like canids gave a phylogenetic tree with the side-striped jackal and the black-backed jackal being the most basal members of this clade, which means that this tree is indicating an African origin for the clade.
Because of this deep divergence between the black-backed jackal and the rest of the "wolf-like" canids, one author has proposed to change the species' generic name from Canis to Lupulella.
In 2017, jackal relationships were further explored, with an mDNA study finding that the two black-backed jackal subspecies had diverged from each other 1.4 million years ago to form the central African and east African populations. The study proposes that due to this long separation, which is longer than the separation of the African golden wolf from the wolf lineage, that the two subspecies might warrant separate species status.
In 2019, a workshop hosted by the IUCN/SSC Canid Specialist Group recommends that because DNA evidence shows the side-striped jackal (Canis adustus) and black-backed jackal (Canis mesomelas) to form a monophyletic lineage that sits outside of the Canis/Cuon/Lycaon clade, that they should be placed in a distinct genus, Lupulella Hilzheimer, 1906 with the names Lupulella adusta and Lupulella mesomelas.
The phylogenetic tree for the wolf-like canids may give conflicting positions for the black-backed jackal and the side-striped jackal relative to the genus Canis members depending on whether the genetic markers were based on mitochondrial DNA or nuclear DNA (from the cell's nucleus).  The explanation proposed is that mitochondrial DNA introgression occurred from an ancient ancestor of Canis into the lineage that led to the black-backed jackal around 6.2–5.2 million years ago.


The black-backed jackal is a fox-like canid with a slender body, long legs, and large ears. It is similar to the closely related side-striped jackal and more distantly related to the golden jackal, though its skull and dentition are more robust and the incisors much sharper. It weighs 6–13 kg (13–29 lb), stands 38–48 cm (15–19 in) at the shoulder, and measures 67.3–81.2 cm (26+1⁄2–32 in) in body length.
The base colour is reddish brown to tan, which is particularly pronounced on the flanks and legs. A black saddle intermixed with silvery hair extends from the shoulders to the base of the tail. A long, black stripe extending along the flanks separates the saddle from the rest of the body, and can be used to differentiate individuals. The tail is bushy and tipped with black. The lips, throat, chest, and inner surface of the limbs are white. The winter coat is a much deeper reddish brown. Albino specimens occasionally occur. The hair of the face measures 10–15 mm in length, and lengthens to 30–40 mm on the rump. The guard hairs of the back are 60 mm on the shoulder, decreasing to 40 mm at the base of the tail. The hairs of the tail are the longest, measuring 70 mm in length.


The black-backed jackal is a monogamous and territorial animal, whose social organisation greatly resembles that of the golden jackal. However, the assistance of elder offspring in helping raise the pups of their parents has a greater bearing on pup survival rates than in the latter species. The basic social unit is a monogamous mated pair which defends its territory through laying faeces and urine on range boundaries. Scent marking is usually done in tandem, and the pair aggressively expels intruders. Such encounters are normally prevented, as the pair vocalises to advertise its presence in a given area. It is a highly vocal species, particularly in Southern Africa. Sounds made by the species include yelling, yelping, woofing, whining, growling, and cackling. It communicates with group members and advertises its presence by a high-pitched, whining howl, and expresses alarm through an explosive cry followed by shorter, high-pitched yelps. This sound is particularly frantic when mobbing a leopard. In areas where the black-backed jackal is sympatric with the African golden wolf, the species does not howl, instead relying more on yelps. In contrast, black-backed jackals in Southern Africa howl much like golden jackals. When trapped, it cackles like a fox.


The mating season takes place from late May to August, with a gestation period of 60 days. Pups are born from July to October. Summer births are thought to be timed to coincide with population peaks of vlei rats and four-striped grass mice, while winter births are timed for ungulate calving seasons. Litters consist of one to 9 pups, which are born blind. For the first three weeks of their lives, the pups are kept under constant surveillance by their dam, while the sire and elder offspring provide food. The pups open their eyes after 8–10 days and emerge from the den at the age of 3 weeks. They are weaned at 8–9 weeks, and can hunt by themselves at the age of 6 months. Sexual maturity is attained at 11 months, though few black-backed jackals reproduce in their first year. Unlike golden jackals, which have comparatively amicable intrapack relationships, black-backed jackal pups become increasingly quarrelsome as they age, and establish more rigid dominance hierarchies. Dominant pups appropriate food, and become independent at an earlier age. The grown pups may disperse at one year of age, though some remain in their natal territories to assist their parents in raising the next generation of pups. The average lifespan in the wild is 7 years, though captive specimens can live twice as long.


Black-backed jackals are omnivores, which feed on invertebrates, such as beetles, grasshoppers, crickets, termites, millipedes, spiders, and scorpions. They also feed on mammals, such as rodents, hares, and young antelopes up to the size of topi calves. They also feed on carrion, birds, lizards and snakes. A pair of black-backed jackals in the Kalahari desert was observed to kill a kori bustard, and on a separate occasion, a black mamba by prolonged harassment of the snake and crushing of the snake's head. Black-backed jackals occasionally feed on fruits and berries. It also feeds on eggs of birds. In coastal areas, they feed on beached marine mammals, seals, fish, and mussels. A single jackal is capable of killing a healthy adult impala. Adult dik-diks and Thomson's gazelles seem to be the upper limit of their killing capacity, though they target larger species if those are sick, with one pair having been observed to harass a crippled bull rhinoceros. They typically kill tall prey by biting at the legs and loins, and frequently go for the throat. In Serengeti woodlands, they feed heavily on African grass rats. In East Africa, during the dry season, they hunt the young of gazelles, impalas, topi, tsessebe, and warthogs. In South Africa, black-backed jackals frequently prey on antelopes (primarily impala and springbok and occasionally duiker, reedbuck, and steenbok), carrion, hares, hoofed livestock, insects, and rodents. They also prey on small carnivores, such as mongooses, polecats, and wildcats. On the coastline of the Namib Desert, jackals feed primarily on marine birds (mainly Cape and white-breasted cormorants and jackass penguins), marine mammals (including Cape fur seals), fish, and insects. Like most canids, the black-backed jackal caches surplus food.


They tend to avoid humans. In areas where the black-backed jackal is sympatric with the larger side-striped jackal, the former species aggressively drives out the latter from grassland habitats into woodlands. This is unique among carnivores, as larger species commonly displace smaller ones. Black-backed jackal pups are vulnerable to African golden wolves, ratels, and spotted and brown hyenas. Adults have few natural predators, save for leopards and African wild dogs, Though there are some reports that martial eagles prey on both juveniles and adults.


Black-backed jackals occasionally hunt domestic animals, including dogs, cats, pigs, goats, sheep, and poultry, with sheep tending to predominate. They rarely target cattle, though cows giving birth may be attacked. Jackals can be a serious problem for sheep farmers, particularly during the lambing season. Sheep losses to black-backed jackals in a 440 km² study area in KwaZulu-Natal consisted of 0.05% of the sheep population. Of 395 sheep killed in a sheep farming area in KwaZulu-Natal, 13% were killed by jackals. Jackals usually kill sheep with a throat bite, and begin feeding by opening the flank and consuming the flesh and skin of the flank, heart, liver, some ribs, haunch of hind leg, and sometimes the stomach and its contents. In older lambs, the main portions eaten are usually heart and liver. Usually, only one lamb per night is killed in any one place, but sometimes two and occasionally three may be killed. The oral history of the Khoikhoi indicates they have been a nuisance to pastoralists long before European settlement. South Africa has been using fencing systems to protect sheep from jackals since the 1890s, though such measures have mixed success, as the best fencing is expensive, and jackals can easily infiltrate cheap wire fences.


Due to livestock losses to jackals, many hunting clubs were opened in South Africa in the 1850s. Black-backed jackals have never been successfully eradicated in hunting areas, despite strenuous attempts to do so with dogs, poison, and gas. Black-backed jackal coursing was first introduced to the Cape Colony in the 1820s by Lord Charles Somerset, who as an avid fox hunter, sought a more effective method of managing jackal populations, as shooting proved ineffective. Coursing jackals also became a popular pastime in the Boer Republics. In the western Cape in the early 20th century, dogs bred by crossing foxhounds, lurchers, and borzoi were used.
Spring traps with metal jaws were also effective, though poisoning by strychnine became more common by the late 19th century. Strychnine poisoning was initially problematic, as the solution had a bitter taste, and could only work if swallowed. Consequently, many jackals learned to regurgitate poisoned baits, thus inciting wildlife managers to use the less detectable crystal strychnine rather than liquid. The poison was usually placed within sheep carcasses or in balls of fat, with great care being taken to avoid leaving any human scent on them. Black-backed jackals were not a popular quarry in the 19th century, and are rarely mentioned in hunter's literature. By the turn of the century, jackals became increasingly popular quarry as they encroached upon human habitations after sheep farming and veld burning diminished their natural food sources. Although poisoning had been effective in the late 19th century, its success rate in eliminating jackals waned in the 20th century, as jackals seemed to be learning to distinguish poisoned foods.
The Tswana people often made hats and cloaks out of black-backed jackal skins. Between 1914 and 1917, 282,134 jackal pelts (nearly 50,000 a year) were produced in South Africa. Demand for pelts grew during the First World War, and were primarily sold in Cape Town and Port Elizabeth. Jackals in their winter fur were in great demand, though animals killed by poison were less valued, as their fur would shed.


Canna indica, commonly known as Indian shot, African arrowroot, edible canna, purple arrowroot, Sierra Leone arrowroot, is a plant species in the family Cannaceae. It is native to much of South America, Central America, the West Indies, and Mexico. It is also naturalized in the southeastern United States (Florida, Texas, Louisiana, and South Carolina), and much of Europe, sub-Saharan Africa, Southeast Asia, and Oceania. Canna indica (achira in Hispanic America, cana-da-índia in Brazil) has been a minor food crop cultivated by indigenous peoples of the Americas for thousands of years.


Canna indica is native to South America: Colombia, Venezuela, Ecuador, Peru, Brazil, Uruguay and Argentina as well as the West Indies and Central America.
In modern times, C. indica is reportedly naturalized in Austria, Portugal, Spain, Azores, Canary Islands, Cape Verde, Madeira, most of tropical Africa, Ascension Island, St. Helena, Madagascar, China, Japan, Taiwan, the Bonin Islands, India, Nepal, Sri Lanka, Cambodia, Laos, Thailand, Vietnam, Burma, Java, Malaysia, the Philippines, Christmas Island, the Bismarck Archipelago, Norfolk Island, New South Wales, Queensland, Fiji, Tonga, Vanuatu, Kiribati, the Cook Islands, the Society Islands, the Caroline Islands and Hawaii.
Canna can be cultivated from sea level to 2,700 m (8,900 ft) above sea level, but thrives in temperate, tropical or subtropical mountain climates, between 1,000 and 2,000 m (3,300 and 6,600 ft) above sea level (in humid tropical climates for higher elevations) at a mean temperature of 14 to 27 °C. The plant prefers a mean annual rainfall between 1,000–4,500 millimetres (3+1⁄2–15 ft), but it can tolerate 500–5,000 mm per year. Canna prefers light sandy-loamy soils, but can also grow on heavy soils, as far as they are not wet. It is indifferent to soil pH. For seeds to germinate, they must soak in water for two to three days.


To cultivate Canna indica, the substrate should be rich, humiferous, and light. The optimal substrate consists of a deep, rich and well-drained soil in a sunny place with a pH between 5.5–7.5. The rhizomes should be planted at 10 cm deep, after the last frost. Although it can tolerate dry periods, occasional irrigation will be beneficial for the yield. C. indica is a plant that can withstand low temperatures (down to -10 °C in regions with a mild climate). Nevertheless, the foliage can already be affected at 0 °C.


The yield varies depending on the region of cultivation and its climate and soil conditions. In certain locations, the yield of Canna indica can be higher than other starchy crops like cassava and arrowroot. The average rhizome yield is believed to be around 22–50 tons per hectare, whereas the starch yield is about 2–5 tons per hectare and can reach up to 10 tons per ha. Observations show that the highest yield in rhizomes does not necessarily correspond to the highest yield in starch.


The plant is native to the higher elevation deserts of the Southwestern United States in Arizona and southeastern California, and of Northwest Mexico in isolated locales in Baja California and Sonora).
Canotia grows in desert scrub environments, primarily on slopes or in washes. It is found in the higher Sonoran Desert across central Arizona, reaching to the Mojave Desert of California and northwest Arizona, and into the lower reaches of the Grand Canyon.


The caper bush has been introduced as a specialized culture in some European countries in the last four decades. The economic importance of the caper plant led to a significant increase in both the area under cultivation and production levels during the late 1980s. The main production areas are in harsh environments found in Iraq, Morocco, the southeastern Iberian Peninsula, Turkey, the Greek island of Santorini, and the Italian island of Pantelleria, and the Aeolian Islands, especially Salina. Capers from Pantelleria and Aelian island are recognized as European PGI products. This species has developed special mechanisms to survive in the Mediterranean conditions, and introduction in semiarid lands may help to prevent the disruption of the equilibrium of those fragile ecosystems.
A harvest duration of at least three months is necessary for profitability. Intense daylight and a long growing period are necessary to secure high yields. The caper bush can withstand temperatures over 40 °C in summer, but it is sensitive to frost during its vegetative period. A caper bush is able to survive low temperatures in the form of stump, as happens in the foothills of the Alps. Caper plants are found even 3,500 m above sea level in Ladakh, though they are usually grown at lower altitudes. Some Italian and Argentine plantings can withstand strong winds without problems, due to the plant's decumbent architecture and the coriaceous consistency of the leaves in some populations.
The known distributions of each species can be used to identify the origin of commercially prepared capers.
The caper bush is a rupicolous species. It is widespread on rocky areas and is grown on different soil associations, including alfisols, regosols, and lithosols. In different Himalayan locations, C. spinosa tolerates both silty clay and sandy, rocky, or gravelly surface soils, with less than 1% organic matter. It grows on bare rocks, crevices, cracks, and sand dunes in Pakistan, in dry calcareous escarpments of the Adriatic region, in dry coastal ecosystems of Egypt, Libya, and Tunisia, in transitional zones between the littoral salt marsh and the coastal deserts of the Asian Red Sea coast, in the rocky arid bottoms of the Jordan valley, in calcareous sandstone cliffs at Ramat Aviv, Israel, and in central west and northwest coastal dunes of Australia. It grows spontaneously in wall joints of antique Roman fortresses, on the Western Wall of Jerusalem's Temple Mount, and on the ramparts of the castle of Santa Bárbara (Alicante, Spain). Clinging caper plants are dominant on the medieval limestone-made ramparts of Alcudia and the bastions of Palma (Majorca, Spain). This aggressive pioneering has brought about serious problems for the protection of monuments.


The Alpine ibex (Capra ibex), also known as the steinbock, bouquetin, or simply ibex, is a species of wild goat that lives in the mountains of the European Alps. It is a sexually dimorphic species: males are larger and carry longer, curved horns than females. Its coat colour is typically brownish grey. Alpine ibex tend to live in steep, rough terrain near the snow line. They are also social, although adult males and females segregate for most of the year, coming together only to mate. Four distinct groups exist; adult male groups, female-offspring groups, groups of young individuals, and mixed-sex groups.
During the breeding season, males fight for access to females, and use their long horns in agonistic behaviour. After being extirpated from most areas by the 19th century, the Alpine ibex was successfully reintroduced to parts of its historical range. All individuals living today descend from the stock in Gran Paradiso National Park in Aosta Valley and Piemonte (Italy), a national park created to help the ibex thrive. The ibex is the emblem of both the Gran Paradiso National Park, and the contiguous Vanoise National Park across the French border. The species is currently listed as of least concern by the IUCN, but went through a population bottleneck of fewer than 100 individuals during its near-extinction event. This has led to very low genetic diversity across populations.


The Alpine ibex was first described by Carl Linnaeus in 1758. It is classified in the genus Capra (Latin for "goat") with at least seven other species of wild goats. Both Capra and Ovis (sheep) descended from a goral-like animal from the Miocene and early Pliocene, whose fossils are found in Kenya, China, and Slovenia. The genus Tossunnoria appears in China during the late Miocene and appears to have been intermediate between gorals and goats. Fossils of Alpine ibex date back to the late Pleistocene, when  the Spanish ibex and it probably evolved from the extinct Pleistocene species Capra camburgensis. The Nubian (C. nubiana), walia (C. walie), and Siberian ibexes (C. sibirica) were previously considered to be subspecies of the Alpine ibex, giving populations in the Alps the trinomial of C. i. ibex.


The Alpine ibex was, at one point, restricted only to the Gran Paradiso National Park in northern Italy, and the contiguous Maurienne Valley in the French Alps but it was reintroduced to most of the Alps. Reintroductions started in 1906 into Switzerland. Alpine ibex are now found in most or all of the Italian and French alpine ranges, southern Germany, Switzerland and Austria. It has also been introduced to Bulgaria and Slovenia.
An excellent climber, its preferred habitat is the rocky region along the snow line above alpine forests, where it occupies steep, rough terrain at elevations of 1,800 to 3,300 m (5,900 to 10,800 ft). Alpine ibex are typically absent from woodland areas, although adult males in densely populated areas may stay in larch and mixed larch-spruce woodland if no snow has fallen. Males spend the winter in coniferous forests. For most of the year, males and females occupy different habitats. Females rely on steep terrain more so than males. Males use lowland meadows during the spring, which is when snow melts and green grass appears. They then climb to alpine meadows during the summer. When winter arrives, both sexes move to steep, rocky slopes that amass little snow. They prefer slopes of 30–45° and use small caves and overhangs for shelter. Home ranges are highly variable, depending on the availability of resources, and vary in size throughout the year. Figures from 180 to 2,800 ha (0.69 to 10.81 sq mi) have been recorded. Home ranges tend to be largest during summer and autumn, smallest in winter, and intermediate in spring. Female home ranges are usually smaller than those of males. Alpine ibex appear to have a low rate of predation and in Gran Paradiso typically die of age, starvation, or disease.


Alpine ibex are strictly herbivorous, with over half of their diet consisting of grasses, and the remainder being a mixture of mosses, flowers, leaves, and twigs. If leaves and shoots are out of reach, they often stand on their rear legs to reach this food. Grass genera that are the most commonly eaten are Agrostis, Avena, Calamagrostis, Festuca, Phleum, Poa, Sesleria, and Trisetum. The climbing ability of the Alpine ibex is such that it has been observed standing on the sheer face of the Cingino Dam in Piedmont, Italy, where it licks the stonework to obtain mineral salts.


Although the Alpine ibex is a social species, they segregate sexually and spatially depending on the season. Four types of groups exist: Adult male groups, female-offspring groups, groups of young individuals 2–3 years old, and mixed-sex groups. Young groups are numerous at the beginning of summer, but are expelled by females at the end of their gestation period. Female and offspring groups occur year-round, at least in an area of the French Alps. Mixed-sex groups of adult males and females occur during breeding, which lasts from December to January. By April and May, the adults separate. The largest aggregations of either sex occur during June and July. Gatherings of males begin to decrease during October and November, and are lowest from the rut from December to March. The males then leave their separate wintering areas and gather again.
A linear dominance hierarchy exists among males. In small populations, which are more cohesive, males know their place in the hierarchy based on memories of past encounters, while in mobile and large groups, where encounters with strangers are common, rank is based on horn size. Antagonistic behavior in males can come in the form of "direct" or "indirect" aggression. With direct aggression, one male bumps another with his horns or places himself in front of his opponent. He stands on his hind legs and comes down on his opponent with his horns. This may signal that he is ready to clash or may be attempting a real clash.  Indirect aggression is mostly intimidation displays.


The Alpine ibex historically ranged through France, Italy, Switzerland, Liechtenstein, Bavaria, Austria and Slovenia. Starting in the early 16th century and with firearms becoming common, the overall population declined due to overexploitation and poaching. The ibex became extirpated in Switzerland and Germany by the 18th century, and was extirpated in Austria and northeastern Italy by the 19th century. They remained only in and around the adjacent Gran Paradiso and Vanoise Massifs, then both part of the Kingdom of Sardinia. Located in the western Italian Alps and the Maurienne valley in the north eastern French alps, bordering the Vanoise and Gran Paradiso Massif, the park was declared a royal hunting reserve in 1854 with the name of "Royal hunting reserve of Gran Paradiso" by Victor Emmanuel II, the first king of a united Italy.
Ibex were protected from poaching and their numbers increased, reaching 3,020 in 1914. The ibex enjoyed further protection when Gran Paradiso was made into a national park in 1922. Animals from this population naturally dispersed into surrounding regions. However, reintroductions have been the predominant source of new populations. Today, the total population of Alpine ibex is over 30,000 and is considered to be of Least Concern by the IUCN. However, Alpine ibex have low genetic diversity putting them at risk of inbreeding depression.


The European nightjar (Caprimulgus europaeus), common goatsucker, Eurasian nightjar or just nightjar, is a crepuscular and nocturnal bird in the nightjar family that breeds across most of Europe and the Palearctic to Mongolia and Northwestern China. The Latin generic name refers to the old myth that the nocturnal nightjar suckled from goats, causing them to cease to give milk. The six subspecies differ clinally, the birds becoming smaller and paler towards the east of the range. All populations are migratory, wintering in sub-Saharan Africa. Their densely patterned grey and brown plumage makes individuals difficult to see in the daytime when they rest on the ground or perch motionless along a branch, although the male shows white patches in the wings and tail as he flies at night.
The preferred habitat is dry, open country with some trees and small bushes, such as heaths, forest clearings or newly planted woodland. The male European nightjar occupies a territory in spring and advertises his presence with a distinctive sustained churring trill from a perch. He patrols his territory with wings held in a V and tail fanned, chasing intruders while wing-clapping and calling. Wing clapping also occurs when the male chases the female in a spiralling display flight. The European nightjar does not build a nest, and its two grey and brown blotched eggs are laid directly on the ground; they hatch after about 17–21 days and the downy chicks fledge in another 16–17 days.
The European nightjar feeds on a wide variety of flying insects, which it seizes in flight, often fly-catching from a perch. It hunts by sight, silhouetting its prey against the night sky. Its eyes are relatively large, each with a reflective layer, which improves night vision. It appears not to rely on its hearing to find insects and does not echolocate. Drinking and bathing take place during flight. Although it suffers a degree of predation and parasitism, the main threats to the species are habitat loss, disturbance and a reduction of its insect prey through pesticide use. Despite population decreases, its large numbers and huge breeding range mean that it is classified by the International Union for Conservation of Nature as being of least concern.


There are six recognised subspecies, although the differences are mainly clinal; birds become smaller and paler in the east of the range and the males have larger white wing spots. Birds of intermediate appearance occur where the subspecies' ranges overlap.
The fossil record is deficient, but it is likely that these poorly defined subspecies diverged as global temperature rose over the last 10,000 years or so. Only one record of the species possibly dates back to the late Eocene.


The European nightjar is 24.5–28 cm (9.6–11.0 in) long, with a 52–59 cm (20–23 in) wingspan. The male weighs 51–101 g (1.8–3.6 oz) and the female 67–95 g (2.4–3.4 oz). The adult of the nominate subspecies has greyish-brown upperparts with dark streaking, a pale buff hindneck collar and a white moustachial line. The closed wing is grey with buff spotting, and the underparts are greyish-brown, with brown barring and buff spots. The bill is blackish, the iris is dark brown and the legs and feet are brown.
The flight on long pointed wings is noiseless, due to their soft plumage, and very buoyant. Flying birds can be sexed since the male has a white wing patch across three primary feathers and white tips to the two outer tail feathers, whereas females do not show any white in flight. Chicks have downy brown and buff plumage, and the fledged young are similar in appearance to the adult female. Adults moult their body feathers from June onwards after breeding, suspend the process while migrating, and replace the tail and flight feathers on the wintering grounds. Moult is completed between January and March. Immature birds follow a similar moult strategy to the adults unless they are from late broods, in which case the entire moult may take place in Africa.
Other nightjar species occur in parts of the breeding and wintering ranges. The red-necked nightjar breeds in Iberia and northwest Africa; it is larger, greyer and longer winged than the European nightjar, and has a broad buff collar and more conspicuous white markings on the wings and tail. Wintering European nightjars in Africa may overlap with the related rufous-cheeked and sombre nightjars. Both have a more prominent buff hind-neck collar and more spotting on the wing coverts. The sombre nightjar is also much darker than its European cousin. Given their nocturnal habits, cryptic plumage and difficulty of observation, nightjar observation "is as much a matter of fortune as effort or knowledge".


The male European nightjar's song is a sustained churring trill, given continuously for up to 10 minutes with occasional shifts of speed or pitch. It is delivered from a perch, and the male may move around its territory using different song posts. Singing is more frequent at dawn and dusk than during the night, and is reduced in poor weather. The song may end with a bubbling trill and wing-clapping, perhaps indicating the approach of a female. Migrating or wintering birds sometimes sing. Individual male nightjars can be identified by analysing the rate and length of the pulses in their songs. Even a singing male may be hard to locate; the perched bird is difficult to spot in low light conditions, and the song has a ventriloquial quality as the singer turns his head. The song is easily audible at 200 m (660 ft), and can be heard at 600 m (2,000 ft) in good conditions; it can be confused with the very similar sound of the European mole cricket.
The female does not sing, but when on the wing, both sexes give a short cuick, cuick call, also used when chasing predators. Other calls include variations on a sharp chuck when alarmed, hisses given by adults when handled or chicks when disturbed, and an assortment of wuk, wuk, wuk, muffled oak, oak and murmurs given at the nest. Large young have a threat display with the mouth opened wide while hissing loudly.
Despite the name, wing-clapping does not involve contact between the two wing tips over the bird's back as was once thought. The sound is produced in a whiplash-like way as each wing cracks down.


The breeding range of the European nightjar comprises Europe north to around latitude 64°N and Asia north to about 60°N and east to Lake Baikal and eastern Mongolia. The southern limits are northwestern Africa, Iraq, Iran and the northwestern Himalayas. This nightjar formerly bred in Syria and Lebanon.
All populations are migratory, and most birds winter in Africa south of the Sahara, with just a few records from Pakistan, Morocco and Israel. Migration is mainly at night, singly or in loose groups of up to twenty birds. European breeders cross the Mediterranean and North Africa, whereas eastern populations move through the Middle East and East Africa. Some Asian birds may therefore cross 100° of longitude on their travels. Most birds start their migration at the time of a full moon.
Most birds winter in eastern or southeastern Africa, although individuals of the nominate race have been recently discovered wintering in the Democratic Republic of the Congo; records elsewhere in West Africa may be wintering birds of this subspecies or C. e. meridionalis. Most autumn migration takes place from August to September, and the birds return to the breeding grounds by May. Recent tracking data has revealed that European nightjars have a loop migration from Western Europe to sub-equatorial Africa where they have to cross several ecological barriers (the Mediterranean Sea, the Sahara and the Central African Tropical Rainforest). Individuals use similar stop-over sites as do other European migrants. Vagrants have occurred in Iceland, the Faroe Islands, the Seychelles, the Azores, Madeira and the Canary Islands.
The European nightjar is a bird of dry, open country with some trees and small bushes, such as heaths, commons, moorland, forest clearings or felled or newly planted woodland. When breeding, it avoids treeless or heavily wooded areas, cities, mountains, and farmland, but it often feeds over wetlands, cultivation or gardens. In winter it uses a wider range of open habitats including acacia steppe, sandy country and highlands. It has been recorded at altitudes of 2,800 m (9,200 ft) on the breeding grounds and 5,000 m (16,000 ft) in the wintering areas.


The European nightjar is crepuscular and nocturnal. During the day it rests on the ground, often in a partly shaded location, or perches motionless lengthwise along an open branch or a similar low perch. The cryptic plumage makes it difficult to see in the daytime, and birds on the ground, if they are not already in shade, will turn occasionally to face the sun thereby minimising their shadow. If it feels threatened, the nightjar flattens itself to the ground with eyes almost closed, flying only when the intruder is 2–5 m (7–16 ft) away. It may call or wing clap as it goes, and land as far as 40 m (130 ft) from where it was flushed. In the wintering area it often roosts on the ground but also uses tree branches up to 20 m (66 ft) high. Roost sites at both the breeding and wintering grounds are used regularly if they are undisturbed, sometimes for weeks at a time.
Like other nightjars, it will sit on roads or paths during the night and hover to investigate large intruders such as deer or humans. It may be mobbed by birds while there is still light, and by bats, other nightjar species or Eurasian woodcocks during the night. Owls and other predators such as red foxes will be mobbed by both male and female European nightjars. Like other aerial birds, such as swifts and swallows, nightjars make a quick plunge into water to wash. They have a unique serrated comb-like structure on the middle claw, which is used to preen and perhaps to remove parasites.
In cold or inclement weather, several nightjar species can slow their metabolism and go into torpor, notably the common poorwill, which will maintain that state for weeks. The European nightjar has been observed in captivity to be able to maintain a state of torpor for at least eight days without harm, but the relevance of this to wild birds is unknown.


Breeding is normally from late May to August, but may be significantly earlier in northwest Africa or western Pakistan. Returning males arrive about two weeks before the females and establish territories which they patrol with wings held in a V-shape and tail fanned, chasing intruders while wing-clapping and calling. Fights may take place in flight or on the ground. The male's display flight involves a similar wing and tail position with frequent wing clapping as he follows the female in a rising spiral. If she lands, he continues to display with bobbing and fluttering until the female spreads her wings and tail for copulation. Mating occasionally takes place on a raised perch instead of the ground. In good habitat, there may be 20 pairs per square kilometre (50 per square mile).
The European nightjar is normally monogamous. There is no nest, and the eggs are laid on the ground among plants or tree roots, or beneath a bush or tree. The site may be bare ground, leaf litter or pine needles, and is used for a number of years. The clutch is usually one or two whitish eggs, rarely unmarked, but normally blotched with browns and greys. The eggs average 32 mm × 22 mm (1.26 in × 0.87 in) and weigh 8.4 g (0.30 oz), of which 6% is shell.
Several nightjar species are known to be more likely to lay in the two weeks before the full moon than the during the waning moon, possibly because insect food may be easier to catch as the moon waxes. A study specifically looking at the European nightjar showed that the phase of the moon is a factor for birds laying in June, but not for earlier breeders. This strategy means that a second brood in July would also have a favourable lunar aspect.
Eggs are laid 36–48 hours apart, and incubation, mainly by the female, starts with the first egg. The male may incubate for short periods, especially around dawn or dusk, but spends the day roosting, sometimes outside his territory or close to other males. If the female is disturbed while breeding, she runs or flutters along the ground feigning injury until she has drawn the intruder away. She may also move the eggs a short distance with her bill. Each egg hatches after about 17–21 days. The semi-precocial downy chicks are mobile when hatched, but are brooded to keep them warm. They fledge in 16–17 days and become independent of the adults around 32 days after hatching. A second brood may be raised by early nesting pairs, in which case the female leaves the first brood a few days before they fledge; the male then cares for the first brood and assists with the second. Both adults feed the young with balls of insects which are either regurgitated into the chick's mouth or pecked by the chick from the adult's open bill.
Broods that fail tend to do so during incubation. One English study showed that only 14.5% of eggs survived to hatching, but once that stage was reached the chances of fledging successfully were high. European nightjars breed when aged one year, and typically live four years. The adult annual survival rate is 70%, but that for juveniles is unknown. The maximum known age in the wild is just over 12 years.


The goldfish (Carassius auratus) is a freshwater fish in the family Cyprinidae of order Cypriniformes. It is commonly kept as a pet in indoor aquariums, and is one of the most popular aquarium fish. Goldfish released into the wild have become an invasive pest in parts of North America.
Native to East Asia, the goldfish is a relatively small member of the carp family (which also includes the Prussian carp and the crucian carp). It was first selectively bred for color in imperial China more than 1,000 years ago, and several distinct breeds have since been developed. Goldfish breeds vary greatly in size, body shape, fin configuration, and coloration (various combinations of white, yellow, orange, red, brown, and black are known).


Goldfish have one of the most studied senses of vision in fishes. Goldfish have four kinds of cone cells, which are respectively sensitive to different colors: red, green, blue and ultraviolet. The ability to distinguish between four different primary colors classifies them as tetrachromats.


Goldfish can only grow to sexual maturity with enough water and the right nutrition. Most goldfish breed in captivity, particularly in pond settings. Breeding usually happens after a significant temperature change, often in spring. Males chase gravid female goldfish (females carrying eggs), and prompt them to release their eggs by bumping and nudging them.
Goldfish, like all cyprinids, are egg-layers. Their eggs are adhesive and attach to aquatic vegetation, typically dense plants such as Cabomba or Elodea or a spawning mop. The eggs hatch within 48 to 72 hours.
Within a week or so, the fry begins to assume its final shape, although a year may pass before they develop a mature goldfish color; until then they are a metallic brown like their wild ancestors. In their first weeks of life, the fry grow quickly—an adaptation born of the high risk of getting devoured by the adult goldfish (or other fish and insects) in their environment.
Some highly selectively bred goldfish can no longer breed naturally due to their altered shape. The artificial breeding method called "hand stripping" can assist nature, but can harm the fish if not done correctly. In captivity, adults may also eat young that they encounter.
Breeding goldfish by the hobbyist is the process of selecting adult fish to reproduce, allowing them to reproduce and then raising the resulting offspring while continually removing fish that do not approach the desired pedigree.


Goldfish are gregarious, displaying schooling behavior, as well as displaying the same types of feeding behaviors. Goldfish may display similar behaviors when responding to their reflections in a mirror.
Goldfish have learned behaviors, both as groups and as individuals, that stem from native carp behavior. They are a generalist species with varied feeding, breeding, and predator avoidance behaviors that contribute to their success. As fish, they can be described as "friendly" towards each other. Very rarely does a goldfish harm another goldfish, nor do the males harm the females during breeding. The only real threat that goldfish present to each other is competing for food. Commons, comets, and other faster varieties can easily eat all the food during a feeding before fancy varieties can reach it. This can lead to stunted growth or possible starvation of fancier varieties when they are kept in a pond with their single-tailed brethren. As a result, care should be taken to combine only breeds with similar body type and swim characteristics.


Goldfish have strong associative learning abilities, as well as social learning skills. In addition, their visual acuity allows them to distinguish between individual humans. Owners may notice that fish react favorably to them (swimming to the front of the glass, swimming rapidly around the tank, and going to the surface mouthing for food) while hiding when other people approach the tank. Over time, goldfish learn to associate their owners and other humans with food, often "begging" for food whenever their owners approach.
Goldfish that have constant visual contact with humans also stop considering them to be a threat. After being kept in a tank for several weeks, sometimes months, it becomes possible to feed a goldfish by hand without it shying away.
Goldfish have a memory-span of at least three months and can distinguish between different shapes, colors, and sounds.  By using positive reinforcement, goldfish can be trained to recognize and to react to light signals of different colors or to perform tricks. Fish respond to certain colors most evidently in relation to feeding. Fish learn to anticipate feedings provided they occur at around the same time every day.


Chinese tradition classifies goldfish into four main types. These classifications are not commonly used in the West.
Crucian (also called "grass") — Goldfish without fancy anatomical features, similar to Crucian carp or grass carp except for their coloration. These include the common goldfish, comet goldfish and Shubunkin.
Wen — Goldfish having a fancy tail, e.g., Fantails and Veiltails ("Wen" is also the name of the characteristic headgrowth on such strains as Oranda and Lionhead)
Dragon Eye — Goldfish having extended eyes, e.g., Black Moor, Bubble Eye, and Telescope Eye
Egg — Goldfish having no dorsal fin, usually with an 'egg-shaped' body, e.g., Lionhead. This group includes a Bubble Eye without a dorsal fin.


Like most species in the carp family, goldfish produce a large amount of waste both in their feces and through their gills, releasing harmful chemicals into the water. Build-up of this waste to toxic levels can occur in a relatively short period of time, and can easily cause a goldfish's death. For common and comet varieties, each goldfish should have about 20 US gallons (76 l; 17 imp gal) of water. Fancy goldfish (which are smaller) should have about 10 US gallons (38 l; 8.3 imp gal) per goldfish. The water surface area determines how much oxygen diffuses and dissolves into the water. A general rule is have 1 square foot (0.093 m²). Active aeration by way of a water pump, filter or fountain effectively increases the surface area.
The goldfish is classified as a coldwater fish, and can live in unheated aquaria at a temperature comfortable for humans. However, rapid changes in temperature (for example in an office building in winter when the heat is turned off at night) can kill them, especially if the tank is small. Care must also be taken when adding water, as the new water may be of a different temperature. Temperatures under about 10 °C (50 °F) are dangerous to fancy varieties, though commons and comets can survive slightly lower temperatures. Extremely high temperatures (over 30 °C (86 °F)) can also harm goldfish. However, higher temperatures may help fight protozoan infestations by accelerating the parasite's life-cycle—thus eliminating it more quickly. The optimum temperature for goldfish is between 20 °C (68 °F) and 22 °C (72 °F).
Like all fish, goldfish do not like to be petted. In fact, touching a goldfish can endanger its health, because it can cause the protective slime coat to be damaged or removed, exposing the fish's skin to infection from bacteria or water-born parasites. However, goldfish respond to people by surfacing at feeding time, and can be trained or acclimated to taking pellets or flakes from human fingers. The reputation of goldfish dying quickly is often due to poor care. The lifespan of goldfish in captivity can extend beyond 10 years.
If left in the dark for a period of time, goldfish gradually change color until they are almost gray. Goldfish produce pigment in response to light, similarly to how human skin becomes tanned in the sun. Fish have cells called chromatophores that produce pigments which reflect light, and give the fish coloration. The color of a goldfish is determined by which pigments are in the cells, how many pigment molecules there are, and whether the pigment is grouped inside the cell or is spaced throughout the cytoplasm.
Because goldfish eat live plants, their presence in a planted aquarium can be problematic. Only a few aquarium plant species (for example Cryptocoryne and Anubias) can survive around goldfish, but they require special attention so that they are not uprooted. Plastic plants are more durable.


Goldfish are popular pond fish, since they are small, inexpensive, colorful and very hardy. In an outdoor pond or water garden, they may even survive for brief periods if ice forms on the surface, as long as there is enough oxygen remaining in the water and the pond does not freeze solid. Common, London and Bristol shubunkins, jikin, wakin, comet and some hardier fantail goldfish can be kept in a pond all year round in temperate and subtropical climates. Moor, veiltail, oranda and lionhead can be kept safely in outdoor ponds year-round only in more tropical climates and only in summer elsewhere.
Compatible fish include rudd, tench, orfe and koi, but the last require specialized care. Ramshorn snails are helpful by eating any algae that grows in the pond. Without some form of animal population control, goldfish ponds can easily become overstocked. Fish such as orfe consume goldfish eggs.
Ponds small and large are fine in warmer areas (although it ought to be noted that goldfish can "overheat" in small volumes of water in the summer in tropical climates). In frosty climes, the depth should be at least 80 centimeters (31 in) to preclude freezing. During winter, goldfish become sluggish, stop eating and often stay on the bottom of the pond. This is normal; they become active again in the spring. Unless the pond is large enough to maintain its own ecosystem without interference from humans, a filter is important to clear waste and keep the pond clean. Plants are essential as they act as part of the filtration system, as well as a food source for the fish. Plants are further beneficial since they raise oxygen levels in the water.
Like their wild ancestors, common and comet goldfish as well as shubunkin can survive, and even thrive, in any climate that can support a pond, whereas fancy goldfish are unlikely to survive in the wild as their bright colors and long fins make them easy prey.


In the wild, the diet of goldfish consists of crustaceans, insects, and various plant matter. Like most fish, they are opportunistic feeders and do not stop eating on their own accord. Overfeeding can be deleterious to their health, typically by blocking the intestines. This happens most often with selectively bred goldfish, which have a convoluted intestinal tract. When excess food is available, they produce more waste and feces, partly due to incomplete protein digestion. Overfeeding can sometimes be diagnosed by observing feces trailing from the fish's cloaca.
Goldfish-specific food has less protein and more carbohydrate than conventional fish food. Enthusiasts may supplement this diet with shelled peas (with outer skins removed), blanched green leafy vegetables, and bloodworms. Young goldfish benefit from the addition of brine shrimp to their diet. As with all animals, goldfish preferences vary.


Like some other popular aquarium fish, such as the guppy, goldfish and other carp are frequently added to stagnant bodies of water to reduce mosquito populations. They are used to prevent the spread of West Nile virus, which relies on mosquitoes to migrate. However, introducing goldfish has often had negative consequences for local ecosystems.


Fishbowls are detrimental to the health of goldfish and are prohibited by animal welfare legislation in several municipalities. The practice of using bowls as permanent fish housing originated from a misunderstanding of Chinese "display" vessels: goldfish which were normally housed in ponds were, on occasion, temporarily displayed in smaller containers to be better admired by guests.
Goldfish kept in bowls or "mini-aquariums" suffer from death, disease, and stunting, due primarily to the low oxygen and very high ammonia/nitrite levels inherent in such an environment. In comparison to other common aquarium fish, goldfish have high oxygen needs and produce a large amount of waste due to the fact they lack a stomach; therefore they require a substantial volume of well-filtered water to thrive. In addition, all goldfish varieties have the potential to reach 5 inches (12.7 cm) in total length, with single-tailed breeds often exceeding one foot (30.5 cm). Single-tailed varieties include common and comet goldfish.
In many countries, carnival and fair operators commonly give goldfish away in plastic bags as prizes. In late 2005 Rome banned the use of goldfish and other animals as carnival prizes. Rome has also banned the use of "goldfish bowls", on animal cruelty grounds, as well as Monza, Italy, in 2004. In the United Kingdom, the government proposed banning this practice as part of its Animal Welfare Bill, though this has since been amended to only prevent goldfish being given as prizes to unaccompanied minors.
In Japan, during summer festivals and religious holidays (ennichi), a traditional game called goldfish scooping is played, in which a player scoops goldfish from a basin with a special scooper. Sometimes bouncy balls are substituted for goldfish.
Although edible and closely related to some fairly widely eaten species, goldfish are rarely eaten. A fad among American college students for many years was swallowing goldfish as a stunt and as a fraternity initiation process. The first recorded instance was in 1939 at Harvard University. The practice gradually fell out of popularity over the course of several decades and is rarely practiced today.
Some animal advocates have called for boycotts of goldfish purchases, citing industrial farming and low survival rates of the fish.


In Chinese history, goldfish was seen "as a symbol of luck and fortune". Moreover, only members of the Song Dynasty could own goldfish. In Iran and among the international Iranian diaspora, goldfish is a traditional part of Nowruz celebrations. Goldfish is usually used in Haft-sin table as the symbol of progress.


Because the sand tiger shark is worldwide in distribution, it has many common names. The term "sand tiger shark" actually refers to four different sand tiger shark species in the family Odontaspididae. Furthermore, the name creates confusion with the unrelated tiger shark Galeocerdo cuvier. The grey nurse shark, the name used in Australia and the United Kingdom, is the second-most used name for the shark, and in India it is known as blue-nurse sand tiger. However, there are unrelated nurse sharks in the family Ginglymostomatidae. The most unambiguous and descriptive English name is probably the South African one, spotted ragged-tooth shark.


C. maenas can live in all types of protected and semiprotected marine and estuarine habitats, including those with mud, sand, or rock substrates, submerged aquatic vegetation, and emergent marsh, although soft bottoms are preferred. C. maenas is euryhaline, meaning that it can tolerate a wide range of salinities (from 4 to 52 ‰), and survive in temperatures of 0 to 30 °C (32 to 86 °F). The wide salinity range allows C. maenas to survive in the lower salinities found in estuaries, and the wide temperature range allows it to survive in extremely cold climates beneath the ice in winter. A molecular biological study using the COI gene found genetic differentiation between the North Sea and the Bay of Biscay, and even more strongly between the populations in Iceland and the Faroe Islands and those elsewhere. This suggests that C. maenas is unable to cross deeper water.
Females can produce up to 185,000 eggs, and larvae develop offshore in several stages before their final moult to juvenile crabs in the intertidal zone. Young crabs live among seaweeds and seagrasses, such as Posidonia oceanica, until they reach adulthood.
C. maenas has the ability to disperse by a variety of mechanisms, including ballast water, ships' hulls, packing materials (seaweeds) used to ship live marine organisms, bivalves moved for aquaculture, rafting, migration of crab larvae on ocean currents, and the movement of submerged aquatic vegetation for coastal zone management initiatives.  C. maenas dispersed in Australia mainly by rare long-distance events, possibly caused by human actions.
C. maenas is a predator, feeding on many organisms, particularly bivalve molluscs (such as clams – up to 40 1⁄2 inch (13 mm) clams per day, oysters, and mussels), polychaetes, and small crustaceans – including other crabs up to their own size. They are primarily diurnal, although activity also depends on the tide, and crabs can be active at any time of day. In California, preferential predation of C. maenas on native clams (Nutricola spp.) resulted in the decline of the native clams and an increase of a previously introduced clam (the amethyst gem clam, Gemma gemma), although C. maenas also voraciously preys on introduced clams such as Potamocorbula amurensis. The soft-shell clam (Mya arenaria) is a preferred prey species of C. maenas. Consequently, it has been implicated in the destruction of the soft-shell clam fisheries on the east coast of the United States and Canada, and the reduction of populations of other commercially important bivalves (such as scallops, Argopecten irradians, and northern quahogs, Mercenaria mercenaria). The prey of C. maenas includes the young of bivalves and fish, although the effect of its predation on winter flounder, Pseudopleuronectes americanus is minimal. C. maenas can, however, have substantial negative impacts on local commercial and recreational fisheries, by preying on the young of species, such as oysters (adults' shells are too tough for C. maenas to crack) and the Dungeness crab, or competing with them for resources Colder water temperatures reduce overall feeding rates of C. maenas. and eating the Zostera marina that Dungeness and juvenile salmon depend upon for habitat.
To protect itself against predators, C. maenas uses different camouflage strategies depending on their habitat: crabs in mudflats try to resemble their surroundings with colours similar to the mud while crabs in rock pool use disruptive coloration.


Cardamine concatenata, the cutleaved toothwort, crow's toes, pepper root or purple-flowered toothwort, is a flowering plant in Brassicaceae. It owes its name to the tooth-like appearance of its rhizome. It is a  perennial woodland wildflower native to eastern North America. It is considered a spring ephemeral and blooms in March, April, and/or May.


The vegetative parts of this plant, which can reach 20–40 cm, arise from a segmented rhizome. The leaves are on long petioles, deeply and palmately dissected into five segments with large "teeth" on the margins. The white to pinkish flowers are held above the foliage in a spike. Fruit is an elongated pod which can be up to 4 cm long. Its native habitats include rich woods, wooded bottomlands, limestone outcrops, and rocky banks and bluffs.


Cardamine pratensis, the cuckoo flower, lady's smock, mayflower, or milkmaids, is a flowering plant in the family Brassicaceae. It is a perennial herb native throughout most of Europe and Western Asia. The specific name pratensis is Latin for "meadow".


The Canada warbler is sometimes called the "necklaced warbler," because of the band of dark streaks across its chest. The adults have minimal sexual dimorphism, although the male's "necklace" is darker and more conspicuous and also has a longer tail. Adults are 12–15 cm (4.7–5.9 in) long, have a wingspan of 17–22 cm (6.7–8.7 in) and weigh 9–13 g (0.32–0.46 oz).
The chest, throat and belly of the bird is yellow, and its back is dark grey. It has no wingbars or tail spots, but the underside of the tail is white. It has a yellow line in front of its eye in the direction of the beak, but the most striking facial feature is the white eyerings or "spectacles." Immature specimens have similar coloration as adults but duller and with less pronounced facial features.