The Andean gull is known from Chile, Peru, Argentina, Bolivia, Ecuador and Colombia where it breeds in mountainous areas, usually at elevations of over 3,000 metres (9,800 ft). In areas with harsh winters it migrates to lower elevations in winter, sometimes as far as the coast. Flocks of these gulls are common on the coast in southern Peru and northern Chile, but It has yet to be established where these particular birds breed. In the wetlands of the Altiplano it is resident and breeds at 4,000 metres (13,000 ft) while in southern Argentina it may breed at as little as 2,000 metres (6,600 ft) but this is in similar habitats to the birds further north. The Andean Gull nests in small scattered colonies, sometimes even as solitary pairs, in small, isolated ponds. Unusually for a gull of this size, it becomes sexually mature at the age of two years rather than three.
It feeds on a variety of prey items, including insects and worms taken on fields and grasslands; it also sometimes catches aerial insects and eats some garbage when available.


Chrysaora fuscescens is commonly found along the coasts of California and Oregon, though some reside in the waters north to the Gulf of Alaska, west to the seas around Japan and south to the Baja Peninsula. The populations reach their peak during the late summer. In recent years, C. fuscescens has become overly abundant off the coast of Oregon, which is thought to be an indicator of climate change. However, others suspect that the population is increasing because of human influences to coastal regions.


Chrysaora fuscescens is capable of both sexual reproduction in the medusa stage and asexual reproduction in the polyp stage. The life cycle of C. fuscescens begins when females catch sperm released by the males to fertilize the eggs she has produced and is holding in her mouth. These fertilized eggs remain attached to her oral arms, and there they grow into flat bean-shaped planula. Once they grow into flower-shaped polyps, they are released into the ocean where they attach themselves to a solid surface and undergo asexual reproduction. The polyp makes identical copies of itself by means of budding, where the new polyp grows from its side. After the new polyp is fully formed, it too is released into the ocean and undergoes metamorphosis as it grows, developing a bell, arms, and tentacles until it is a fully formed medusa.


The painted turtle (Chrysemys picta) is the most widespread native turtle of North America. It lives in slow-moving fresh waters, from southern Canada to northern Mexico, and from the Atlantic to the Pacific. They have been shown to prefer large wetlands with long periods of innundation and emergent vegetation. This turtle is a member of the genus Chrysemys, which is part of the pond turtle family Emydidae. Fossils show that the painted turtle existed 15 million years ago. Three regionally based subspecies (the eastern, midland, and western) evolved during the last ice age. The southern painted turtle (C. dorsalis) is alternately considered the only other species in Chrysemys, or another subspecies of C. picta.
The adult painted turtle is 13–25 cm (5–10 in) long; the male is smaller than the female. The turtle's top shell is dark and smooth, without a ridge. Its skin is olive to black with red, orange, or yellow stripes on its extremities. The subspecies can be distinguished by their shells: the eastern has straight-aligned top shell segments; the midland has a large gray mark on the bottom shell; the western has a red pattern on the bottom shell.
The turtle eats aquatic vegetation, algae, and small water creatures including insects, crustaceans, and fish. Painted turtles primarily feed while in water and are able to locate and subdue prey even in heavily clouded conditions. Although they are frequently consumed as eggs or hatchlings by rodents, canines, and snakes, the adult turtles' hard shells protect them from most predators. Reliant on warmth from its surroundings, the painted turtle is active only during the day when it basks for hours on logs or rocks. During winter, the turtle hibernates, usually in the mud at the bottom of water bodies. The turtles mate in spring and autumn. Females dig nests on land and lay eggs between late spring and mid-summer. Hatched turtles grow until sexual maturity: 2–9 years for males, 6–16 for females.
In the traditional tales of Algonquian tribes, the colorful turtle played the part of a trickster. In modern times, four U.S. states (Colorado, Illinois, Michigan, and Vermont) have named the painted turtle their official reptile. While habitat loss and road killings have reduced the turtle's population, its ability to live in human-disturbed settings has helped it remain the most abundant turtle in North America. Adults in the wild can live for more than 55 years.


Originally described in 1783 by Johann Gottlob Schneider as Testudo picta, the painted turtle was called Chrysemys picta first by John Edward Gray in 1855. Four subspecies were then recognized: the eastern by Schneider in 1783, the western by Gray in 1831, and the midland and southern by Louis Agassiz in 1857, though the southern painted turtle is now generally considered a full species.


Although its evolutionary history—what the forerunner to the species was and how the close relatives branched off—is not well understood, the painted turtle is common in the fossil record. The oldest samples, found in Nebraska, date to about 15 million years ago. Fossils from 15 million to about 5 million years ago are restricted to the Nebraska-Kansas area, but more recent fossils are gradually more widely distributed. Fossils newer than 300,000 years old are found in almost all the United States and southern Canada.


Adult painted turtles can grow to 13–25 cm (5–10 in) long, with males being smaller. The shell is oval, smooth with little grooves where the large scale-like plates overlap, and flat-bottomed. The color of the top shell (carapace) varies from olive to black. Darker specimens are more common where the bottom of the water body is darker. The bottom shell (plastron) is yellow, sometimes red, sometimes with dark markings in the center. Similar to the top shell, the turtle's skin is olive to black, but with red and yellow stripes on its neck, legs, and tail. As with other pond turtles, such as the bog turtle, the painted turtle's feet are webbed to aid swimming.
The head of the turtle is distinctive. The face has only yellow stripes, with a large yellow spot and streak behind each eye, and on the chin two wide yellow stripes that meet at the tip of the jaw. The turtle's upper jaw is shaped into an inverted "V" (philtrum), with a downward-facing, tooth-like projection on each side.
The hatchling has a proportionally larger head, eyes, and tail, and a more circular shell than the adult. The adult female is generally longer than the male, 10–25 cm (4–10 in) versus 7–15 cm (3–6 in). For a given length, the female has a higher (more rounded, less flat) top shell. The female weighs around 500 g (18 oz) on average, against the males' average adult weight of roughly 300 g (11 oz). The female's greater body volume supports her egg-production. The male has longer foreclaws and a longer, thicker tail, with the anus (cloaca) located further out on the tail.


The most widespread North American turtle, the painted turtle is the only turtle whose native range extends from the Atlantic to the Pacific. It is native to eight of Canada's ten provinces, forty-five of the fifty United States, and one of Mexico's thirty-one states. On the East Coast, it lives from the Canadian Maritimes to the U.S. state of Georgia. On the West Coast, it lives in British Columbia, Washington, and Oregon and offshore on southeast Vancouver Island. The northernmost American turtle, its range includes much of southern Canada. To the south, its range reaches the U.S. Gulf Coast in Louisiana and Alabama. In the southwestern United States there are only dispersed populations. It is found in one river in extreme northern Mexico. It is absent in a part of southwestern Virginia and the adjacent states as well as in north-central Alabama.
The borders between the four subspecies are not sharp, because the subspecies interbreed. Many studies have been performed in the border regions to assess the intermediate turtles, usually by comparing the anatomical features of hybrids that result from intergradation of the classical subspecies. Despite the imprecision, the subspecies are assigned nominal ranges.


The eastern painted turtle ranges from southeastern Canada to Georgia with a western boundary at approximately the Appalachians. At its northern extremes, the turtle tends to be restricted to the warmer areas closer to the Atlantic Ocean. It is uncommon in far north New Hampshire and in Maine is common only in a strip about 50 miles from the coast. In Canada, it lives in New Brunswick and Nova Scotia but not in Quebec or Prince Edward Island. To the south it is not found in the coastal lowlands of southern North Carolina, South Carolina, or Georgia, or in southern Georgia in general or at all in Florida.
The eastern subspecies's range extends slightly into east central Alabama, where it intergrades with the southern subspecies. In the northeast, there is extensive mixing with the midland subspecies, and some writers have called these turtles a "hybrid swarm". In the southeast, the border between the eastern and midland is more sharp as mountain chains separate the subspecies to different drainage basins.


The midland painted turtle lives from southern Ontario and Quebec, through the eastern U.S. Midwest states, to Kentucky, Tennessee and northwestern Alabama, where it intergrades with the southern painted turtle. It also is found eastward through West Virginia, western Maryland and Pennsylvania. The midland painted turtle appears to be moving east, especially in Pennsylvania. To the northeast it is found in western New York and much of Vermont, and it intergrades extensively with the eastern subspecies.


The western painted turtle's northern range includes southern parts of western Canada from Ontario through Manitoba, Saskatchewan, Alberta and British Columbia. In Ontario, the western subspecies is found north of Minnesota and directly north of Lake Superior, but there is a 130 km (80 mi) gap to the east of Lake Superior (in the area of harshest winter climate) where no painted turtles of any subspecies occur. Thus Ontario's western subspecies does not intergrade with the midland painted turtle of southeastern Ontario. In Manitoba, the turtle is numerous and ranges north to Lake Manitoba and the lower part of Lake Winnipeg. The turtle is also common in south Saskatchewan, but in Alberta, there may only be 100 individuals, all found very near the U.S. border, mostly in the southeast.
In British Columbia, populations exist in the interior in the vicinity of the Kootenai, Columbia, Okanagan, and Thompson river valleys. At the coast, turtles occur near the mouth of the Fraser and a bit further north, as well as the bottom of Vancouver Island, and some other nearby islands. Within British Columbia, the turtle's range is not continuous and can better be understood as northward extensions of the range from the United States. High mountains present barriers to east–west movement of the turtles within the province or from Alberta. Some literature has shown isolated populations much further north in British Columbia and Alberta, but these were probably pet-releases.
In the United States, the western subspecies forms a wide intergrade area with the midland subspecies covering much of Illinois as well as a strip of Wisconsin along Lake Michigan and part of the Upper Peninsula of Michigan (UP). Further west, the rest of Illinois, Wisconsin and the UP are part of the range proper, as are all of Minnesota and Iowa, as well as all of Missouri except a narrow strip in the south. All of North Dakota is within range, all of South Dakota except a very small area in the west, and all of Nebraska. Almost all of Kansas is in range; the border of that state with Oklahoma is roughly the species range border, but the turtle is found in three counties of north central Oklahoma.
To the northwest, almost all of Montana is in range. Only a narrow strip in the west, along most of the Idaho border (which is at the Continental Divide) lacks turtles. Wyoming is almost entirely out of range; only the lower elevation areas near the eastern and northern borders have painted turtles. In Idaho, the turtles are found throughout the far north (upper half of the Idaho Panhandle). Recently, separate Idaho populations have been observed in the southwest (near the Payette and Boise rivers) and the southeast (near St. Anthony). In Washington state, turtles are common throughout the state within lower elevation river valleys. In Oregon, the turtle is native to the northern part of the state throughout the Columbia River Valley as well as the Willamette River Valley north of Salem.
To the southwest, the painted turtle's range is fragmented. In Colorado, while range is continuous in the eastern, prairie, half of the state, it is absent in most of the western, mountainous, part of the state. However, the turtle is confirmed present in the lower elevation southwest part of the state (Archuleta and La Plata counties), where a population ranges into northern New Mexico in the San Juan River basin. In New Mexico, the main distribution follows the Rio Grande and the Pecos River, two waterways that run in a north–south direction through the state. Within the aforementioned rivers, it is also found in the northern part of Far West Texas. In Utah, the painted turtle lives in an area to the south (Kane County) in streams draining into the Colorado River, although it is disputed if they are native. In Arizona, the painted turtle is native to an area in the east, Lyman Lake. The painted turtle is not native to Nevada or California.
In Mexico, painted turtles have been found about 50 miles south of New Mexico near Galeana in the state of Chihuahua. There, two expeditions found the turtles in the Rio Santa Maria which is in a closed basin.


Pet releases are starting to establish the painted turtle outside its native range. It has been introduced into waterways near Phoenix, Arizona, and to Germany, Indonesia, the Philippines, and Spain.


To thrive, painted turtles need fresh waters with soft bottoms, basking sites, and aquatic vegetation. They find their homes in shallow waters with slow-moving currents, such as creeks, marshes, ponds, and the shores of lakes. The subspecies have evolved different habitat preferences.
The eastern painted turtle is very aquatic, leaving the immediate vicinity of its water body only when forced by drought to migrate. Along the Atlantic, painted turtles have appeared in brackish waters.
The midland and southern painted turtles seek especially quiet waters, usually shores and coves. They favor shallows that contain dense vegetation and have an unusual toleration of pollution.
The western painted turtle lives in streams and lakes, similar to the other painted turtles, but also inhabits pasture ponds and roadside pools. It is found as high as 1,800 m (5,900 ft).


Within much of its range, the painted turtle is the most abundant turtle species. Population densities range from 10 to 840 turtles per hectare (2.5 acres) of water surface. Warmer climates produce higher relative densities among populations, and habitat desirability also influences density. Rivers and large lakes have lower densities because only the shore is desirable habitat; the central, deep waters skew the surface-based estimates. Also, lake and river turtles have to make longer linear trips to access equivalent amounts of foraging space.
Adults outnumber juveniles in most populations, but gauging the ratios is difficult because juveniles are harder to catch; with current sampling methods, estimates of age distribution vary widely. Annual survival rate of painted turtles increases with age. The probability of a painted turtle surviving from the egg to its first birthday is only 19%. For females, the annual survival rate rises to 45% for juveniles and 95% for adults. The male survival rates follow a similar pattern, but are probably lower overall than females, as evidenced by the average male age being lower than that of the female. Natural disasters can confound age distributions. For instance, a hurricane can destroy many nests in a region, resulting in fewer hatchlings the next year. Age distributions may also be skewed by migrations of adults.
To understand painted turtle adult age distributions, researchers require reliable methods. Turtles younger than four years (up to 12 years in some populations) can be aged based on "growth rings" in their shells. For older turtles, some attempts have been made to determine age based on size and shape of their shells or legs using mathematical models, but this method is more uncertain. The most reliable method to study the long-lived turtles is to capture them, permanently mark their shells by notching with a drill, release the turtles, and then recapture them in later years. The longest-running study, in Michigan, has shown that painted turtles can live more than 55 years.
Adult sex ratios of painted turtle populations average around 1:1. Many populations are slightly male-heavy, but some are strongly female-imbalanced; one population in Ontario has a female to male ratio of 4:1. Hatchling sex ratio varies based on egg temperature. During the middle third of incubation, temperatures of 23–27 °C (73–81 °F) produce males, and anything above or below that, females. It does not appear that females choose nesting sites to influence the sex of the hatchlings; within a population, nests will vary sufficiently to give both male and female-heavy broods.


The painted turtle hunts along water bottoms. It quickly juts its head into and out of vegetation to stir potential victims out into the open water, where they are pursued. The turtle holds large prey in its mouth and tears the prey apart with its forefeet. It also consumes plants and skims the surface of the water with its mouth open to catch small particles of food.
Although all subspecies of painted turtle eat both plants and animals (in the form of leaves, algae, fish, crustaceans, aquatic insects and carrion), their specific diets vary.
Painted turtles obtain coloration from carotenoids in their natural diet by eating algae and a variety of aquatic plants from their environment.Stripes and spots increase red and yellow chroma and decrease UV chroma and brightness in turtles with large amounts of carotenoids in their diet compared to the stripes and spots of turtles with only moderate amounts of carotenoids in their diet.
The eastern painted turtle's diet is the least studied. It prefers to eat in the water, but has been observed eating on land. The fish it consumes are typically dead or injured.
The midland painted turtle eats mostly aquatic insects and both vascular and non-vascular plants.
The western painted turtle's consumption of plants and animals changes seasonally. In early summer, 60% of its diet comprises insects. In late summer, 55% includes plants. Of note, the western painted turtle aids in the dispersal of white water-lily seeds. The turtle consumes the hard-coated seeds, which remain viable after passing through the turtle, and disperses them through its feces.


Painted turtles are most vulnerable to predators when young. Nests are frequently ransacked and the eggs eaten by garter snakes, crows, chipmunks, thirteen-lined ground and gray squirrels, skunks, groundhogs, raccoons, badgers, gray and red fox, and humans. The small and sometimes bite-size, numerous hatchlings fall prey to water bugs, bass, catfish, bullfrogs, snapping turtles, three types of snakes (copperheads, racers and water snakes), herons, rice rats, weasels, muskrats, minks, and raccoons. As adults, the turtles' armored shells protect them from many potential predators, but they still occasionally fall prey to alligators, ospreys, crows, red-shouldered hawks, bald eagles, and especially raccoons.
Painted turtles defend themselves by kicking, scratching, biting, or urinating. In contrast to land tortoises, painted turtles can right themselves if they are flipped upside down.


The painted turtles mate in spring and fall in waters of 10–25 °C (50–77 °F). Males start producing sperm in early spring, when they can bask to an internal temperature of 17 °C (63 °F). Females begin their reproductive cycles in mid-summer, and ovulate the following spring.
Courtship begins when a male follows a female until he meets her face-to-face. He then strokes her face and neck with his elongated front claws, a gesture returned by a receptive female. The pair repeat the process several times, with the male retreating from and then returning to the female until she swims to the bottom, where they copulate. As the male is smaller than the female, he is not dominant. Although not directly observed, evidence indicates that the male will inflict injury on the female in attempts of coercion. Males will use their tooth-like cusps on their beaks and their foreclaws during this act of coercion with the female. The female stores sperm, to be used for up to three clutches, in her oviducts; the sperm may remain viable for up to three years. A single clutch may have multiple fathers.


Nesting is done, by the females only, between late May and mid-July. The nests are vase-shaped and are usually dug in sandy soil, often at sites with southern exposures. Nests are often within 200 m (220 yd) of water, but may be as far away as 600 m (660 yd), with older females tending to nest further inland. Nest sizes vary depending on female sizes and locations but are about 5–11 cm (2–4 in) deep. Females may return to the same sites several consecutive years, but if several females make their nests close together, the eggs become more vulnerable to predators. Female eastern painted turtles have been shown to nest together, possibly even participating in communal nesting.
The female's optimal body temperature while digging her nest is 29–30 °C (84–86 °F). If the weather is unsuitable, for instance a too hot night in the Southeast, she delays the process until later at night. Painted turtles in Virginia have been observed waiting three weeks to nest because of a hot drought.
While preparing to dig her nest, the female sometimes exhibits a mysterious preliminary behavior. She presses her throat against the ground of different potential sites, perhaps sensing moisture, warmth, texture, or smell, although her exact motivation is unknown. She may further temporize by excavating several false nests as the wood turtles also do.
The female relies on her hind feet for digging. She may accumulate so much sand and mud on her feet that her mobility is reduced, making her vulnerable to predators. To lighten her labors, she lubricates the area with her bladder water. Once the nest is complete, the female deposits into the hole. The freshly laid eggs are white, elliptical, porous, and flexible. From start to finish, the female's work may take four hours. Sometimes she remains on land overnight afterwards, before returning to her home water.
Females can lay five clutches per year, but two is a normal average after including the 30–50% of a population's females that do not produce any clutches in a given year. In some northern populations, no females lay more than one clutch per year. Bigger females tend to lay bigger eggs and more eggs per clutch. Clutch sizes of the subspecies vary, although the differences may reflect different environments, rather than different genetics. The two more northerly subspecies, western and midland, are larger and have more eggs per clutch—11.9 and 7.6, respectively—than the eastern (4.9). Within subspecies, also, the more northerly females lay larger clutches.


Incubation lasts 72–80 days in the wild and for a similar period in artificial conditions. In August and September, the young turtle breaks out from its egg, using a special projection of its jaw called the egg tooth. Not all offspring leave the nest immediately, though. Hatchlings north of a line from Nebraska to northern Illinois to New Jersey typically arrange themselves symmetrically in the nest and overwinter to emerge the following spring.
The hatchling's ability to survive winter in the nest has allowed the painted turtle to extend its range farther north than any other American turtle. The painted turtle is genetically adapted to survive extended periods of subfreezing temperatures with blood that can remain supercooled and skin that resists penetration from ice crystals in the surrounding ground. The hardest freezes nevertheless kill many hatchlings.
Immediately after hatching, turtles are dependent on egg yolk material for sustenance. About a week to a week and a half after emerging from their eggs (or the following spring if emergence is delayed), hatchlings begin feeding to support growth. The young turtles grow rapidly at first, sometimes doubling their size in the first year. Growth slows sharply at sexual maturity and may stop completely. Likely owing to differences of habitat and food by water body, growth rates often differ from population to population in the same area. Among the subspecies, the western painted turtles are the quickest growers.
Females grow faster than males overall, and must be larger to mature sexually. In most populations males reach sexual maturity at 2–4 years old, and females at 6–10. Size and age at maturity increase with latitude; at the northern edge of their range, males reach sexual maturity at 7–9 years of age and females at 11–16.


In the spring, when the water reaches 15–18 °C (59–64 °F), the turtle begins actively foraging. However, if the water temperature exceeds 30 °C (86 °F), the turtle will not feed. In fall, the turtle stops foraging when temperatures drop below the spring set-point.
During the winter, the turtle hibernates. In the north, the inactive season may be as long as from October to March, while the southernmost populations may not hibernate at all. While hibernating, the body temperature of the painted turtle averages 6 °C (43 °F). Periods of warm weather bring the turtle out of hibernation, and even in the north, individuals have been seen basking in February.
The painted turtle hibernates by burying itself, either on the bottom of a body of water, near water in the shore-bank or the burrow of a muskrat, or in woods or pastures. When hibernating underwater, the turtle prefers shallow depths, no more than 2 m (7 ft). Within the mud, it may dig down an additional 1 m (3 ft). In this state, the turtle does not breathe, although if surroundings allow, it may get some oxygen through its skin. The species is one of the best-studied vertebrates able to survive long periods without oxygen. Adaptations of its blood chemistry, brain, heart, and particularly its shell allow the turtle to survive extreme lactic acid buildup while oxygen-deprived.


Searching for water, food, or mates, the painted turtles travel up to several kilometers at a time. During summer, in response to heat and water-clogging vegetation, the turtles may vacate shallow marshes for more permanent waters. Short overland migrations may involve hundreds of turtles together. If heat and drought are prolonged, the turtles will bury themselves and, in extreme cases, die.
Foraging turtles frequently cross lakes or travel linearly down creeks. Daily crossings of large ponds have been observed. Tag and release studies show that sex also drives turtle movement. Males travel the most, up to 26 km (16 mi), between captures; females the second most, up to 8 km (5 mi), between captures; and juveniles the least, less than 2 km (1.2 mi), between captures. Males move the most and are most likely to change wetlands because they seek mates.
The painted turtles, through visual recognition, have homing capabilities. Many individuals can return to their collection points after being released elsewhere, trips that may require them to traverse land. One experiment placed 98 turtles varying several-kilometer distances from their home wetland; 41 returned. When living in a single large body of water, the painted turtles can home from up to 6 km (4 mi) away. Another experiment found that if placed far enough away from water the turtles will just walk in straight paths and not orient towards water or in any specific direction which indicates a lack of homing ability. Females may use homing to help locate suitable nesting sites.
Eastern painted turtle movements may contribute to aquatic plant seed dispersal. A study done in Massachusetts found that the quantity of intact macrophyte seeds defecated by Eastern painted turtles can be high and that the seeds of specifically Nymphaea ordorata that were found in feces were capable of moderate to high level germination. As turtles move between ponds and habitats, they carry seeds along with them to new locations.


The species is currently classified as least concern by the IUCN but populations have been subject to decline locally.
The decline in painted turtle populations is not a simple matter of dramatic range reduction, like that of the American bison. Instead the turtle is classified as G5 (demonstrably widespread) in its Natural Heritage Global Rank, and the IUCN rates it as a species of least concern. The painted turtle's high reproduction rate and its ability to survive in polluted wetlands and artificially made ponds have allowed it to maintain its range, but the post-Columbus settlement of North America has reduced its numbers.
Only within the Pacific Northwest is the turtle's range eroding. Even there, in Washington, the painted turtle is designated S5 (demonstrably widespread). However, in Oregon, the painted turtle is designated S2 (imperiled), and in British Columbia, the turtle's populations in the Coast and Interior regions are labeled "endangered" and "of special concern", respectively.
Much is written about the different factors that threaten the painted turtle, but they are unquantified, with only inferences of relative importance. A primary threat category is habitat loss in various forms. Related to water habitat, there is drying of wetlands, clearing of aquatic logs or rocks (basking sites), and clearing of shoreline vegetation, which allows more predator access or increased human foot traffic. Related to nesting habitat, urbanization or planting can remove needed sunny soils.
Another significant human impact is roadkill—dead turtles, especially females, are commonly seen on summer roads. In addition to direct killing, roads genetically isolate some populations. Localities have tried to limit roadkill by constructing underpasses, highway barriers, and crossing signs. Oregon has introduced public education on turtle awareness, safe swerving, and safely assisting turtles across the road.
In the West, human-introduced bass, bullfrogs, and especially snapping turtles, have increased the predation of hatchlings. Outside the Southeast, where sliders are native, released pet red-eared slider turtles increasingly compete with painted turtles. In cities, increased urban predators (raccoons, canines, and felines) may impact painted turtles by eating their eggs.
Other factors of concern for the painted turtles include over-collection from the wild, released pets introducing diseases or reducing genetic variability, pollution, boating traffic, angler's hooks (the turtles are noteworthy bait-thieves), wanton shooting, and crushing by agricultural machines or golf course lawnmowers or all-terrain vehicles. Gervais and colleagues note that research itself impacts the populations and that much funded turtle trapping work has not been published. They advocate discriminating more on what studies are done, thereby putting fewer turtles into scientists' traps. Global warming represents an uncharacterized future threat.
As the most common turtle in Nova Scotia, the eastern painted turtle is not listed under the Species at Risk Act for conservation requirements.


Commercial harvesting of painted turtles in the wild is controversial and, increasingly, restricted. Wisconsin formerly had virtually unrestricted trapping of painted turtles but based on qualitative observations forbade all commercial harvesting in 1997. Neighboring Minnesota, where trappers collected more than 300,000 painted turtles during the 1990s, commissioned a study of painted turtle harvesting. Scientists found that harvested lakes averaged half the painted turtle density of off-limit lakes, and population modeling suggested that unrestricted harvests could produce a large decline in turtle populations. In response, Minnesota forbade new harvesters in 2002 and limited trap numbers. Although harvesting continued, subsequent takes averaged half those of the 1990s. As of 2009, painted turtles faced virtually unlimited harvesting in Arkansas, Iowa, Missouri, Ohio, and Oklahoma; since then, Missouri has prohibited their harvesting.
Individuals who trap painted turtles typically do so to earn additional income, selling a few thousand a year at $1–2 each. Many trappers have been involved in the trade for generations, and value it as a family activity. Some harvesters disagree with limiting the catch, saying the populations are not dropping.
Many U.S. state fish and game departments allow non-commercial taking of painted turtles under a creel limit, and require a fishing (sometimes hunting) license; others completely forbid the recreational capture of painted turtles. Trapping is not allowed in Oregon, where western painted turtle populations are in decline, and in Missouri, where there are populations of both southern and western subspecies. In Canada, Ontario protects both subspecies present, the midland and western, and British Columbia protects its dwindling western painted turtles.
Capture methods are also regulated by locality. Typically trappers use either floating "basking traps" or partially submerged, baited "hoop traps". Trapper opinions, commercial records, and scientific studies show that basking traps are more effective for collecting painted turtles, while the hoop traps work better for collecting "meat turtles" (snapping turtles and soft-shell turtles). Nets, hand capture, and fishing with set lines are generally legal, but shooting, chemicals, and explosives are forbidden.


The adults are an iridescent blue-green with a metallic copper, golden or crimson shine. Dogbane beetles range from 8 to 11 mm in length, and they are oblong and convex in shape. The head contains two antennae that are widely set apart and located between the beetle's eyes. They are twelve-jointed with eleven segments, and the basal segment is shorter than that of the second segment.
The larvae have white bodies and brown heads.


Because the dogbane beetle's main source of food is dogbane leaves, they live in areas where this plant is abundant, such as fields, forests, railroads and lakeshores.
Chrysochus auratus is generally found in eastern North America, spanning the entire eastern United States and into adjacent southern Canada west of the Rocky Mountains. At the western edge of its range, it extends west of the Rocky Mountains into Arizona and Utah. The related species Chrysochus cobaltinus, in contrast, is found exclusively in western North America, and the two species were historically considered to have allopatric distributions. Recently, at least two narrow regions in western North America have been documented where both C. auratus and C. cobaltinus occur and apparently interbreed.


Chrysomela populi can reach a length of about 9–13 mm. The female is slightly larger than the male. These beetles show a black, dark blue or dark green body, that is round and ladybird-like. Head and pronotum are black, while elytra is bright red, with a black stain at the base. Some beetles come as orange coloured.
It can be distinguished from Chrysolina grossa by its shorter antennae and less estensive pronotum. It is also rather similar to Chrysomela saliceti  and Chrysomela tremula.


Chrysomela vigintipunctata, or the spotted willow leaf beetle, is a species of broad-shouldered leaf beetle belonging to the family Chrysomelidae, subfamily Chrysomelinae.
The length of the beetles varies from 6.5 to 9.2 mm. Both larvae and adults feed on the leaves of its host plants, especially Salicaceae species. Elytra usually are yellowish-green, with ten black spots on each side, while the head and the pronotum are black, with yellow sides.
Beetles live mostly in forests, along lakes, swamps and feed on willow and Populus.
They are found in most of Europe, Mongolia, Russian Far East and Japan. A single dead specimen was found, for the first time in Britain, at Southwold beach, east Suffolk on 26 May 2016. A search in suitable habitat for more specimens was unsuccessful.


Chytolita is a monotypic litter moth genus of the family Erebidae erected by Augustus Radcliffe Grote in 1873. Its only species, Chytolita morbidalis, the morbid owlet moth or morbid owlet, was first described by Achille Guenée in 1854. It is found in large parts of North America, from coast to coast in the north and south to North Carolina, Texas and Florida in the west. The habitat consists of deciduous woods and edges.
The wingspan is 29–35 mm. The forewings are pale grayish white with diffuse, brownish or orangish lines. The hindwings are even paler with faint grayish veins. Adults are on wing from May to August. There seems to be one generation per year.
The larvae feed on the leaves of various deciduous trees and have been reared on dandelion, grass, hazel and lettuce. There are also records for dead leaves, including dried white oak leaves.


The mottled owl is a medium-sized owl with adults reaching 280 to 355 mm (11.0 to 14.0 in) in length. Females are considerably larger than males; the mottled owl shows the greatest degree of sexual dimorphism of any species of owl. The crown, nape and back are mottled in several shades of darkish brown, the facial disc is pale brown and the throat, breast and belly are off-white with distinctive vertical brown streaks. The large eyes are brown, the beak is greyish-yellow or greyish-blue, and the legs and feet are greyish-yellow. There is a darker form of the bird with a buff breast and belly. Mottled owls produce a range of calls which include a hoot used in maintaining territory boundaries, and various whistles, screeches and hisses.


The mottled owl is native to Central and South America. Its range extends from Mexico south to Argentina and Brazil and it is found at elevations up to about 2,500 m (8,200 ft). It inhabits a variety of wooded habitats including rainforest, woodland verges, dry thorn forest, pine/oak woodland and plantations and also open countryside with scattered trees. In some parts of its range it is common and it is often found close to human habitations.


The mottled owl is nocturnal and spends the day in dense vegetation where it may be mobbed by other birds. Its large eyes are adapted for sight at low levels of light, and its hearing is also acute. It is a predator and at night often perches on a branch beside a glade or at the edge of woodland on the lookout for prey. When it detects a small moving object, it swoops down from its perch on silent wings and pounces on its target, which may be a small mammal, a bird, reptile or amphibian or a large beetle, grasshopper or other insect. It is known to prey on bats.
The mottled owl breeds between February and May in Colombia and in September to November in Argentina. It usually nests in a hole in a tree but may also choose an empty nest built by another species. One or usually two white eggs are laid and incubated by the female and both parents care for the young.


Cicindela aurulenta reaches about 15–18 millimetres (0.59–0.71 in) in length. C. aurulenta have polychromatic and iridescent external appearance. This species has blue-green elytra, with six large yellowish-white or bluish spots and two smaller spots on the shoulders. The medial large spot is somewhat more crescent-shaped compared to the anterior and posterior spots. Its elytra appears reddish-orange along the base and margins. When viewed dorsally, head and thorax are blue-green, with appearances of red-orange under certain light conditions due to its iridescent colouration. It has two prominent compound eyes and large, predominantly black mandibles with yellowish-white marking at its base. It has a labrum that's ivory-white with black base and margins. Its antennae is filiform. Its scape, pedicel, and first two flagellum are blue-green; while the remaining flagellum are dull black. It has pale colored hairs on the abdomen and legs.


C. aurulenta is widespread throughout the Indo-Malaya region. This species is distributed across Nepal, Bhutan, India, China, and Southeast Asia. It ranges as far south as the islands of Borneo, Sumatra, and the Lesser Sundas, such as Bali. Several subspecies variations co-occur within these regions.


C. aurulenta can be found in a wide range of sandy habitats, including riverbars, forest trails, and sand dunes near shorelines and mangroves.


Cicindela repanda, commonly known as the bronzed tiger beetle or common shore tiger beetle, is a species of tiger beetle that measures 10–13 millimetres (0.39–0.51 in) long, lives in most of North America. Its labrum is small with one tooth and the pronotum is coppery and hairy. The shoulder marking touches or nearly touches the middle band. It is usually seen in spring and summer and it lives in sand, gravel, or clay soil. Its food is many insects and some fruit. The species have a two-year life cycle. It can be found all across sand dunes around the Great Lakes. The species comprises three subspecies: C. repanda repanda, C. repanda novascotiae, and C. repanda tanneri.


The white stork (Ciconia ciconia) is a large bird in the stork family, Ciconiidae. Its plumage is mainly white, with black on the bird's wings. Adults have long red legs and long pointed red beaks, and measure on average 100–115 cm (39–45 in) from beak tip to end of tail, with a 155–215 cm (61–85 in) wingspan. The two subspecies, which differ slightly in size, breed in Europe (north to Finland), northwestern Africa, southwestern Asia (east to southern Kazakhstan) and southern Africa. The white stork is a long-distance migrant, wintering in Africa from tropical Sub-Saharan Africa to as far south as South Africa, or on the Indian subcontinent. When migrating between Europe and Africa, it avoids crossing the Mediterranean Sea and detours via the Levant in the east or the Strait of Gibraltar in the west, because the air thermals on which it depends for soaring do not form over water.
A carnivore, the white stork eats a wide range of animal prey, including insects, fish, amphibians, reptiles, small mammals and small birds. It takes most of its food from the ground, among low vegetation, and from shallow water. It is a monogamous breeder, but does not pair for life. Both members of the pair build a large stick nest, which may be used for several years. Each year the female can lay one clutch of usually four eggs, which hatch asynchronously 33–34 days after being laid. Both parents take turns incubating the eggs and both feed the young. The young leave the nest 58–64 days after hatching, and continue to be fed by the parents for a further 7–20 days.
The white stork has been rated as least concern by the International Union for Conservation of Nature (IUCN). It benefited from human activities during the Middle Ages as woodland was cleared, but changes in farming methods and industrialisation saw it decline and disappear from parts of Europe in the 19th and early 20th centuries. Conservation and reintroduction programs across Europe have resulted in the white stork resuming breeding in the Netherlands, Belgium, Switzerland, Sweden and the United Kingdom. It has few natural predators, but may harbour several types of parasite; the plumage is home to chewing lice and feather mites, while the large nests maintain a diverse range of mesostigmatic mites. This conspicuous species has given rise to many legends across its range, of which the best-known is the story of babies being brought by storks.


The white stork is a large bird. It has a length of 100–115 cm (39–45 in), and a standing height of 100–125 cm (39–49 in). The wingspan is 155–215 cm (61–85 in) and its weight is 2.3–4.5 kg (5.1–9.9 lb). Like all storks, it has long legs, a long neck and a long straight pointed beak. The sexes are identical in appearance, except that males are larger than females on average. The plumage is mainly white with black flight feathers and wing coverts; the black is caused by the pigment melanin. The breast feathers are long and shaggy forming a ruff which is used in some courtship displays. The irises are dull brown or grey, and the peri-orbital skin is black. The adult has a bright red beak and red legs, the colouration of which is derived from carotenoids in the diet. In parts of Spain, studies have shown that the pigment is based on astaxanthin obtained from an introduced species of crayfish (Procambarus clarkii) and the bright red beak colours show up even in nestlings, in contrast to the duller beaks of young white storks elsewhere.
As with other storks, the wings are long and broad enabling the bird to soar. In flapping flight its wingbeats are slow and regular. It flies with its neck stretched forward and with its long legs extended well beyond the end of its short tail. It walks at a slow and steady pace with its neck upstretched. In contrast, it often hunches its head between its shoulders when resting. Moulting has not been extensively studied, but appears to take place throughout the year, with the primary flight feathers replaced over the breeding season.
Upon hatching, the young white stork is partly covered with short, sparse, whitish down feathers. This early down is replaced about a week later with a denser coat of woolly white down. By three weeks, the young bird acquires black scapulars and flight feathers. On hatching the chick has pinkish legs, which turn to greyish-black as it ages. Its beak is black with a brownish tip. By the time it fledges, the juvenile bird's plumage is similar to that of the adult, though its black feathers are often tinged with brown, and its beak and legs are a duller brownish-red or orange. The beak is typically orange or red with a darker tip. The bills gain the adults' red colour the following summer, although the black tips persist in some individuals. Young storks adopt adult plumage by their second summer.


The nominate race of the white stork has a wide although disjunct summer range across Europe, clustered in the Iberian Peninsula and North Africa in the west, and much of eastern and central Europe, with 25% of the world's population concentrated in Poland, as well as parts of western Asia. The asiatica population of about 1450 birds is restricted to a region in central Asia between the Aral Sea and Xinjiang in western China. The Xinjiang population is believed to have become extinct around 1980. Migration routes extend the range of this species into many parts of Africa and India. Some populations adhere to the eastern migration route, which passes across Israel into eastern and central Africa.
A few records of breeding from South Africa have been known since 1933 at Calitzdorp, and about 10 birds have been known to breed since the 1990s around Bredasdorp. A small population of white storks winters in India and is thought to derive principally from the C. c. asiatica population as flocks of up to 200 birds have been observed on spring migration in the early 1900s through the Kurram Valley. However, birds ringed in Germany have been recovered in western (Bikaner) and southern (Tirunelveli) India. An atypical specimen with red orbital skin, a feature of the Oriental white stork, has been recorded and further study of the Indian population is required. North of the breeding range, it is a passage migrant or vagrant in Finland, Iceland, Ireland, Norway and Sweden, and west to the Azores and Madeira. In recent years, the range has expanded into western Russia.
The white stork's preferred feeding grounds are grassy meadows, farmland and shallow wetlands. It avoids areas overgrown with tall grass and shrubs. In the Chernobyl area of northern Ukraine, white stork populations declined after the 1986 nuclear accident there as farmland was succeeded by tall grass shrubs. In parts of Poland, poor natural foraging grounds have forced birds to seek food at rubbish dumps since 1999. White storks have also been reported foraging in rubbish dumps in the Middle East, North Africa and South Africa.
The white stork breeds in greater numbers in areas with open grasslands, particularly grassy areas which are wet or periodically flooded, and less in areas with taller vegetation cover such as forest and shrubland. They make use of grasslands, wetlands, and farmland on the wintering grounds in Africa. White storks were probably aided by human activities during the Middle Ages as woodland was cleared and new pastures and farmland were created, and they were found across much of Europe, breeding as far north as Sweden. The population in Sweden is thought to have established in the 16th century after forests were cut down for agriculture. About 5000 pairs were estimated to breed in the 18th century which declined subsequently. The first accurate census in 1917 found 25 pairs and the last pair failed to breed around 1955. The white stork has been a rare visitor to the British Isles, with about 20 birds seen in Britain every year, and prior to 2020 there were no records of nesting since a pair nested atop St Giles High Kirk in Edinburgh, Scotland in 1416. In 2020, a pair bred in the United Kingdom for the first time in over 600 years, as part of an re-introduction initiative called the White Stork Project.
A decline in population began in the 19th century due to industrialisation and changes in agricultural methods. White storks no longer nest in many countries, and the current strongholds of the western population are in Portugal, Spain, Ukraine and Poland. In the Iberian Peninsula, populations are concentrated in the southwest, and have also declined due to agricultural practices. A study published in 2005 found that the Podhale region in the uplands of southern Poland had seen an influx of white storks, which first bred there in 1931 and have nested at progressively higher altitudes since, reaching 890 m (3000 ft) in 1999. The authors proposed that this was related to climate warming and the influx of other animals and plants to higher altitudes. White storks arriving in Poznań province (Greater Poland Voivodeship) in western Poland in spring to breed did so some 10 days earlier in the last twenty years of the 20th century than at the end of the 19th century.


White storks fly south from their summer breeding grounds in Europe in August and September, heading for Africa. There, they spend the winter in savannah from Kenya and Uganda south to the Cape Province of South Africa. In these areas, they congregate in large flocks which may exceed a thousand individuals. Some diverge westwards into western Sudan and Chad, and may reach Nigeria.
In spring, the birds return north; they are recorded from Sudan and Egypt from February to April. They arrive back in Europe around late March and April, after an average journey of 49 days. By comparison, the autumn journey is completed in about 26 days. Tailwinds and scarcity of food and water en route (birds fly faster over regions lacking resources) increase average speed.
To avoid a long sea crossing over the Mediterranean, birds from central Europe either follow an eastern migration route by crossing the Bosphorus in Turkey, traversing the Levant, then bypassing the Sahara Desert by following the Nile valley southwards, or follow a western route over the Strait of Gibraltar. These migration corridors maximise help from the thermals and thus save energy.
In winter 2013–2014, white storks were observed in southern India's Mudumalai National Park for the first time.
The eastern route is by far the more important with 530,000 white storks using it annually, making the species the second commonest migrant there (after the European honey buzzard). The flocks of migrating raptors, white storks and great white pelicans can stretch for 200 km (125 mi). The eastern route is twice as long as the western, but storks take the same time to reach the wintering grounds by either.
Juvenile white storks set off on their first southward migration in an inherited direction but, if displaced from that bearing by weather conditions, they are unable to compensate, and may end up in a new wintering location. Adults can compensate for strong winds and adjust their direction to finish at their normal winter sites, because they are familiar with the location. For the same reason, all spring migrants, even those from displaced wintering locations, can find their way back to the traditional breeding sites. An experiment with young birds raised in captivity in Kaliningrad and released in the absence of wild storks to show them the way revealed that they appeared to have an instinct to fly south, although the scatter in direction was large.


The white stork is a gregarious bird; flocks of thousands of individuals have been recorded on migration routes and at wintering areas in Africa. Non-breeding birds gather in groups of 40 or 50 during the breeding season. The smaller dark-plumaged Abdim's stork is often encountered with white stork flocks in southern Africa. Breeding pairs of white stork may gather in small groups to hunt, and colony nesting has been recorded in some areas. However, groups among white stork colonies vary widely in size and the social structure is loosely defined; young breeding storks are often restricted to peripheral nests, while older storks attain higher breeding success while occupying the better quality nests toward the centres of breeding colonies. Social structure and group cohesion is maintained by altruistic behaviours such as allopreening. White storks exhibit this behaviour exclusively at the nest site. Standing birds preen the heads of sitting birds, sometimes these are parents grooming juveniles, and sometimes juveniles preen each other. Unlike most storks, it never adopts a spread-winged posture, though it is known to droop its wings (holding them away from its body with the primary feathers pointing downwards) when its plumage is wet.

A white stork's droppings, containing faeces and uric acid, are sometimes directed onto its own legs, making them appear white. The resulting evaporation provides cooling and is termed urohidrosis. Birds that have been ringed can sometimes be affected by the accumulation of droppings around the ring leading to constriction and leg trauma. The white stork has also been noted for tool use by squeezing moss in the beak to drip water into the mouths of its chicks.


The white stork breeds in open farmland areas with access to marshy wetlands, building a large stick nest in trees, on buildings, or on purpose-built man-made platforms. Each nest is 1–2 m (3.3–6.6 ft) in depth, 0.8–1.5 m (2.6–4.9 ft) in diameter, and 60–250 kg (130–550 lb) in weight. Nests are built in loose colonies. Not persecuted as it is viewed as a good omen, it often nests close to human habitation; in southern Europe, nests can be seen on churches and other buildings. The nest is typically used year after year especially by older males. The males arrive earlier in the season and choose the nests. Larger nests are associated with greater numbers of young successfully fledged, and appear to be sought after. Nest change is often related to a change in the pairing and failure to raise young the previous year, and younger birds are more likely to change nesting sites. Although a pair may be found to occupy a nest, partners may change several times during the early stages and breeding activities begin only after a stable pairing is achieved.
Several bird species often nest within the large nests of the white stork. Regular occupants are house sparrows, tree sparrows, and common starlings; less common residents include Eurasian kestrels, little owls, European rollers, white wagtails, black redstarts, Eurasian jackdaws, and Spanish sparrows. Paired birds greet by engaging in up-down and head-shaking crouch displays, and clattering the beak while throwing back the head. Pairs copulate frequently throughout the month before eggs are laid. High-frequency pair copulation is usually associated with sperm competition and high frequency of extra-pair copulation; however, extra-pair copulation is infrequent in white storks.
A white stork pair raises a single brood a year. The female typically lays four eggs, though clutches of one to seven have been recorded. The eggs are white, but often look dirty or yellowish due to a glutinous covering. They typically measure 73 mm × 52 mm (2.9 in × 2.0 in), and weigh 96–129 g (3.4–4.6 oz), of which about 11 g (0.39 oz) is shell. Incubation begins as soon as the first egg is laid, so the brood hatches asynchronously, beginning 33 to 34 days later. The first hatchling typically has a competitive edge over the others. While stronger chicks are not aggressive towards weaker siblings, as is the case in some species, weak or small chicks are sometimes killed by their parents. This behaviour occurs in times of food shortage to reduce brood size and hence increase the chance of survival of the remaining nestlings. White stork nestlings do not attack each other, and their parents' method of feeding them (disgorging large amounts of food at once) means that stronger siblings cannot outcompete weaker ones for food directly, hence parental infanticide is an efficient way of reducing brood size. Despite this, this behaviour has not commonly been observed.
The temperature and weather around the time of hatching in spring is important; cool temperatures and wet weather increase chick mortality and reduce breeding success rates. Somewhat unexpectedly, studies have found that later-hatching chicks which successfully reach adulthood produce more chicks than do their earlier-hatching nestmates. The body weight of the chicks increases rapidly in the first few weeks and reaches a plateau of about 3.4 kg (7.5 lb) in 45 days. The length of the beak increases linearly for about 50 days. Young birds are fed with earthworms and insects, which are regurgitated by the parents onto the floor of the nest. Older chicks reach into the mouths of parents to obtain food. Chicks fledge 58 to 64 days after hatching.
White storks generally begin breeding when about four years old, although the age of first breeding has been recorded as early as two years and as late as seven years. The oldest known wild white stork lived for 39 years after being ringed in Switzerland, while captive birds have lived for more than 35 years.


White storks consume a wide variety of animal prey. They prefer to forage in meadows that are within roughly 5 km (3 mi) of their nest and sites where the vegetation is shorter so that their prey is more accessible. Their diet varies according to season, locality and prey availability. Common food items include insects (primarily beetles, grasshoppers, locusts and crickets), earthworms, reptiles, amphibians, particularly frog species such as the edible frog (Pelophylax kl. esculentus) and common frog (Rana temporaria) and small mammals such as voles, moles and shrews. Less commonly, they also eat bird eggs and young birds, fish, molluscs, crustaceans and scorpions. They hunt mainly during the day, swallowing small prey whole, but killing and breaking apart larger prey before swallowing. Rubber bands are mistaken for earthworms and consumed, occasionally resulting in fatal blockage of the digestive tract.
Birds returning to Latvia during spring have been shown to locate their prey, moor frogs (Rana arvalis), by homing in on the mating calls produced by aggregations of male frogs.
The diet of non-breeding birds is similar to that of breeding birds, but food items are more often taken from dry areas. White storks wintering in western India have been observed to follow blackbuck to capture insects disturbed by them. Wintering white storks in India sometimes forage along with the woolly-necked stork (Ciconia episcopus). Food piracy has been recorded in India with a rodent captured by a western marsh harrier appropriated by a white stork, while Montagu's harrier is known to harass white storks foraging for voles in some parts of Poland. White storks can exploit landfill sites for food during the breeding season, migration period and winter.


White stork nests are habitats for an array of small arthropods, particularly over the warmer months after the birds arrive to breed. Nesting over successive years, the storks bring more material to line their nests and layers of organic material accumulate within them. Not only do their bodies tend to regulate temperatures within the nest, but excrement, food remains and feather and skin fragments provide nourishment for a large and diverse population of free-living mesostigmatic mites. A survey of twelve nests found 13,352 individuals of 34 species, the most common being Macrocheles merdarius, M. robustulus, Uroobovella pyriformis and Trichouropoda orbicularis, which together represented almost 85% of all the specimens collected. These feed on the eggs and larvae of insects and on nematodes, which are abundant in the nest litter. These mites are dispersed by coprophilous beetles, often of the family Scarabaeidae, or on dung brought by the storks during nest construction. Parasitic mites do not occur, perhaps being controlled by the predatory species. The overall effect of the mite population is unclear, the mites may have a role in suppressing harmful organisms (and hence be beneficial), or they may themselves have an adverse effect on nestlings.
The birds themselves host species belonging to more than four genera of feather mites. These mites, including Freyanopterolichus pelargicus, and Pelargolichus didactylus live on fungi growing on the feathers. The fungi found on the plumage may feed on the keratin of the outer feathers or on feather oil. Chewing lice such as Colpocephalum zebra tend to be found on the wings, and Neophilopterus incompletus elsewhere on the body.
The white stork also carries several types of internal parasites, including Toxoplasma gondii and intestinal parasites of the genus Giardia. A study of 120 white stork carcasses from Saxony-Anhalt and Brandenburg in Germany yielded eight species of trematode (fluke), four cestode (tapeworm) species, and at least three species of nematode. One species of fluke, Chaunocephalus ferox, caused lesions in the wall of the small intestine in a number of birds admitted to two rehabilitation centres in central Spain, and was associated with reduced weight. It is a recognised pathogen and cause of morbidity in the Asian openbill (Anastomus oscitans). More recently, the thorough study performed by J. Sitko and P. Heneberg in the Czech Republic in 1962–2013 suggested that the central European white storks host 11 helminth species. Chaunocephalus ferox, Tylodelphys excavata and Dictymetra discoidea were reported to be the dominant ones. The other species found included Cathaemasia hians, Echinochasmus spinulosus, Echinostoma revolutum, Echinostoma sudanense, Duboisia syriaca, Apharyngostrigea cornu, Capillaria sp. and Dictymetra discoidea. Juvenile white storks were shown to host less species, but the intensity of infection was higher in the juveniles than in the adult storks.
West Nile virus (WNV) is mainly a bird infection that is transmitted between birds by mosquitos. Migrating birds appear to be important in spread of the virus, the ecology of which remains poorly known. On 26 August 1998, a flock of about 1,200 migrating white storks that had been blown off course on their southward journey landed in Eilat, in southern Israel. The flock was stressed as it had resorted to flapping flight to return to its migratory route, and a number of birds died. A virulent strain of West Nile virus was isolated from the brains of eleven dead juveniles. Other white storks subsequently tested in Israel have shown anti-WNV antibodies. In 2008 three juvenile white storks from a Polish wildlife refuge yielded seropositive results indicating exposure to the virus, but the context or existence of the virus in Poland is unclear.


The white stork's decline due to industrialisation and agricultural changes (principally the draining of wetlands and conversion of meadows to crops such as maize) began in the 19th century: the last wild individual in Belgium was seen in 1895, in Sweden in 1955, in Switzerland in 1950 and in the Netherlands in 1991. However, the species has since been reintroduced to many regions. It has been rated as least concern by the IUCN since 1994, after being evaluated as near threatened in 1988. The white stork is one of the species to which the Agreement on the Conservation of African-Eurasian Migratory Waterbirds (AEWA) applies. Parties to the agreement are required to engage in a wide range of conservation strategies described in a detailed action plan. The plan is intended to address key issues such as species and habitat conservation, management of human activities, research, education, and implementation. Threats include the continued loss of wetlands, collisions with overhead power lines, use of persistent pesticides (such as DDT) to combat locusts in Africa, and largely illegal hunting on migration routes and wintering grounds.
A large population of white storks breeds in central (Poland, Ukraine and Germany) and southern Europe (Spain and Turkey). In a 2004/05 census, there were 52,500 pairs in Poland, 30,000 pairs in Ukraine, 20,000 pairs in Belarus, 13,000 pairs in Lithuania (the highest known density of this species in the world), 10,700 pairs in Latvia, and 10,200 in Russia. There were around 5,500 pairs in Romania, 5,300 in Hungary and an estimated 4,956 breeding pairs in Bulgaria. In Germany, the majority of the total 4,482 pairs were in the eastern region, especially in the states of Brandenburg and Mecklenburg-Vorpommern (1296 and 863 pairs in 2008 respectively). Apart from Spain and Portugal (33,217 and 7,684 pairs in 2004/05 respectively), populations are generally much less stable. In the eastern Mediterranean region Turkey has a sizeable population of 6,195 pairs, and Greece 2,139 pairs. In Western Europe the white stork remains a rare bird despite conservation efforts. In 2004 France had only 973 pairs, and the Netherlands 528 pairs. In Denmark, the species had consistently bred since the 15th century, peaking at several thousands pairs around 1800. Afterwards it began declining mainly due to habitat loss (especially conversion of wetlands and meadows into modern farming), with only a few tens of breeding pairs in 1974 and none in 2008. Since then, it has reestablished itself and the population has slowly started to increase, reaching 8 pairs in 2022. In Armenia the population of the white stork slightly increased in the period between 2005 and 2015, and by last data reached 652 pairs.
In the early 1980s, the population had fallen to fewer than nine pairs in the entire upper Rhine River valley, an area closely identified with the white stork for centuries. Conservation efforts successfully increased the population of birds there to 270 pairs (in 2008), largely due to the actions of the Association for the Protection and Reintroduction of Storks in Alsace and Lorraine. The reintroduction of zoo-reared birds has halted further declines in Italy, the Netherlands, and Switzerland. There were 601 pairs breeding in Armenia and around 700 pairs in the Netherlands in 2008, and few pairs also breed in South Africa, typically recent colonists from within the normal wintering population. In Poland, electric poles have been modified with a platform at the top to prevent the white stork's large nest from disrupting the electricity supply, and sometimes nests are moved from an electric pole to a man-made platform. Introductions of zoo-reared birds in the Netherlands has been followed up by feeding and nest-building programs by volunteers. Similar reintroduction programs are taking place in Sweden, and Switzerland, where 175 pairs were recorded breeding in 2000. Long-term viability of the population in Switzerland is unclear as breeding success rates are low, and supplementary feeding does not appear to be of benefit. However, as of 2017, 470 adults and 757 young ones were recorded in Switzerland. Historically, the species' northern breeding limit was at Estonia, but it has moved slowly northwards (possibly due to warmer temperatures) into Karelia and in 2015 the first ever known breeding happened in Finland.
In August 2019, 24 juveniles were released at the Knepp Estate in West Sussex, and others at a site near Tunbridge Wells and at the Wintershall Estate, near Godalming, as part of a project to reintroduce the white stork as a breeding species in South East England, for the first time since 1416. In 2020, the program was successful with the birth of five baby storks.


The woolly-necked stork or white-necked stork (Ciconia episcopus) is a large wading bird in the stork family Ciconiidae.  It breeds singly, or in small loose colonies. It is distributed in a wide variety of habitats including marshes in forests, agricultural areas, and freshwater wetlands across Asia and Africa.


It is a widespread tropical species which breeds in Asia, from India to Indonesia, and across much of western, eastern and south-central Africa. It is a resident breeder building nests on trees located on agricultural fields or wetlands, on natural cliffs, and on cell phone towers. They use a variety of freshwater wetlands including seasonal and perennial reservoirs and marshes, crop lands, irrigation canals and rivers, but are mostly seen in agricultural areas and in wetlands outside protected areas across south Asia and Myanmar. They are attracted to fires in grasslands and crop fields where they capture insects trying to escape the fire. They use ponds and marshes inside forests in both Africa and Asia, especially in south-east Asia where they use grassy and marshy areas in clearings in several forest types. In India, they are an uncommon species in coastal habitats. They use coastal areas in Africa also, with birds in Sulawesi observed to be eating sea snakes, and birds on the Kenya coast foraging in coral reefs and mudflats. In an agricultural landscape in north India, woolly-necked storks preferred fallow fields during the summer and monsoon seasons, and natural freshwater wetlands during the winter. Here, irrigation canals were preferentially used during winters when water levels were low, and birds avoided crop fields in all seasons. Assisted by construction of new irrigation canals, this species is spreading to arid areas like the Thar Desert in Rajasthan, India. Across south Asia, woolly-necked storks largely use agricultural landscapes with more numbers seen using unprotected wetlands relative to the amount of wetlands on the landscape, and a majority of individuals use agricultural crops. In KwaZulu Natal, South Africa, they are accustomed to people feeding them, and nest on exotic tree species in sub-urban areas. In Haryana, north India, they nest on trees planted along crop fields and irrigation canals as part of traditional multifunctional agroforestry and generally avoid trees close to human settlements.
Individuals of this species have been sighted at altitudes of 3,790 m above sea level in China (Napahai wetland), and 3,540 m above sea level in Nepal (Annapurna Conservation Area).


The woolly-necked stork walks slowly and steadily on the ground seeking its prey, which like that of most of its relatives, consists of amphibians, reptiles and insects. In suburban South Africa, nestlings were provisioned largely with guttural toad (Sclerophrys gutturalis), but also with common river frogs (Amietia queckettii), fish, snakes, crabs and moles (Amblysomus sp.). Despite being provided with supplementary foods by people in South Africa, nestling diet was largely (>60%) natural animal species. More than two adult birds provisioned chicks of one nest in South Africa providing the first known evidence of cooperative behaviour in woolly-necked storks. Death of two nestlings was attributed to provisioning of processed foods that people fed adult birds.


Typically, a large stick nest is built on a tree, and clutch size is two to six eggs, with five and six eggs being less common. Birds use both forest trees and scattered trees in agricultural areas to build nests. In India, some nests have been being observed in or near urban areas on cell phone towers, but such nesting on artificial human-made structures is not a regular occurrence. Riverside cliffs are occasionally used for nesting. In South Africa, woolly-necked storks nested largely on trees in suburban areas such as gardens with nests largely placed on exotic tree species such as Pinus elliottii, Eucalyptus sp., Melia azedarach, Cinnamommum camphora, and Jacaranda mimosifolia. Very few nests were built on native trees such as Trichilia dregeana, Ficus burkei and Syderoxylon inerme. Three of 30 nests in South Africa were built on anthropogenic structures: one on a roof top of a two-story building, one in an unusual nest box, and one atop an electric pole.
In Haryana, north India, nesting woolly-necked storks used trees close to irrigation canals and far from human habitation for nesting, and were not affected by the presence of natural wetlands and relatively larger patches of trees on the landscape. Very few nests each year were placed on artificial structures such as electricity pylons, and the majority were placed on Dalbergia sissoo, Ficus religiosa and Eucalyptus sp. In Haryana's agricultural landscape, small numbers of woolly-necked stork nests were also found on Acacia nilotica, Azadirachta indica, Mangifera indica, Mitragyna parviflora, Syzhygium cumini and Tectona gradis. Woolly-necked Storks reused over 44% of nest sites for multiple years. Brood size of 42 successful nests in Haryana was relatively high with over three chicks successfully fledging from nests, and a small number of nests each year fledging four and five chicks each with six chicks fledging from one nest. Detailed observations of breeding habits in South Africa and north India suggest that the woolly-necked stork is not an obligate wetland species unlike other stork species that locate their nests close to wetlands.
Two nests of woolly-necked storks were reused after stork chicks had fledged by black kites Milvus migrans in lowland Nepal. Woolly-necked stork nests in Haryana were preferentially reused by dusky eagle-owls Bubo coromandus over other large nests made by other bird species in the area. It seems likely that woolly-necked storks support the well-being of many other species, including large raptors such as dusky eagle-owls, via commensal and other inter-species relationships associated with their nests.


The woolly-necked stork is one of the species to which the Agreement on the Conservation of African-Eurasian Migratory Waterbirds (AEWA) applies. The species was elevated from "Near-threatened" to "Vulnerable" in 2014 based on anecdotal reports of deforestation in south-east Asia potentially leading to catastrophic population declines with the assumption that the species required protected wetlands inside forested reserves. It was, however, down listed to "Near-threatened" in 2019 after concrete evidence emerged from south Asia and Myanmar that the erstwhile population estimate was a severe underestimate, and that most woolly-necked storks used agricultural areas and unprotected wetlands, with abundances being lower inside forested reserves. Agricultural landscapes in north India support considerable numbers of breeding pairs that have relatively large brood sizes and behaviors similar to storks in protected managed reserves suggesting that this species is not an obligate wetland bird and that it is not reliant on undisturbed protected wetlands and forest reserves. An earlier "guesstimate" of the south and south-east Asian population of woolly-necked storks of 25,000 has been revised upwards to an estimated > 2,00,000 storks in south Asia alone.
There have been severe declines in numbers of woolly-necked storks along the Mekong River in Cambodia suggesting that the species is indeed threatened in south-east Asia, perhaps much more than anywhere else in its wide distribution range, likely due to indiscriminate hunting. Modeled distributions of woolly-necked storks strongly overlapped forested reserves in south-east Asia suggesting low ability to survive outside forests in stark contrast to the situation in south Asia and Myanmar. The majority of the protected reserve forests and wetlands in south-east Asia are under threat suggesting that woolly-necked storks face an uncertain future in this region.


The maguari stork (Ciconia maguari) is a large species of stork that inhabits seasonal wetlands over much of South America, and is very similar in appearance to the white stork; albeit slightly larger. It is the only species of its genus to occur in the New World and is one of the only three New World stork species, together with the wood stork and the jabiru.


A distinct characteristic unique to this stork species is intermediately dark plumage in the young, which persists for most of the nestling period. In the chicks’ first few days after hatching, their sparse down is snowy white. The chicks subsequently undergo two basic moults before entering adulthood. The first moult usually begins after four days, in which black semi-plumes on the head and neck begin to emerge; followed shortly by the emergence of greyish-black down feathers over the body after one week of age. Some white down feathers initially remain attached to give a temporary mottled black and white appearance before finally receding to leave a darkish grey downy plumage. During development of the dark down, the bill, legs and feet are shiny black. A pale yellow stripe extends up the venter, the gular pouch is bright orange, and the iris is dark brown. At hatching, nestlings weigh 76-90g.
After the chicks pass their third week of age, the black down develops olive streaks and the only parts of the body that do not appear dark are the bright orange gular skin and a small patch of white down above and below the tail. Just over 10 days later, black flight feathers develop, shortly followed by black contour feathers. At this point, the chicks are fledged and have their first full juvenile plumage after about a month of hatching. Hence, juveniles remain black at fledging, by which time the orange of the gular pouch has also turned pale scarlet.
In the second moult, white down begins to return when the chicks are seven weeks old, followed by white semi-plumes then white contour feathers. The first white basic plumage is complete after three months and resembles that of adults. By this time, the legs and feet are now an incipient pink and the bill has become bicoloured (pale blue and maroon). Although the young now resemble adults, the skin surrounding the eye remains black for about a year before turning red, and the dark brown iris does not begin to yellow until after two years old. These are two potentially useful features that identify juvenile maguari storks; but in the field, the dark iris of the juveniles is the most reliable feature that distinguishes them from adults.
Two hypotheses have been proposed to explain the dark colouration in the juveniles. First, the blackish feathers may serve as camouflage against both ground and aerial predators. Alternatively, the black plumage may serve a thermoregulatory function by enhancing its absorption of the sun's radiation. This could arguably be advantageous because this stork breeds during the winter in the part of its range in the southern hemisphere, so that the chicks can be exposed to low environmental temperatures. However, Thomas  regards the anti-predator explanation for the dark plumage to be more plausible.


Unlike other stork species, this stork is commonly found to nest on the ground, whereas many other stork species habitually nest at higher elevation. The nest always lies near to shallow water amongst tall grass and reeds, since aquatic organisms form the bulk of the nestlings’ diet. The maguari stork's nest is also unusual in extensively comprising grass and reeds. Common species used in nest fabrication include the reed Cyperus giganteus and the marsh grass Zizianopsis bonariensis, alongside other aquatic plants in the families Polygonaceae and Solinaceae. Ground nests found in the southern parts of the maguari stork's range are one-meter-high (3.3 ft) conical structures measuring 1.5–2.5 m in basal diameter, tapering to a flat platform of 1–1.5 m at the top. The nest rises up to two feet above the surface of the water and its location is largely treeless. Hence overall, the maguari stork's nest seems to more closely resemble that of cranes and screamers such as the southern screamer Chauna torquata than that of other stork species.
In the Venezuelan llanos, the maguari stork is also found to nest in low, thick-trunked trees including Ficus pertusa and Randia venezuelansis. These nests comprise sticks that are usually less than a metre long and less than 2 cm in diameter, and often from the palm species Copernicia tectorum. Nevertheless, the species does not seem to naturally perch in trees because unlike its sympatric heterospecifics such as the jabiru, it cannot stand on its hallux. Therefore, it probably only nests opportunistically above the ground. The same nest may be used by a pair in successive years, sometimes for as long as seven years. However, ground nests composed of herbaceous plants usually disintegrate after a year, whereupon the same breeding pair returns to the nest site to rebuild the nest. Both partners participate in building and lining the nest, which continues throughout incubation and brooding. Nest lining usually begins when the base is about a metre wide, and lining material largely consists of wet grass that dries and hardens in the intense sun.
The maguari stork is primarily a colonial nester, although it also less frequently nests solitarily. Solitary nests are found to be less successful than colonial nests in terms of survival of the young, but survival rates in the former appear to be more consistent. Colonies typically consist of 5-15 nests, some of which are within 50 cm apart in tree colonies, but at different heights. One colony from a study on the llanos comprised as many as 40 nests. Colonial and solitary nesters also differ in the way they defend their nests. Colonial nesters are more aggressive than solitary nesters and physically attack intruders with bill jabbing. Contrarily, solitary nesters use a "lead-away" strategy, in which the nesting bird walks up behind the intruder with threatening bill clattering, and ceases to clatter its bill when the intruder moves away from the nest.


Breeding is highly synchronised with the onset of rain during the wet season, which usually lasts from May to November in the llanos of Venezuela. However, most breeding of the maguari stork happens from July to mid-September, which is earlier than breeding in both the sympatric wood stork and the jabiru. Individuals begin migration to the breeding grounds as the rains begin, although some individuals may migrate and arrive early just before the rains. Despite the lengthy wet season in the llanos, the nesting period in one year never lasts for over four months. Elsewhere in the stork's global range, the timing of the breeding season is slightly different and somewhat shorter. In north east Argentina for example, the breeding season extends from June to August; and in east central Argentina extends from July to October, with eggs probably being laid in late June and early July. On Isla Mexiana in eastern Brazil, the breeding season lasts only from August to September. The timing of onset of the seasonal rain is extremely variable across the maguari stork's global range, with rainfall beginning in some years as early as late March and others as late as June. The start of breeding is therefore correspondingly variable; and in Argentina, nesting may commence as late as August with late rains. In the llanos, breeders may lay eggs as early as late May following unusually early and heavy rainfall; whereas young breeders may lay eggs as late as October with late-arriving rain.
The clutch size is typically 3 or 4 eggs, with an average of 3.2 being reported. Eggs are laid on alternate days, so that hatching within clutches is highly asynchronous; with some young hatching up to a week apart. Incubation begins after the 2nd or 3rd egg is laid, is carried out by both parents, and lasts 29–32 days. The eggs are oval or subelliptical, and mean egg measurements are 75.19mm in length and 52.56mm in width; with maximum egg measurements of 77.4mm in length and 56.2mm in breadth having been recorded. The eggs are also said to be disproportionately small compared to the laying female's body mass.
Weight differences between different-aged siblings have been recorded as 500-1400 g. Egg laying is however highly synchronous between nests in a colony, so that relatively large groups of young from different nests fledge together in batches toward the end of the wet season. This may serve as an anti-predator strategy to dilute an individual's risk of being predated. As adults, male maguari storks become sexually mature at three years of age, and females at four years.
After three weeks of age, Maguari stork nestlings develop defensive behaviour if their nest is approached by an intruder; which is not known for nestlings of other stork species. They crouch forward, partially spread their wings and erect the black feathers on their head, neck and back; followed by a shrill, rasping scream and an attempt to grab a persistent intruder with their bill. In many other stork species, akinesia lasts throughout much of the nestlings’ early life, and the constantly white plumage makes them appear as eggs to potential chick predators. However, in maguari stork nestlings, akinesia ceases much earlier, and the unusual aggressiveness of the young has probably developed as a specialised anti-predator strategy in compensation for the young chicks’ inability to leave the nest due to their slow-developing hallux and the vulnerable position of the nest on the ground. However, at 25–35 days old, the hallux is sufficiently developed to allow the chicks to leave ground nests in search of food. Chicks also occasionally beg for food from their parents outside the nest, but parents have never been observed to feed their young outside the nest.
There appears to be little sibling rivalry in maguari stork nestings, with most nestling mortality being due to young falling out of nests containing 3-4 chicks; and egg mortality primarily through predation appears to be higher than chick mortality.
The large congregations of maguari storks in the pre- and post-reproductive periods are considered a reliable indicator that this stork breeds locally in the area in which the groups are sighted.


The primary threats to this species are anthropogenic habitat disturbance and hunting for food. A common human disturbance occurs through habitat destruction via land reclamation from marshes for agriculture, which has occurred especially in southeast Brazil and may therefore evoke conservation concern for the species in this area. Claiming land for agriculture by digging canals, together with land filling and sewage discharges may also threaten dry-season foraging environments for maguari storks especially on the northern Rio de Janeiro coast. The maguari stork is vulnerable to nesting habitat destruction because it shows nest-site fidelity, and will continue to nest in the same place even after onset of recent anthropogenic disturbance. Use of pesticides may also adversely affect the health and breeding success of this species. The capture of individuals for food presents another survival threat and occurs particularly in the Southern Amazon and Venezuela.
Natural enemies of this stork include boa constrictors and crested caracaras (Polyborus plancus), both of which eat this species’ eggs. Both of these predatory species can probably also eat maguari stork chicks that are no more than a few weeks old. Many other potential predators such as jaguars, crocodilians, Pampas cats and maned wolves could also access ground nests.  Although these animals are known to opportunistically feed on birds, no instances of predation on maguari storks have been hitherto recorded; but such predation is considered likely.
The maguari stork is potentially threatened in the Pantanal, which as well as being subject to considerable land reclamation for agriculture, has hosted increased operation of hydroelectric dams, especially in the Parana river basin. Dams retain much water during the dry season, so that natural water features downstream are more likely to dry out completely and thereby lead to a decrease in suitable foraging sites for this stork. Conversely, during the wet season, dams can lead to extensive flooding downstream caused by the release of a large volume of water at once that renders the storks’ usual foraging grounds too deep for them to stand in.


The black stork (Ciconia nigra) is a large bird in the stork family Ciconiidae. It was first described by Carl Linnaeus in the 10th edition of his Systema Naturae. Measuring on average 95 to 100 cm (37 to 39 in) from beak tip to end of tail with a 145-to-155 cm (57-to-61 in) wingspan, the adult black stork has mainly black plumage, with white underparts, long red legs and a long pointed red beak. A widespread but uncommon species, it breeds in scattered locations across Europe (predominantly in Portugal and Spain, and central and eastern parts), and east across the Palearctic to the Pacific Ocean. It is a long-distance migrant, with European populations wintering in tropical Sub-Saharan Africa, and Asian populations in the Indian subcontinent. When migrating between Europe and Africa, it avoids crossing the Mediterranean Sea and detours via the Levant in the east or the Strait of Gibraltar in the west. An isolated, non-migratory, population occurs in Southern Africa.
Unlike the closely related white stork, the black stork is a shy and wary species. It is seen singly or in pairs, usually in marshy areas, rivers or inland waters. It feeds on amphibians, small fish and insects, generally wading slowly in shallow water stalking its prey. Breeding pairs usually build nests in large forest trees—most commonly deciduous but also coniferous—which can be seen from long distances, as well as on large boulders, or under overhanging ledges in mountainous areas. The female lays two to five greyish-white eggs, which become soiled over time in the nest. Incubation takes 32 to 38 days, with both sexes sharing duties, and fledging takes 60 to 71 days.
The black stork is considered to be a species of least concern by the International Union for Conservation of Nature, but its actual status is uncertain. Despite its large range, it is nowhere abundant, and it appears to be declining in parts of its range, such as in India, China and parts of Western Europe, though increasing in others such as the Iberian Peninsula. Various conservation measures have been taken for the black stork, like the Conservation Action Plan for African black storks by Wetlands International. It is also protected under the African-Eurasian Waterbird Agreement and the Convention on International Trade in Endangered Species of Wild Fauna and Flora.


English naturalist Francis Willughby wrote about the black stork in the 17th century, having seen one in Frankfurt. He named it Ciconia nigra, from the Latin words for "stork" and "black" respectively. It was one of the many species originally described by Swedish zoologist Carl Linnaeus in the landmark 1758 10th edition of his Systema Naturae, where it was given the binomial name of Ardea nigra. It was moved to the new genus Ciconia by French zoologist Mathurin Jacques Brisson two years later. The word stork is derived from the Old English word storc, thought to be related to the Old High German storah, meaning "stork", and the Old English stearc, meaning "stiff".
The black stork is a member of the genus Ciconia, or typical storks, a group of seven extant species, characterised by straight bills and mainly black and white plumage. The black stork was long thought to be most closely related to the white stork (C. ciconia). However, genetic analysis via DNA–DNA hybridization and mitochondrial cytochrome b DNA by Beth Slikas in 1997 found that it was basal (an early offshoot) in the genus Ciconia. Fossil remains have been recovered from Miocene beds on Rusinga and Maboko Islands in Kenya, which are indistinguishable from the white and black storks.


The black stork is a large bird, measuring between 95 and 100 cm (37 and 39 in) in length with a 145-to-155 cm (57-to-61 in) wingspan, and weighing around 3 kg (6.6 lb). Standing as tall as 102 cm (40 in), it has long red legs, a long neck and a long, straight, pointed red beak. It bears some resemblance to Abdim's stork (C. abdimii), which can be distinguished by its much smaller build, predominantly green bill, legs and feet, and white rump and lower back. The plumage is black with a purplish green sheen, except for the white lower breast, belly, armpits, axillaries and undertail coverts. The breast feathers are long and shaggy, forming a ruff which is used in some courtship displays. The black stork has brown irises, and bare red skin around its eyes. The sexes are identical in appearance, except that males are larger than females on average. Moulting takes place in spring, with the iridescent sheen brighter in new plumage. It walks slowly and steadily on the ground and like all storks, it flies with its neck outstretched.
The juvenile resembles the adult in plumage, but the areas corresponding to the adult black feathers are browner and less glossy. The scapulars, wing and upper tail coverts have pale tips. The legs, bill and bare skin around the eyes are greyish green. It could possibly be confused with the juvenile yellow-billed stork, but the latter has paler wings and mantle, a longer bill and white under the wings.


During the summer, the black stork is found from Eastern Asia (Siberia and northern China) west to Central Europe, reaching Estonia in the north, Poland, Lower Saxony and Bavaria in Germany, the Czech Republic, Hungary, Italy and Greece in the south, with an outlying population in the central-southwest region of the Iberian Peninsula (Extremadura and surrounding provinces of Spain, plus Portugal). It is migratory, wintering in tropical Africa and Asia, although certain populations of black storks are sedentary or dispersive. An isolated population exists in Southern Africa, where the species is more numerous in the east, in eastern South Africa and Mozambique, and is also found in Zimbabwe, Eswatini, Botswana and less commonly Namibia.
Most of the black storks that summer in Europe migrate to Africa, with those from western Germany and points west heading south via the Iberian Peninsula and the rest via Turkey and the Levant. Those flying via Spain spend winter in the Falémé River basin of eastern Senegal, Guinea, southern Mauritania, Ivory Coast, Sierra Leone and western and central Mali, while those flying via the Sinai end up in northern Ethiopia, the Kotto River basin in the Central African Republic, the Mbokou river basin in Chad and northeastern Nigeria. Black storks summering in western Asia migrate to northern and northeastern India, ranging mainly from Punjab south to Karnataka, and Africa. They are occasional visitors to Sri Lanka. Those summering further east in eastern Russia and China winter mainly in southern China, and occasionally in Hong Kong, Myanmar, northern Thailand, and Laos. They were first recorded in western Myanmar in 1998.
The black stork prefers more wooded areas than the better-known white stork, and breeds in large marshy wetlands with interspersed coniferous or broadleaved woodlands, but also inhabits hills and mountains with sufficient networks of creeks. It usually inhabits ponds, rivers, edges of lakes, estuaries and other freshwater wetlands. The black stork does inhabit more agricultural areas in the Caspian lowlands, but even here it avoids close contact with people. Its wintering habitat in India comprises reservoirs or rivers with nearby scrub or forest, which provide trees that black storks can roost in at night. In southern Africa it is found in shallow water in rivers or lakes, or swamps, but is occasionally encountered on dry land.
After disappearing from Belgium before the onset of the 20th century, it has returned to breed in the Belgian Ardennes, Luxembourg and Burgundy, France, by 2000. It appears to be increasing in numbers in Spain and Portugal, where the population was estimated at 405 to 483 pairs in 2006. The black stork is a rare vagrant to the British Isles, turning up in the warmer months—particularly in spring—generally in the south and east. Sightings have become more common since the 1970s as its breeding range moves northwards. It has been recorded in Scotland six times between 1946 and 1983, including from Shetland, Orkney and the Highlands, as well as the Scottish Borders (Peebles). It is not abundant in the western parts of its distribution, but more densely inhabits eastern Transcaucasia. Further east, it has been recorded from locations across Iran, though little is known about its habits there; breeding has been recorded from near Aliabad in Fars province, Khabr National Park in Kerman province, Karun river in Khuzestan province, Qaranqu River in East Azarbaijan province, and Aliabad river in Razavi Khorasan province. The population has declined in Iran due to draining of wetlands. East of the Ural Mountains, the black stork is patchily found in forested and mountainous areas up to 60°‒63° N across Siberia to the Pacific Ocean. South of Siberia, it breeds in Xinjiang, northwestern China, northern Mongolia south to the Altai Mountains, and northeastern China south to the vicinity of Beijing. In the Korean Peninsula, the black stork is an uncommon summer visitor, no longer breeding in the south since 1966. Birds have been seen in the northeast but it is not known whether they breed there. Similarly it has been seen in the summer in Afghanistan, but its breeding status is uncertain.