The Roman Empire was split in 395 upon the death of Theodosius I into an Eastern and Western Roman Empire in part because of the increasing pressure from threats during the Barbarian Invasions. From the 6th century into the 7th century, the Slavs crossed the Danube and largely absorbed the indigenous Ancient Greeks, Illyrians and Thracians in the Balkans; thus, the Illyrians were mentioned for the last time in historical records in the 7th century.
In the 11th century, the Great Schism formalised the break of communion between the Eastern Orthodox and Western Catholic Church that is reflected in Albania through the emergence of a Catholic north and Orthodox south. The Albanian people inhabited the west of Lake Ochrida and the upper valley of River Shkumbin and established the Principality of Arbanon in 1190 under the leadership of Progon of Kruja. The realm was succeeded by his sons Gjin and Dhimitri.
Upon the death of Dhimiter, the territory came under the rule of the Albanian-Greek Gregory Kamonas and subsequently under the Golem of Kruja. In the 13th century, the principality was dissolved. Arbanon is considered to be the first sketch of an Albanian state, that retained a semi-autonomous status as the western extremity of the Byzantine Empire, under the Byzantine Doukai of Epirus or Laskarids of Nicaea.
Towards the end of the 12th and beginning of the 13th centuries, Serbs and Venetians started to take possession over the territory. The ethnogenesis of the Albanians is uncertain; however the first undisputed mention of Albanians dates back in historical records from 1079 or 1080 in a work by Michael Attaliates, who referred to the Albanoi as having taken part in a revolt against Constantinople. At this point the Albanians were fully Christianised.
Few years after the dissolution of Arbanon, Charles of Anjou concluded an agreement with the Albanian rulers, promising to protect them and their ancient liberties. In 1272, he established the Kingdom of Albania and conquered regions back from the Despotate of Epirus. The kingdom claimed all of central Albania territory from Dyrrhachium along the Adriatic Sea coast down to Butrint. A catholic political structure was a basis for the papal plans of spreading Catholicism in the Balkan Peninsula. This plan found also the support of Helen of Anjou, a cousin of Charles of Anjou. Around 30 Catholic churches and monasteries were built during her rule mainly in northern Albania. Internal power struggles within the Byzantine Empire in the 14th century enabled Serbs' most powerful medieval ruler, Stefan Dusan, to establish a short-lived empire that included all of Albania except Durrës. In 1367, various Albanian rulers established the Despotate of Arta. During that time, several Albanian principalities were created, notably the Principality of Albania, Principality of Kastrioti, Lordship of Berat and Principality of Dukagjini. In the first half of the 15th century, the Ottoman Empire invaded most of Albania, and the League of Lezhë was held under Skanderbeg as a ruler, who became the national hero of the Albanian medieval history.


Albania (/ælˈbeɪniə, ɔːl-/ (listen) a(w)l-BAY-nee-ə; Albanian: Shqipëri or Shqipëria), officially the Republic of Albania (Albanian: Republika e Shqipërisë), is a country in Southeastern Europe. It is located on the Adriatic and Ionian Seas within the Mediterranean Sea and shares land borders with Montenegro to the northwest, Kosovo to the northeast, North Macedonia to the east and Greece to the south. Tirana is its capital and largest city, followed by Durrës, Vlorë, and Shkodër.
Albania displays varied climatic, geological, hydrological, and morphological conditions, defined in an area of 28,748 km² (11,100 sq mi). It possesses significant diversity with the landscape ranging from the snow-capped mountains in the Albanian Alps as well as the Korab, Skanderbeg, Pindus and Ceraunian Mountains to the hot and sunny coasts of the Albanian Adriatic and Ionian Sea along the Mediterranean Sea.
Albania has been inhabited by different civilisations over time, such as the Illyrians, Thracians, Greeks, Romans, Byzantines, Venetians, and Ottomans. The Albanians established the autonomous Principality of Arbër in the 12th century. The Kingdom of Albania and Principality of Albania formed between the 13th and 14th centuries. Prior to the Ottoman conquest of Albania in the 15th century, the Albanian resistance to Ottoman expansion into Europe led by Skanderbeg won them acclaim over most of Europe. Albania remained under Ottoman rule for nearly five centuries, during which many Albanians (known as Arnauts) attained high-ranking offices in the empire, especially in the Southern Balkans and Egypt. Between the 18th and 19th centuries, cultural developments, widely attributed to Albanians having gathered both spiritual and intellectual strength, conclusively led to the Albanian Renaissance. After the defeat of the Ottomans in the Balkan Wars, the modern nation state of Albania declared independence in 1912. In the 20th century, the Kingdom of Albania was invaded by Italy, which formed Greater Albania before becoming a protectorate of Nazi Germany. Enver Hoxha formed the People's Socialist Republic of Albania after World War II, modeled under the terms of Hoxhaism. The Revolutions of 1991 concluded the fall of communism in Albania and eventually the establishment of the current Republic of Albania.
Albania is a unitary parliamentary constitutional republic and a developing country with an upper-middle income economy dominated by the service sector, followed by manufacturing. It went through a process of transition following the end of communism in 1990, from centralised planning to a market-based economy. Albania provides universal health care and free primary and secondary education to its citizens. Albania is a member of the United Nations, World Bank, UNESCO, NATO, WTO, COE, OSCE, and OIC. It has been an official candidate for membership in the European Union since 2014. It is one of the founding members of the Energy Community, including the Organization of the Black Sea Economic Cooperation and Union for the Mediterranean.


In ancient times, the incorporated territory of Albania was historically inhabited by Indo-European peoples, among them numerous Illyrian tribes, Ancient Greeks and Thracians. In view of the Illyrian tribes, there is no evidence that these tribes used any collective nomenclature for themselves, while it is regarded to be unlikely that they used a common endonym. The endonym Illyrians seems to be the name applied to a specific Illyrian tribe, which was the first to come in liaison with the Ancient Greeks resulting in the endonym Illyrians to be applied pars pro toto to all people of similar language and customs.
The territory referred to as Illyria corresponded roughly to the area east of the Adriatic Sea in the Mediterranean Sea extending in the south to the mouth of the Vjosë. The first account of the Illyrian groups comes from Periplus of the Euxine Sea, an ancient Greek text written in the middle of the 4th century BC. The west was inhabited by the Thracian tribe of the Bryges while the south was inhabited by the Ancient Greek-speaking tribe of the Chaonians, whose capital was at Phoenice. Other colonies such as Apollonia, Epidamnos, and Amantia, were established by Ancient Greek city-states on the coast by the 7th century BC.
The Illyrian Ardiaei tribe, centred in Montenegro, ruled over most of the territory of Albania. Their Ardiaean Kingdom reached its greatest extent under King Agron, the son of Pleuratus II. Agron extended his rule over other neighbouring tribes as well. Following Agron's death in 230 BC, his wife, Teuta, inherited the Ardiaean kingdom. Teuta's forces extended their operations further southwards to the Ionian Sea. In 229 BC, Rome declared war on the kingdom for extensively plundering Roman ships. The war ended in Illyrian defeat in 227 BC. Teuta was eventually succeeded by Gentius in 181 BC. Gentius clashed with the Romans in 168 BC, initiating the Third Illyrian War. The conflict resulted in Roman conquest of the region by 167 BC. The Romans split the region into three administrative divisions.


The term Albania is the medieval Latin name of the country. It may be derived from the Illyrian tribe of Albani (Albanian: Albanët) recorded by Ptolemy, the geographer and astronomer from Alexandria, who drafted a map in 150 AD which shows the city of Albanopolis located northeast of Durrës. The term may have a continuation in the name of a medieval settlement called Albanon or Arbanon, although it is not certain that this was the same place. In his history written in the 10th century, the Byzantine historian Michael Attaliates was the first to refer to Albanoi as having taken part in a revolt against Constantinople in 1043 and to the Arbanitai as subjects of the Duke of Dyrrachium. During the Middle Ages, the Albanians called their country Arbëri or Arbëni and referred to themselves as Arbëreshë or Arbëneshë.
Nowadays, Albanians call their country Shqipëri or Shqipëria. The words Shqipëri and Shqiptar are attested from 14th century onwards, but it was only at the end of 17th and beginning of the early 18th centuries that the placename Shqipëria and the ethnic demonym Shqiptarë gradually replaced Arbëria and Arbëreshë amongst Albanian speakers. The two terms are popularly interpreted as "Land of the Eagles" and "Children of the Eagles".


The chief environmental factors affecting the distribution of living organisms are temperature, humidity, climate, soil and light intensity, and the presence or absence of all the requirements that the organism needs to sustain it. Generally speaking, animal communities are reliant on specific types of plant communities.
Some plants and animals have habitat requirements which are met in a wide range of locations. The small white butterfly Pieris rapae for example is found on all the continents of the world apart from Antarctica. Its larvae feed on a wide range of Brassicas and various other plant species, and it thrives in any open location with diverse plant associations. The large blue butterfly Phengaris arion is much more specific in its requirements; it is found only in chalk grassland areas, its larvae feed on Thymus species and because of complex lifecycle requirements it inhabits only areas in which Myrmica ants live.
Disturbance is important in the creation of biodiverse habitat types. In the absence of disturbance, a climax vegetation cover develops that prevents the establishment of other species. Wildflower meadows are sometimes created by conservationists but most of the flowering plants used are either annuals or biennials and disappear after a few years in the absence of patches of bare ground on which their seedlings can grow. Lightning strikes and toppled trees in tropical forests allow species richness to be maintained as pioneering species move in to fill the gaps created. Similarly coastal habitat types can become dominated by kelp until the seabed is disturbed by a storm and the algae swept away, or shifting sediment exposes new areas for colonisation. Another cause of disturbance is when an area may be overwhelmed by an invasive introduced species which is not kept under control by natural enemies in its new habitat.


Habitat fragmentation describes the emergence of discontinuities (fragmentation) in an organism's preferred environment (habitat), causing population fragmentation and  ecosystem decay. Causes of habitat fragmentation include geological processes that slowly alter the layout of the physical environment (suspected of being one of the major causes of speciation), and human activity such as land conversion, which can alter the environment much faster and causes the extinction of many species. More specifically, habitat fragmentation is a process by which large and contiguous habitats get divided into smaller, isolated patches of habitats.


Whether from natural processes or the activities of man, landscapes and their associated habitat types change over time. There are the slow geomorphological changes associated with the geologic processes that cause tectonic uplift and subsidence, and the more rapid changes associated with earthquakes, landslides, storms, flooding, wildfires, coastal erosion, deforestation and changes in land use. Then there are the changes in habitat types brought on by alterations in farming practices, tourism, pollution, fragmentation and climate change.
Loss of habitat is the single greatest threat to any species. If an island on which an endemic organism lives becomes uninhabitable for some reason, the species will become extinct. Any type of habitat surrounded by a different habitat is in a similar situation to an island. If a forest is divided into parts by logging, with strips of cleared land separating woodland blocks, and the distances between the remaining fragments exceeds the distance an individual animal is able to travel, that species becomes especially vulnerable. Small populations generally lack genetic diversity and may be threatened by increased predation, increased competition, disease and unexpected catastrophe. At the edge of each forest fragment, increased light encourages secondary growth of fast-growing species and old growth trees are more vulnerable to logging as access is improved. The birds that nest in their crevices, the epiphytes that hang from their branches and the invertebrates in the leaf litter are all adversely affected and biodiversity is reduced. Habitat fragmentation can be ameliorated to some extent by the provision of wildlife corridors connecting the fragments. These can be a river, ditch, strip of trees, hedgerow or even an underpass to a highway. Without the corridors, seeds cannot disperse and animals, especially small ones, cannot travel through the hostile territory, putting populations at greater risk of local extinction.
Habitat disturbance can have long-lasting effects on the environment. Bromus tectorum is a vigorous grass from Europe which has been introduced to the United States where it has become invasive. It is highly adapted to fire, producing large amounts of flammable detritus and increasing the frequency and intensity of wildfires. In areas where it has become established, it has altered the local fire regimen to such an extant that native plants cannot survive the frequent fires, allowing it to become even more dominant. A marine example is when sea urchin populations "explode" in coastal waters and destroy all the macroalgae present. What was previously a kelp forest becomes an urchin barren that may last for years and this can have a profound effect on the food chain. Removal of the sea urchins, by disease for example, can result in the seaweed returning, with an over-abundance of fast-growing kelp.


In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical manifestation of its ecological niche. Thus "habitat" is a species-specific term, fundamentally different from concepts such as environment or vegetation assemblages, for which the term "habitat-type" is more appropriate.
The physical factors may include (for example): soil, moisture, range of temperature, and light intensity. Biotic factors will include the availability of food and the presence or absence of predators. Every species has particular habitat requirements, with habitat generalist species able to thrive in a wide array of environmental conditions while habitat specialist species requiring a very limited set of factors to survive. The habitat of a species is not necessarily found in a geographical area, it can be the interior of a stem, a rotten log, a rock or a clump of moss; a parasitic organism has as its habitat the body of its host, part of the host's body (such as the digestive tract), or a single cell within the host's body.
Habitat types are environmental categorizations of different environments based on the characteristics of a given geographical area, particularly vegetation and climate. Thus habitat types do not refer to a single species but to multiple species living in the same area. For example, terrestrial habitat types include forest, steppe, grassland, semi-arid or desert. Fresh-water habitat types include marshes, streams, rivers, lakes, and ponds; marine habitat types include salt marshes, the coast, the intertidal zone, estuaries, reefs, bays, the open sea, the sea bed, deep water and submarine vents. Habitat types may change over time. Causes of change may include a violent event (such as the eruption of a volcano, an earthquake, a tsunami, a wildfire or a change in oceanic currents); or change may occur more gradually over millennia with alterations in the climate, as ice sheets and glaciers advance and retreat, and as different weather patterns bring changes of precipitation and solar radiation. Other changes come as a direct result of human activities, such as deforestation, the plowing of ancient grasslands, the diversion and damming of rivers, the draining of marshland and the dredging of the seabed. The introduction of alien species can have a devastating effect on native wildlife – through increased predation, through competition for resources or through the introduction of pests and diseases to which the indigenous species have no immunity.


Terrestrial habitat types include forests, grasslands, wetlands and deserts. Within these broad biomes are more specific habitat types with varying climate types, temperature regimes, soils, altitudes and vegetation. Many of these habitat types grade into each other and each one has its own typical communities of plants and animals. A habitat-type may suit a particular species well, but its presence or absence at any particular location depends to some extent on chance, on its dispersal abilities and its efficiency as a colonizer.
Freshwater habitat types include rivers, streams, lakes, ponds, marshes and bogs. Although some organisms are found across most of these habitat types, the majority have more specific requirements. The water velocity, its temperature and oxygen saturation are important factors, but in river systems, there are fast and slow sections, pools, bayous and backwaters which provide a range of habitat types. Similarly, aquatic plants can be floating, semi-submerged, submerged or grow in permanently or temporarily saturated soils besides bodies of water. Marginal plants provide important habitat for both invertebrates and vertebrates, and submerged plants provide oxygenation of the water, absorb nutrients and play a part in the reduction of pollution.
Marine habitats include brackish water, estuaries, bays, the open sea, the intertidal zone, the sea bed, reefs and deep / shallow water zones. Further variations include rock pools, sand banks, mudflats, brackish lagoons, sandy and pebbly beaches, and seagrass beds, all supporting their own flora and fauna. The benthic zone or seabed provides a home for both static organisms, anchored to the substrate, and for a large range of organisms crawling on or burrowing into the surface. Some creatures float among the waves on the surface of the water, or raft on floating debris, others swim at a range of depths, including organisms in the demersal zone close to the seabed, and myriads of organisms drift with the currents and form the plankton.
A desert is not the kind of habitat that favours the presence of amphibians, with their requirement for water to keep their skins moist and for the development of their young. Nevertheless, some frogs live in deserts, creating moist habitat types underground and hibernating while conditions are adverse. Couch's spadefoot toad (Scaphiopus couchii) emerges from its burrow when a downpour occurs and lays its eggs in the transient pools that form; the tadpoles develop with great rapidity, sometimes in as little as nine days, undergo metamorphosis, and feed voraciously before digging a burrow of their own.
Other organisms cope with the drying up of their aqueous habitat in other ways. Vernal pools are ephemeral ponds that form in the rainy season and dry up afterwards. They have their specially-adapted characteristic flora, mainly consisting of annuals, the seeds of which survive the drought, but also some uniquely adapted perennials. Animals adapted to these extreme habitat types also exist; fairy shrimps can lay "winter eggs" which are resistant to desiccation, sometimes being blown about with the dust, ending up in new depressions in the ground. These can survive in a dormant state for as long as fifteen years. Some killifish behave in a similar way; their eggs hatch and the juvenile fish grow with great rapidity when the conditions are right, but the whole population of fish may end up as eggs in diapause in the dried up mud that was once a pond.
Many animals and plants have taken up residence in urban environments. They tend to be adaptable generalists and use the town's features to make their homes. Rats and mice have followed man around the globe, pigeons, peregrines, sparrows, swallows and house martins use the buildings for nesting, bats use roof space for roosting, foxes visit the garbage bins and squirrels, coyotes, raccoons and skunks roam the streets. About 2,000 coyotes are thought to live in and around Chicago. A survey of dwelling houses in northern European cities in the twentieth century found about 175 species of invertebrate inside them, including 53 species of beetle, 21 flies, 13 butterflies and moths, 13 mites, 9 lice, 7 bees, 5 wasps, 5 cockroaches, 5 spiders, 4 ants and a number of other groups. In warmer climates, termites are serious pests in the urban habitat; 183 species are known to affect buildings and 83 species cause serious structural damage.


The development of maritime trade has rapidly affected the way marine organisms are transported within the ocean. Two ways marine organisms are transported to new environments are hull fouling and ballast water transport. In fact, Molnar et al. 2008 documented the pathways of hundreds of marine invasive species and found that shipping was the dominant mechanism for the transfer of invasive species.
Many marine organisms have the capacity to attach themselves to vessel hulls. Therefore, these organisms are easily transported from one body of water to another and are a significant risk factor for a biological invasion event. Controlling for vessel hull fouling is voluntary and there are no regulations currently in place to manage hull fouling. However, the governments of California and New Zealand have announced more stringent control for vessel hull fouling within their respective jurisdictions.
The other main vector for the transport of non-native aquatic species is ballast water. Ballast water taken up at sea and released in port by transoceanic vessels is the largest vector for non-native aquatic species invasions. In fact, it is estimated that 10,000 different species, many of which are non-indigenous, are transported via ballast water each day. Many of these species are considered harmful and can negatively affect their new environment. For example, freshwater zebra mussels, native to the Black, Caspian and Azov seas, most likely reached the Great Lakes via ballast water from a transoceanic vessel. Zebra mussels outcompete other native organisms for oxygen and food, such as algae. Although the zebra mussel invasion was first noted in 1988, and a mitigation plan was successfully implemented shortly thereafter, the plan had a serious flaw or loophole, whereby ships loaded with cargo when they reached the Seaway were not tested because their ballast water tanks were empty. However, even in an empty ballast tank, there remains a puddle of water filled with organisms that could be released at the next port (when the tank is filled with water after unloading the cargo, the ship takes on ballast water which mixes with the puddles and then everything including the living organisms in the puddles is discharged at the next port). Current regulations for the Great Lakes rely on ‘salinity shock’ to kill freshwater organisms left in ballast tanks.
Even though ballast water regulations are in place to protect against potentially invasive species, there exists a loophole for organisms in the 10–50 micron size class. For organisms between 10 and 50 microns, such as certain types of phytoplankton, current regulations allow less than 10 cells per milliliter be present in discharge from treatment systems. The discharge gets released when a ship takes on cargo at a port so the discharged water is not necessarily the same as the receiving body of water. Since many species of phytoplankton are less than 10 microns in size and reproduce asexually, only one cell released into the environment could exponentially grow into many thousands of cells over a short amount of time. This loophole could have detrimental effects to the environment. For example, some species in the genus Pseudo-nitzschia are smaller than 10 microns in width and contain domoic acid, a neurotoxin. If toxic Pseudo-nitzschia spp. are alive in ballast discharge and get released into their “new environment” they could cause domoic acid poisoning in shellfish, marine mammals and birds. Fortunately, human deaths related to domoic acid poisoning have been prevented because of stringent monitoring programs that arose after a domoic acid outbreak in Canada in 1987. Ballast water regulations need to be more rigorous to prevent future ramifications associated with the potential release of toxic and invasive phytoplankton.
Another important factor to consider about marine invasive species is the role of environmental changes associated with climate change, such as an increase in ocean temperature. There have been multiple studies suggesting an increase in ocean temperature will cause range shifts in organisms, which could have detrimental effects on the environment as new species interactions emerge. For example, Hua and Hwang proposed that organisms in a ballast tank of a ship traveling from the temperate zone through tropical waters can experience temperature fluctuations as much as 20 °C. To further examine the effects of temperature on organisms transported on hulls or in ballast water, Lenz et al. (2018) carried out study where they conducted a double heat stress experiment. Their results suggest that heat challenges organisms face during transport may enhance the stress tolerance of species in their non-native range by selecting for genetically adapted genotypes that will survive a second applied heat stress, such as increased ocean temperature in the founder population. Due to the complexity of climate-change-induced variations, it is difficult to predict the nature of temperature-based success of non-native species in-situ. Since some studies have suggested increased temperature tolerance of “hijackers” on ships’ hulls or in ballast water, it is necessary to develop more comprehensive fouling and ballast water management plans in an effort to prevent against future possible invasions as environmental conditions continue to change around the world.


Non-native species have many vectors, including biogenic vectors, but most invasions are associated with human activity. Natural range extensions are common in many species, but the rate and magnitude of human-mediated extensions in these species tend to be much larger than natural extensions, and humans typically carry specimens greater distances than natural forces.
An early human vector occurred when prehistoric humans introduced the Pacific rat (Rattus exulans) to Polynesia.
Vectors include plants or seeds imported for horticulture. The pet trade moves animals across borders, where they can escape and become invasive. Organisms stow away on transport vehicles. Among professionals in invasion biology, the overwhelming consensus is that incidental human assisted transfer is the main cause of introductions – other than for polar regions. Diseases may also be vectored by invasive insects such as the Asian citrus psyllid and the bacterial disease citrus greening.
The arrival of invasive propagules to a new site is a function of the site's invasibility.
Species have also been introduced intentionally. For example, to feel more "at home," American colonists formed "Acclimation Societies" that repeatedly imported birds that were native to Europe to North America and other distant lands. In 2008, U.S. postal workers in Pennsylvania noticed noises coming from inside a box from Taiwan; the box contained more than two dozen live beetles. Agricultural Research Service entomologists identified them as the rhinoceros beetle, Hercules beetle, and king stag beetle. Because these species were not native to the U.S., they could have threatened native ecosystems. To prevent exotic species from becoming a problem in the U.S., special handling and permits are required when living materials are shipped from foreign countries. USDA programs such as Smuggling Interdiction and Trade Compliance (SITC) attempt to prevent exotic species outbreaks in America. The intentional spread of domesticated plants to other favourable environments has been described as biological globalization.
Many invasive species, once they are dominant in the area, are essential to the ecosystem of that area. If they are removed from the location it could be harmful to that area.
Economics plays a major role in exotic species introduction. High demand for the valuable Chinese mitten crab is one explanation for the possible intentional release of the species in foreign waters.


Invasive species often exploit disturbances to an ecosystem (wildfires, roads, foot trails) to colonize an area. Large wildfires can sterilize soils, while adding a variety of nutrients. In the resulting free-for-all, formerly entrenched species lose their advantage, leaving more room for invasives. In such circumstances, plants that can regenerate from their roots have an advantage. Non-natives with this ability can benefit from a low intensity fire burn that removes surface vegetation, leaving natives that rely on seeds for propagation to find their niches occupied when their seeds finally sprout.
Wildfires often occur in remote areas, needing fire suppression crews to travel through pristine forest to reach the site. The crews can bring invasive seeds with them. If any of these stowaway seeds become established, a thriving colony of invasives can erupt in as few as six weeks, after which controlling the outbreak can need years of continued attention to prevent further spread. Also, disturbing the soil surface, such as cutting firebreaks, destroys native cover, exposes soil, and can accelerate invasions. In suburban and wildland-urban interface areas, the vegetation clearance and brush removal ordinances of municipalities for defensible space can result in excessive removal of native shrubs and perennials that exposes the soil to more light and less competition for invasive plant species.
Fire suppression vehicles are often major culprits in such outbreaks, as the vehicles are often driven on back roads overgrown with invasive plant species. The undercarriage of the vehicle becomes a prime vessel of transport. In response, on large fires, washing stations "decontaminate" vehicles before engaging in suppression activities. Large wildfires attract firefighters from remote places, further increasing the potential for seed transport.


An Asturian Visigothic noble named Pelagius of Asturias in 718 was elected leader by many of the ousted Visigoth nobles. Pelagius called for the remnant of the Christian Visigothic armies to rebel against the Moors and regroup in the unconquered northern Asturian highlands, better known today as the Cantabrian Mountains, in what is today the small mountain region in north-western Spain, adjacent to the Bay of Biscay.
Pelagius' plan was to use the Cantabrian mountains as a place of refuge and protection from the invading Moors. He then aimed to regroup the Iberian Peninsula's Christian armies and use the Cantabrian mountains as a springboard from which to regain their lands. In the process, after defeating the Moors in the Battle of Covadonga in 722, Pelagius was proclaimed king, thus founding the Christian Kingdom of Asturias and starting the war of Christian reconquest known in Portuguese as the Reconquista Cristã.
At the end of the 9th century, the region of Portugal, between the rivers Minho and Douro, was reconquered from the Moors by the nobleman and knight Vímara Peres on the orders of King Alfonso III of Asturias. Finding that the region had previously had two major cities – Portus Cale in the coast and Braga in the interior, with many towns that were now deserted – he decided to repopulate and rebuild them with Portuguese and Galician refugees and other Christians.
Apart from the Arabs from the South, the coastal regions in the North were also attacked by Norman and Viking raiders mainly from 844. The last great invasion, through the Minho (river), ended with the defeat of Olaf II Haraldsson in 1014 against the Galician nobility who also stopped further advances into the County of Portugal.
Count Vímara Peres organized the region he had reconquered, and elevated it to the status of County, naming it the County of Portugal after the region's major port city – Portus Cale or modern Porto. One of the first cities Vimara Peres founded at this time is Vimaranes, known today as Guimarães – the "birthplace of the Portuguese nation" or the "cradle city" (Cidade Berço in Portuguese).
After annexing the County of Portugal into one of the several counties that made up the Kingdom of Asturias, King Alfonso III of Asturias knighted Vímara Peres, in 868, as the First Count of Portus Cale (Portugal). The region became known as Portucale, Portugale, and simultaneously Portugália – the County of Portugal.
Later the Kingdom of Asturias was divided into a number of Christian Kingdoms in Northern Iberia due to dynastic divisions of inheritance among the king's offspring. With the forced abdication of Alfonso III "the Great" of Asturias by his sons in 910, the Kingdom of Asturias split into three separate kingdoms. The three kingdoms were eventually reunited in 924 under the crown of León.
In 1093, Alfonso VI of León bestowed the county to Henry of Burgundy and married him to his illegitimate daughter, Teresa of León, for his role in reconquering the land from Moors. Henry based his newly formed county in Bracara Augusta (modern Braga), capital city of the ancient Roman province, and also previous capital of several kingdoms over the first millennia.


In the early 5th century, Germanic tribes, namely the Suebi and the Vandals (Silingi and Hasdingi) together with their allies, the Sarmatians and Alans invaded the Iberian Peninsula where they would form their kingdom. The Kingdom of the Suebi was the Germanic post-Roman kingdom, established in the former Roman provinces of Gallaecia-Lusitania. 5th-century vestiges of Alan settlements were found in Alenquer (from old Germanic Alan kerk, temple of the Alans), Coimbra and Lisbon.
About 410 and during the 6th century it became a formally declared Kingdom of the Suebi, where king Hermeric made a peace treaty with the Gallaecians before passing his domains to Rechila, his son. In 448 Rechila died, leaving the state in expansion to Rechiar.
After the defeat against the Visigoths, the Suebian kingdom was divided, with Frantan and Aguiulfo ruling simultaneously. Both reigned from 456 to 457, the year in which Maldras (457–459) reunified the kingdom. He was assassinated after a failed Roman-Visigothic conspiracy.
Although the conspiracy did not achieve its true purposes, the Suebian Kingdom was again divided between two kings:
Frumar (Frumario 459–463) and Remismund (Remismundo, son of Maldras) (459–469) who would re-reunify his father's kingdom in 463. He would be forced to adopt Arianism in 465 due to the Visigoth influence.
By the year 500, the Visigothic Kingdom had been installed in Iberia, it was based in Toledo and advancing westwards. They became a threat to the Suebian rule.
After the death of Remismund in 469 a dark period set in, where virtually all written texts and accounts disappear. This period lasted until 550. The only thing known about this period is that Theodemund (Teodemundo) most probably ruled the Suebians.
The dark period ended with the reign of Karriarico (550–559) who reinstalled Catholic Christianity in 550. He was succeeded by Theodemar (559–570) during whose reign the 1st Council of Braga (561) was held.
The councils represented an advance in the organization of the territory (paroeciam suevorum (Suebian parish) and the Christianization of the pagan population (De correctione rusticorum) under the auspices of Saint Martin of Braga (São Martinho de Braga).
After the death of Teodomiro, Miro (570–583) was his successor. During his reign, the 2nd Council of Braga (572) was held.
The Visigothic civil war began in 577. Miro intervened. Later in 583 he also organized an unsuccessful expedition to reconquer Seville. During the return from this failed operation Miro died.
In the Suebian Kingdom many internal struggles continued to take place. Eborico (Eurico, 583–584) was dethroned by Andeca (Audeca 584–585), who failed to prevent the Visigothic invasion led by Leovigildo. The Visigothic invasion, completed in 585, turned the once rich and fertile kingdom of the Suebi into the sixth province of the Gothic kingdom.Leovigild was crowned King of Gallaecia, Hispania and Gallia Narbonensis.
For the next 300 years and by the year 700, the entire Iberian Peninsula was ruled by the Visigoths. Under the Visigoths, Gallaecia was a well-defined space governed by a doge of its own. Doges at this time were related to the monarchy and acted as princes in all matters. Both 'governors' Wamba and Wittiza (Vitiza) acted as doge (they would later become kings in Toledo). These two became known as the 'vitizians', who headquartered in the northwest and called on the Arab invaders from the South to be their allies in the struggle for power in 711. King Roderic (Rodrigo) was killed while opposing this invasion, thus becoming the last Visigothic king of Iberia. From the various Germanic groups who settled in western Iberia, the Suebi left the strongest lasting cultural legacy in what is today Portugal, Galicia and western fringes of Asturias.
According to Dan Stanislawski, the Portuguese way of living in regions North of the Tagus is mostly inherited from the Suebi, in which small farms prevail, distinct from the large properties of Southern Portugal.
Bracara Augusta, the modern city of Braga and former capital of Gallaecia, became the capital of the Suebi. Apart from cultural and some linguistic traces, the Suebians left the highest Germanic genetic contribution of the Iberian Peninsula in Portugal and Galicia. Orosius, at that time resident in Hispania, shows a rather pacific initial settlement, the newcomers working their lands or serving as bodyguards of the locals.
Another Germanic group that accompanied the Suebi and settled in Gallaecia were the Buri. They settled in the region between the rivers Cávado and Homem, in the area known as Terras de Bouro (Lands of the Buri).


Romans first invaded the Iberian Peninsula in 219 BC. The Carthaginians, Rome's adversary in the Punic Wars, were expelled from their coastal colonies. During the last days of Julius Caesar, almost the entire peninsula was annexed to the Roman Republic.
The Roman conquest of what is now part of Portugal took almost two hundred years and took many lives of young soldiers and
the lives of those who were sentenced to a certain death in the slave mines when not sold as slaves to other parts of the empire. It suffered a severe setback in 155 BC, when a rebellion began in the north. The Lusitanians and other native tribes, under the leadership of Viriathus, wrested control of all of western Iberia.
Rome sent numerous legions and its best generals to Lusitania to quell the rebellion, but to no avail – the Lusitanians kept conquering territory. The Roman leaders decided to change their strategy. They bribed Viriathus's allies to kill him. In 139 BC, Viriathus was assassinated and Tautalus became leader of the Lusitanians.
Rome installed a colonial regime. The complete Romanization of Lusitania only took place in the Visigothic era.
In 27 BC, Lusitania gained the status of Roman province. Later, a northern province of Lusitania was formed, known as Gallaecia, with capital in Bracara Augusta, today's Braga.
There are still many ruins of castros (hill forts) throughout modern Portugal and remains of the Castro culture.
Some urban remains are quite large, like Conímbriga and Mirobriga. The former, beyond being one of the largest Roman settlements in Portugal, is also classified as a National Monument. Conímbriga lies 16 kilometres (10 miles) from Coimbra, which in turn was the ancient Aeminium. The site also has a museum that displays objects found by archaeologists during their excavations.
Several works of engineering, such as baths, temples, bridges, roads, circuses, theatres and laymen's homes are preserved throughout the country. Coins, some coined in Lusitanian land, as well as numerous pieces of ceramics, were also found. Contemporary historians include Paulus Orosius (c. 375–418) and Hydatius (c. 400–469), bishop of Aquae Flaviae, who reported on the final years of the Roman rule and arrival of the Germanic tribes.


On 24 June 1128, the Battle of São Mamede occurred near Guimarães. Afonso Henriques, Count of Portugal, defeated his mother Countess Teresa and her lover Fernão Peres de Trava, thereby establishing himself as sole leader. Afonso then turned his arms against the Moors in the south.
Afonso's campaigns were successful and, on 25 July 1139, he obtained an overwhelming victory in the Battle of Ourique, and straight after was unanimously proclaimed King of Portugal by his soldiers. This is traditionally taken as the occasion when the County of Portugal, as a fief of the Kingdom of León, was transformed into the independent Kingdom of Portugal.
Afonso then established the first of the Portuguese Cortes at Lamego, where he was crowned by the Archbishop of Braga, though the validity of the Cortes of Lamego has been disputed and called a myth created during the Portuguese Restoration War. Afonso was recognized in 1143 by King Alfonso VII of León, and in 1179 by Pope Alexander III.
During the Reconquista period, Christians reconquered the Iberian Peninsula from Moorish domination. Afonso Henriques and his successors, aided by military monastic orders, pushed southward to drive out the Moors. At this time, Portugal covered about half of its present area. In 1249, the Reconquista ended with the capture of the Algarve and complete expulsion of the last Moorish settlements on the southern coast, giving Portugal its present-day borders, with minor exceptions.
In one of these situations of conflict with the kingdom of Castile, Dinis I of Portugal signed with the king Fernando IV of Castile (who was represented, when a minor, by his mother the queen Maria de Molina) the Treaty of Alcañices (1297), which stipulated that Portugal abolished agreed treaties against the kingdom of Castile for supporting the infant Juan de Castilla. This treaty established among other things the border demarcation between the kingdom of Portugal and the kingdom of Leon, where the disputed town of Olivenza was included.
The reigns of Dinis I (Denis I), Afonso IV (Alphons IV), and Pedro I (Peter I) for the most part saw peace with the Christian kingdoms of Iberia.
In 1348 and 1349 Portugal, like the rest of Europe, was devastated by the Black Death. In 1373, Portugal made an alliance with England, which is one of the oldest standing alliances in the world. Over time, this went far beyond geo-political and military cooperation (protecting both nations' interests in Africa, the Americas and Asia against French, Spanish and Dutch rivals) and maintained strong trade and cultural ties between the two old European allies. In the Oporto region, in particular, there is visible English influence to this day.


The early history of Portugal is shared with the rest of the Iberian Peninsula located in south-western Europe. The name of Portugal derives from the joined Romano-Celtic name Portus Cale. The region was settled by Pre-Celts and Celts, giving origin to peoples like the Gallaeci, Lusitanians, Celtici and Cynetes (also known as Conii), visited by Phoenicians-Carthaginians and Ancient Greeks, was incorporated in the Roman Republic dominions as Lusitania and part of Gallaecia, after 45 BC until 298 AD.
The region of present-day Portugal was inhabited by Neanderthals and then by Homo sapiens, who roamed the border-less region of the northern Iberian peninsula. These were subsistence societies and although they did not establish prosperous settlements, they did form organized societies. Neolithic Portugal experimented with domestication of herding animals, the raising of some cereal crops and fluvial or marine fishing.
It is believed by some scholars that early in the first millennium BC, several waves of Celts invaded Portugal from Central Europe and inter-married with the local populations, forming different tribes. Another theory suggests that Celts inhabited western Iberia / Portugal well before any large Celtic migrations from Central Europe. In addition, a number of linguists expert in ancient Celtic have presented compelling evidence that the Tartessian language, once spoken in parts of SW Spain and SW Portugal, is at least proto-Celtic in structure.
Modern archaeology and research shows a Portuguese root to the Celts in Portugal and elsewhere. During that period and until the Roman invasions, the Castro culture (a variation of the Urnfield culture also known as Urnenfelderkultur) was prolific in Portugal and modern Galicia.
This culture, together with the surviving elements of the Atlantic megalithic culture and the contributions that come from the more Western Mediterranean cultures, ended up in what has been called the Cultura Castreja or Castro Culture. This designation refers to the characteristic Celtic populations called 'dùn', 'dùin' or 'don' in Gaelic and that the Romans called castrae in their chronicles.
Based on the Roman chronicles about the Callaeci peoples, along with the Lebor Gabála Érenn narrations and the interpretation of the abundant archaeological remains throughout the northern half of Portugal and Galicia, it is possible to infer that there was a matriarchal society, with a military and religious aristocracy probably of the feudal type. The figures of maximum authority were the chieftain (chefe tribal), of military type and with authority in his Castro or clan, and the druid, mainly referring to medical and religious functions that could be common to several castros. The Celtic cosmogony remained homogeneous due to the ability of the druids to meet in councils with the druids of other areas, which ensured the transmission of knowledge and the most significant events.
The first documentary references to Castro society are provided by chroniclers of Roman military campaigns such as Strabo, Herodotus and Pliny the Elder among others, about the social organization, and describing the inhabitants of these territories, the Gallaeci of Northern Portugal as:
"A group of barbarians who spend the day fighting and the night eating, drinking and dancing under the moon."
There were other similar tribes, and chief among them were the Lusitanians; the core area of these people lay in inland central Portugal, while numerous other related tribes existed such as the Celtici of Alentejo, and the Cynetes or Conii of the Algarve. Among the tribes or sub-divisions were the Bracari, Coelerni, Equaesi, Grovii, Interamici, Leuni, Luanqui, Limici, Narbasi, Nemetati, Paesuri, Quaquerni, Seurbi, Tamagani, Tapoli, Turduli, Turduli Veteres, Turdulorum Oppida, Turodi, and Zoelae. A few small, semi-permanent, commercial coastal settlements (such as Tavira) were also founded in the Algarve region by Phoenicians–Carthaginians.


Today's continental Portugal, along with most of modern Spain, was part of al-Andalus between 726 and 1249, following the Umayyad Caliphate conquest of the Iberian Peninsula. This rule lasted from some decades in the North to five centuries in the South.
After defeating the Visigoths in only a few months, the Umayyad Caliphate started expanding rapidly in the peninsula. Beginning in 726, the land that is now Portugal became part of the vast Umayyad Caliphate's empire of Damascus, which stretched from the Indus river in the Indian sub-continent up to the South of France, until its collapse in 750. That year the west of the empire gained its independence under Abd-ar-Rahman I with the establishment of the Emirate of Córdoba. After almost two centuries, the Emirate became the Caliphate of Córdoba in 929, until its dissolution a century later in 1031 into no less than 23 small kingdoms, called Taifa kingdoms.
The governors of the taifas each proclaimed themselves Emir of their provinces and established diplomatic relations with the Christian kingdoms of the north. Most of present-day Portugal fell into the hands of the Taifa of Badajoz of the Aftasid Dynasty, and after a short spell of an ephemeral Taifa of Lisbon in 1022, fell under the dominion of the Taifa of Seville of the Abbadids poets. The Taifa period ended with the conquest of the Almoravids who came from Morocco in 1086 winning a decisive victory at the Battle of Sagrajas, followed a century later in 1147, after the second period of Taifa, by the Almohads, also from Marrakesh.
Al-Andaluz was divided into different districts called Kura. Gharb Al-Andalus at its largest was constituted of ten kuras, each with a distinct capital and governor. The main cities of the period in Portugal were in the southern half of the country: Beja, Silves, Alcácer do Sal, Santarém and Lisbon.
The Muslim population of the region consisted mainly of native Iberian converts to Islam (the so-called Muwallad or Muladi) and Arabs. The Arabs were principally noblemen from Syria and Oman; and though few in numbers, they constituted the elite of the population. The Berbers were originally from the Rif and Atlas mountains region of North Africa and were nomads.


July 6 – Daniel Morgan, American pioneer, Congressman from Virginia, and general (b. 1736)
July 22 – Xavier Bichat, French anatomist and pathologist (b. 1771)
July 24 – Joseph Ducreux, French noble, portrait painter, pastelist, miniaturist, and engraver (b. 1735)
July 25 – Friedrich Karl Joseph von Erthal, Archbishop of Mainz (b. 1719)
August 10 – Franz Aepinus, German philosopher (b. 1724)
August 12 – Louis Lebègue Duportail French military leader in the Continental Army during the American Revolutionary War (b. 1743)
September 19 – Princess Luisa of Naples and Sicily (b. 1773)
September 26 – Jurij Vega, Slovenian mathematician, physicist, and soldier (b. 1754)
October 5 – Suzanne Bélair, Haitian national heroine (b. 1781)
October 8 – Emmanuele Vitale, Maltese military leader (b.1758)
October 31 – Sir William Parker, 1st Baronet, of Harburn, British admiral (b. 1743)
November 9 – Thomas Girtin, English artist (b. 1775)
November 15 – George Romney, English artist (b. 1734)
November 16 – André Michaux, French botanist (b. 1746)
December 5 – Lemuel Francis Abbott, English portrait painter (b. 1716)
December 31 – Francis Lewis, signer of the United States Declaration of Independence (b. 1713)


1802 (MDCCCII) was a common year starting on Friday of the Gregorian calendar and a common year starting on Wednesday of the Julian calendar, the 1802nd year of the Common Era (CE) and Anno Domini (AD) designations, the 802nd  year of the 2nd millennium, the 2nd  year of the 19th century, and the  3rd   year of the 1800s decade. As of the start of 1802, the Gregorian calendar was 12 days ahead of the Julian calendar, which remained in localized use until 1923.


July 5 – Parliamentary elections begin in the United Kingdom, with voting continuing until August 28; the Tories, led by Henry Addington, win control of the House of Commons.
July 19 – Éleuthère Irénée du Pont founds E. I. du Pont de Nemours and Company, the modern DuPont chemical company, as a gunpowder manufactory near Wilmington, Delaware.
July 22 – Gia Long captures Hanoi, completing his unification of Vietnam.
July 31 – William Wordsworth, leaving London for Dover and Calais with his sister Dorothy, witnesses the early morning scene which he captures in his sonnet "Composed upon Westminster Bridge".
August 2 – In a plebiscite, Napoleon Bonaparte is confirmed as the First Consul of France.
September 11 – The Italian region of Piedmont becomes a part of the French First Republic.


1806 (MDCCCVI) was a common year starting on Wednesday of the Gregorian calendar and a common year starting on Monday of the Julian calendar, the 1806th year of the Common Era (CE) and Anno Domini (AD) designations, the 806th  year of the 2nd millennium, the 6th  year of the 19th century, and the  7th   year of the 1800s decade. As of the start of 1806, the Gregorian calendar was 12 days ahead of the Julian calendar, which remained in localized use until 1923.


April 8 – Stéphanie de Beauharnais, adopted daughter of Napoleon Bonaparte, marries Prince Karl Ludwig Friedrich of Baden.
April 25 – Rana Bahadur Shah is killed by his step-brother Sher Bahadur Shah in the late night meeting which triggers two weeks' long massacre in Bhandarkhal garden.
May 30 – Future President of the United States Andrew Jackson fights his second duel, killing a man who had accused Jackson's wife of bigamy.
June 5 – Louis Bonaparte is appointed as King of Holland by his brother, Emperor Napoleon, replacing the Batavian Republic.


January 1
The French Republican Calendar is abolished.
The Kingdom of Bavaria is established by Napoleon.
January 5 – The body of Horatio Nelson, 1st Viscount Nelson, lies in state in the Painted Hall of Greenwich Hospital, London, prior to his funeral.
January 8 – Battle of Blaauwberg: British infantry force troops of the Batavian Republic in the Dutch Cape Colony to withdraw.
January 9
The Dutch commandant of Cape Town surrenders to British forces. On January 10, formal capitulation is signed under the Treaty Tree in  Papendorp (modern-day Woodstock).
Lord Nelson is given a state funeral and interment at St Paul's Cathedral in London, attended by the Prince of Wales.
January 18 – The Dutch Cape Colony capitulates to British forces, the origin of its status as a colony within the British Empire.
January 23
Grenville succeeds his cousin William Pitt the Younger as wartime Prime Minister of the United Kingdom upon Pitt's death this day amidst worsening health, caused by the stresses of the Napoleonic Wars.
Invasion of Naples: Ferdinand I of the Two Sicilies flees to the Kingdom of Sicily under British protection.
February 6 – Battle of San Domingo: The British Royal Navy gains a victory over the French off Santo Domingo.
February 15 – Invasion of Naples: Joseph Bonaparte enters Naples.
March 23 – Explorers Lewis and Clark and their Corps of Discovery, having reached the Pacific Ocean after traveling through the Louisiana Purchase, begin their journey home.
March 28 – Washington College (modern-day Washington & Jefferson College) is chartered by the Pennsylvania General Assembly.
March 29 – Construction is authorized of the National Road, the first United States federal highway.
March 30 – Kingdom of Naples (Napoleonic) proclaimed.


Species Plantarum was published on 1 May 1753 by Laurentius Salvius in Stockholm, in two volumes. A second edition was published in 1762–1763, and a third edition in 1764, although this "scarcely differed" from the second. Further editions were published after Linnaeus' death in 1778, under the direction of Karl Ludwig Willdenow, the director of the Berlin Botanical Garden; the fifth edition (1800) was published in four volumes.


Its full title is Species plantarum, exhibentes plantas rite cognitas ad genera relatas, cum differentiis specificis, nominibus trivialibus, synonymis selectis, locis natalibus, secundum systema sexuale digestas.
The book was actually published in two volumes, the first being on 24 May and the second on 16 August. However, for practical purposes, the dates of issue for volumes was arbitrarily set on 1 May, see Stearn, W.T. (1957), The preparation of the Species Plantarum and the introduction of binomial nomenclature, in: Species Plantarum, A Facsimile of the first edition, London, Ray Society: 72 and ICN (Melbourne Code) Art. 13.4 Note 1: "The two volumes of Linnaeus' Species plantarum, ed. 1 (1753), which appeared in May and August, 1753, respectively, are treated as having been published simultaneously on 1 May 1753."
Now considered a synonym of G. glabra.


Species Plantarum was the first botanical work to consistently apply the binomial nomenclature system of naming to any large group of organisms (Linnaeus' tenth edition of Systema Naturae would apply the same technique to animals for the first time in 1758). Prior to this work, a plant species would be known by a long polynomial, such as Plantago foliis ovato-lanceolatis pubescentibus, spica cylindrica, scapo tereti (meaning "plantain with pubescent ovate-lanceolate leaves, a cylindrical spike and a terete scape") or Nepeta floribus interrupte spicatis pedunculatis (meaning "Nepeta with flowers in a stalked, interrupted spike"). In Species Plantarum, these cumbersome names were replaced with two-part names, consisting of a single-word genus name, and a single-word specific epithet or "trivial name"; the two examples above became Plantago media and Nepeta cataria, respectively. The use of binomial names had originally been developed as a kind of shorthand in a student project about the plants eaten by cattle.
After the specific epithet, Linnaeus gave a short description of each species, and a synonymy. The descriptions were careful and terse, consisting of few words in small genera; in Glycyrrhiza, for instance, the three species (Glycyrrhiza echinata, Glycyrrhiza glabra and "Glycyrrhiza hirsuta", respectively) were described as "leguminibus echinatis", "leguminibus glabris" and "leguminibus hirsutis".
Because it is the first work in which binomial nomenclature was consistently applied, Species Plantarum was chosen as the "starting point" for the nomenclature of most plants (the nomenclature of some non-vascular plants and all fungi uses later starting points).


Species Plantarum contained descriptions of the thousands of plant species known to Linnaeus at the time. In the first edition, there were 5,940 names, from Acalypha australis to Zygophyllum spinosum. In his introduction, Linnaeus estimated that there were fewer than 10,000 plant species in existence; there are now thought to be around 400,000 species of flowering plants alone.
The species were arranged in around a thousand genera, which were grouped into 24 classes, according to Linnaeus' sexual system of classification. There are no descriptions of the genera in Species Plantarum; these are supplied in the companion volume Genera Plantarum ("the genera of plants"), the fifth edition of which was printed at a similar time to the first edition of Species Plantarum. Linnaeus' sexual system is now acknowledged to be an artificial system, rather than one which accurately reflects shared ancestry, but the system's simplicity made it easier for non-specialists to rapidly find the correct class, being based on simple counts of floral parts such as stigmas and stamens.


Species Plantarum (Latin for "The Species of Plants") is a book by Carl Linnaeus, originally published in 1753, which lists every species of plant known at the time, classified into genera. It is the first work to consistently apply binomial names and was the starting point for the naming of plants.


The zone of the sclerophyll vegetation lies in the border area between the subtropics and the temperate zone, approximately between the 30th and 40th degree of latitude (in the northern hemisphere also up to the 45th degree of latitude). Their presence is limited to the coastal western sides of the continents, but nonetheless can typical in any regions of a continent with scarce annual precipitation or frequent seasonal droughts and poor soils that are heavily leached.
The sclerophyll zone often merges into temperate deciduous forests towards the poles, on the coasts also into temperate rainforests and towards the equator in hot semi-deserts or deserts. The Mediterranean areas, which have a very high biodiversity, are under great pressure from the population. This is especially true for the Mediterranean region since ancient times. Through overexploitation (logging, grazing, agricultural use) and frequent fires caused by people, the original forest vegetation is converted. In extreme cases, the hard-leaf vegetation disappears completely and is replaced by open rock heaths.
Some sclerophyll areas are closer to the equator than the Mediterranean zone—for example, the interior of Madagascar, the dry half of New Caledonia, the lower edge areas of the Madrean pine-oak woodlands of the Mexican highlands between 800 and 1800/2000 m or around 2000 m high plateaus of the Asir Mountains on the western edge of the Arabian Peninsula.


Sclerophyll is a type of vegetation that is adapted to long periods of dryness and heat. The plants feature hard leaves, short internodes (the distance between leaves along the stem) and leaf orientation which is parallel or oblique to direct sunlight. The word comes from the Greek sklēros (hard) and phyllon (leaf). The term was coined by A.F.W. Schimper in 1898 (translated in 1903), originally as a synonym of xeromorph, but the two words were later differentiated.
Sclerophyllous plants occur in many parts of the world, but are most typical of areas with low rainfall or seasonal droughts, such as Australia, Africa, and western North and South America. They are prominent throughout Australia, parts of Argentina, the Cerrado biogeographic region of Bolivia, Paraguay and Brazil, and in the Mediterranean biomes that cover the Mediterranean Basin, California, Chile, and the Cape Province of South Africa.
In the Mediterranean basin, holm oak, cork oak and olives are typical hardwood trees. In addition, there are several species of pine under the trees in the vegetation zone. The shrub layer contains numerous herbs such as rosemary, thyme and lavender. In relation to the potential natural vegetation, around 2% of the earth's land surface is covered by sclerophyll woodlands, and a total of 10% of all plant species on earth live there.


The leaching of highly water soluble tannins from decaying vegetation and leaves along a stream may produce what is known as a blackwater river. Water flowing out of bogs has a characteristic brown color from dissolved peat tannins. The presence of tannins (or humic acid) in well water can make it smell bad or taste bitter, but this does not make it unsafe to drink.
Tannins leaching from an unprepared driftwood decoration in an aquarium can cause pH lowering and coloring of the water to a tea-like tinge. A way to avoid this is to boil the wood in water several times, discarding the water each time. Using peat as an aquarium substrate can have the same effect. Many hours of boiling the driftwood may need to be followed by many weeks or months of constant soaking and many water changes before the water will stay clear. Adding baking soda to the water to raise its pH level will accelerate the process of leaching, as the more alkaline solution can draw out tannic acid from the wood faster than the pH-neutral water.
Softwoods, while in general much lower in tannins than hardwoods, are usually not recommended for use in an aquarium so using a hardwood with a very light color, indicating a low tannin content, can be an easy way to avoid tannins. Tannic acid is brown in color, so in general white woods have a low tannin content. Woods with a lot of yellow, red, or brown coloration to them (like cedar, redwood, red oak, etc.) tend to contain a lot of tannin.


Tannins are distributed in species throughout the plant kingdom. They are commonly found in both gymnosperms and angiosperms. Mole studied the distribution of tannin in 180 families of dicotyledons and 44 families of monocotyledons (Cronquist). Most families of dicot contain tannin-free species (tested by their ability to precipitate proteins). The best known families of which all species tested contain tannin are: Aceraceae, Actinidiaceae, Anacardiaceae, Bixaceae, Burseraceae, Combretaceae, Dipterocarpaceae, Ericaceae, Grossulariaceae, Myricaceae for dicot and Najadaceae and Typhaceae in Monocot. To the family of the oak, Fagaceae, 73% of the species tested contain tannin. For those of acacias, Mimosaceae, only 39% of the species tested contain tannin, among Solanaceae rate drops to 6% and 4% for the Asteraceae. Some families like the Boraginaceae, Cucurbitaceae, Papaveraceae contain no tannin-rich species.
The most abundant polyphenols are the condensed tannins, found in virtually all families of plants, and comprising up to 50% of the dry weight of leaves.


1813 (MDCCCXIII) was a common year starting on Friday of the Gregorian calendar and a common year starting on Wednesday of the Julian calendar, the 1813th year of the Common Era (CE) and Anno Domini (AD) designations, the 813th  year of the 2nd millennium, the 13th  year of the 19th century, and the  4th   year of the 1810s decade. As of the start of 1813, the Gregorian calendar was 12 days ahead of the Julian calendar, which remained in localized use until 1923.


January 18–January 23 – War of 1812: The Battle of Frenchtown is fought in modern-day Monroe, Michigan between the United States and a British and Native American alliance.
January 24 – The Philharmonic Society (later the Royal Philharmonic Society) is founded in London.
January 28 – Jane Austen's Pride and Prejudice is published anonymously in London.
January 31 – The Assembly of the Year XIII is inaugurated in Buenos Aires.
February – War of 1812 in North America: General William Henry Harrison sends out an expedition to burn the British vessels at Fort Malden by going across Lake Erie via the Bass Islands in sleighs, but the ice is not hard enough, and the expedition returns.
February 3 – Argentine War of Independence: José de San Martín and his Regiment of Mounted Grenadiers gain a largely symbolic victory against a Spanish royalist army in the Battle of San Lorenzo.
February 7 – Napoleonic Wars: Action of 7 February 1813 – The French frigate Aréthuse (1812) and the British ship HMS Amelia (1796) engage in battle in the Îles de Los on the Guinea Coast; both ships retire unbeaten.
February 9 – Prussia abolishes the canton system.
February 11 – War of 1812: Construction begins on Fort Meigs in Ohio, under the command of General William Henry Harrison. Major Amos Stoddard assumes command of its artillery.
March 4
Napoleonic Wars: The French garrison evacuates Berlin, leaving Russian troops able to reach and take the city without a fight.
Cyril VI of Constantinople is elected Ecumenical Patriarch.
James Madison is sworn in for a second term as President of the United States.
March 17 – Napoleonic Wars: Prussia declares war on France, and introduces the Iron Cross military award (backdated to March 10).
March 28 – 1813–1814 Malta plague epidemic spreads from Egypt.
March 29 – Mexican War of Independence: Battle of Rosillo Creek – The Republican Army of the North defeats the Spanish Royalist Army in modern-day Bexar County, Texas.


January 1 –Gioacchino Navarro, Maltese priest and poet (b. 1748)
January 6 –Louis Baraguey d'Hilliers, French general (b. 1764)
January 15 –Anton Bernolák, Slovak linguist (b. 1762)
January 20 –Christoph Martin Wieland, German writer (b. 1733)
January 24 –George Clymer, American signer of the Declaration of Independence (b. 1739)
February 13 –Samuel Ashe, Governor of North Carolina (b. 1725)
February 26 –Robert Livingston, American signer of the Declaration of Independence (b. 1746)
March 23 – Princess Augusta of Great Britain, elder sibling of King George III (b. 1737)
April 3 – Friederike Brion, first great love of Johann Wolfgang Goethe (b. 1752)
April 10 –Joseph-Louis Lagrange, Italian mathematician (b. 1736)
April 19 – Benjamin Rush, Founding Father of the United States (b. 1746)
April 27 –Zebulon Pike, American general (b. 1779)
April 28 –Mikhail Kutuzov, Russian field marshal (b. 1745)
April 29 –John Andrews, American clergyman, Provost of the University of Pennsylvania, considered America's first scholar (b. 1746)
May 1 –Jean-Baptiste Bessières, French marshal (killed in action) (b. 1768)
May 21 – José Antonio Pareja, Spanish admiral (b. 1757)
May 23 –Géraud Duroc, French general (mortally wounded in action) (b. 1772)
June 6
Alexandre-Théodore Brongniart, French architect (b. 1739)
Antonio Cachia, Maltese architect, engineer and archaeologist (b. 1739)
June 17 –Charles Middleton, 1st Baron Barham, English sailor, politician (b. 1726)
July 6 – Granville Sharp, English abolitionist (b. 1735)
July 17 – Fredrica Löf, Swedish actress (b. 1760)
June 28 –Gerhard von Scharnhorst, Prussian general (b. 1755)


October 2 – The Philomathean Society of the University of Pennsylvania is founded (the oldest continuously existing literary society in the United States).
October 5 – War of 1812 –Battle of the Thames in Upper Canada: William Henry Harrison defeats the British, and native leader Tecumseh is killed in battle.
October 14 – After a ceremony in Caracas, Venezuela, the municipality gives Simón Bolívar the title of El Libertador.
October 16–19 – Napoleonic Wars – Battle of Leipzig: Napoleon is defeated by the forces of the Sixth Coalition. More than 600,000 troops are in the field, with well over 10% killed, wounded or missing. Many of the German states forming the Confederation of the Rhine defect from Napoleon to the Coalition, as a result of the battle.
October 24–November 5 – Persia and Russia sign the Treaty of Gulistan at the end of the Russo-Persian War, by which Persia loses modern-day Georgia, Dagestan and most of Azerbaijan to Russia.
October 26 – War of 1812 – Battle of the Chateauguay: Charles de Salaberry defeats an American invasion.
November 11 –War of 1812 – Battle of Crysler's Farm: An outnumbered British–Canadian force repels an American attack, forcing the Americans to give up their attempt to capture Montreal.
November 21 – An independent government is restored in the Netherlands.
December 8 – Ludwig van Beethoven's Symphony No. 7, together with his Wellington's Victory, are premiered in Vienna under the composer's baton, in a benefit concert for Austrian and Bavarian soldiers wounded at the Battle of Hanau.
December 18–19 – War of 1812: British soldiers and native allies invade the United States, and are successful in the Capture of Fort Niagara, and attack Lewiston, New York.
December 29 – War of 1812: British soldiers burn Buffalo, New York.


Rainforests are characterized by a closed and continuous tree canopy, moisture-dependent vegetation, the presence of epiphytes and lianas and the absence of wildfire. Rainforest can be classified as tropical rainforest or temperate rainforest, but other types have been described.
Estimates vary from 40% to 75% of all biotic species being indigenous to the rainforests. There may be many millions of species of plants, insects and microorganisms  still undiscovered in tropical rainforests. Tropical rainforests have been called the "jewels of the Earth" and the "world's largest pharmacy", because over one quarter of natural medicines have been discovered there.
Rainforests as well as endemic rainforest species are rapidly disappearing due to deforestation, the resulting habitat loss and pollution of the atmosphere.


The Scramble for Africa, also called the Partition of Africa, or the Conquest of Africa, was the invasion, annexation, division, and colonization of most of Africa by seven Western European powers during a short period known as New Imperialism (between 1881 and 1914). The 10 percent of Africa that was under formal European control in 1870 increased to almost 90 percent by 1914, with only Ethiopia (Abyssinia) and Liberia remaining independent, though Ethiopia would later be invaded and occupied by Italy from 1936 to 1941.

The Berlin Conference of 1884, which regulated European colonization and trade in Africa, is usually accepted as the beginning. In the last quarter of the 19th century, there were considerable political rivalries among the empires of the European continent, leading to the African continent being partitioned without wars between European nations. The later years of the 19th century saw a transition from "informal imperialism" — military influence and economic dominance — to direct rule.


The size of the Sahara has historically been extremely variable, with its area rapidly fluctuating and at times disappearing depending on global climatic conditions. At the end of the Ice ages, estimated to have been around 10,500 BC, the Sahara had again become a green fertile valley, and its African populations returned from the interior and coastal highlands in Sub-Saharan Africa, with rock art paintings depicting a fertile Sahara and large populations discovered in Tassili n'Ajjer dating back perhaps 10 millennia. However, the warming and drying climate meant that by 5000 BC, the Sahara region was becoming increasingly dry and hostile. Around 3500 BC, due to a tilt in the earth's orbit, the Sahara experienced a period of rapid desertification. The population trekked out of the Sahara region towards the Nile Valley below the Second Cataract where they made permanent or semi-permanent settlements. A major climatic recession occurred, lessening the heavy and persistent rains in Central and Eastern Africa. Since this time, dry conditions have prevailed in Eastern Africa and, increasingly during the last 200 years, in Ethiopia.
The domestication of cattle in Africa preceded agriculture and seems to have existed alongside hunter-gatherer cultures. It is speculated that by 6000 BC, cattle were domesticated in North Africa. In the Sahara-Nile complex, people domesticated many animals, including the donkey and a small screw-horned goat which was common from Algeria to Nubia.
Between 10,000 and 9,000 BC, pottery was independently invented in the region of Mali in the savannah of West Africa.
In the steppes and savannahs of the Sahara and Sahel in Northern West Africa, people possibly ancestral to modern Nilo-Saharan and Mandé cultures started to collect wild millet, around 8000 to 6000 BC. Later, gourds, watermelons, castor beans, and cotton were also collected. Sorghum was first domesticated in Eastern Sudan around 4000 BC, in one of the earliest instances of agriculture in human history. Its cultivation would gradually spread across Africa, before spreading to India around 2000 BC. Sorghum was first domesticated in the. They also started making pottery and built stone settlements (e.g., Tichitt, Oualata). Fishing, using bone-tipped harpoons, became a major activity in the numerous streams and lakes formed from the increased rains. In West Africa, the wet phase ushered in an expanding rainforest and wooded savanna from Senegal to Cameroon. Between 9,000 and 5,000 BC, Niger–Congo speakers domesticated the oil palm and raffia palm. Black-eyed peas and voandzeia (African groundnuts), were domesticated, followed by okra and kola nuts. Since most of the plants grew in the forest, the Niger–Congo speakers invented polished stone axes for clearing forest.
Around 4000 BC, the Saharan climate started to become drier at an exceedingly fast pace. This climate change caused lakes and rivers to shrink significantly and caused increasing desertification. This, in turn, decreased the amount of land conducive to settlements and encouraged migrations of farming communities to the more tropical climate of West Africa. During the first millennium BC, a reduction in wild grain populations related to changing climate conditions facilitated the expansion of farming communities and the rapid adoption of rice cultivation around the Niger River.
By the first millennium BC, ironworking had been introduced in Northern Africa. Around that time it also became established in parts of sub-Saharan Africa, either through independent invention there or diffusion from the north and vanished under unknown circumstances around 500 AD, having lasted approximately 2,000 years, and by 500 BC, metalworking began to become commonplace in West Africa. Ironworking was fully established by roughly 500 BC in many areas of East and West Africa, although other regions didn't begin ironworking until the early centuries AD. Copper objects from Egypt, North Africa, Nubia, and Ethiopia dating from around 500 BC have been excavated in West Africa, suggesting that Trans-Saharan trade networks had been established by this date.


Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km² (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area and 20% of its land area. With 1.4 billion people as of 2021, it accounts for about 18% of the world's human population. Africa's population is the youngest amongst all the continents; the median age in 2012 was 19.7, when the worldwide median age was 30.4. Despite a wide range of natural resources, Africa is the least wealthy continent per capita and second-least wealthy by total wealth, behind Oceania. Scholars have attributed this to different factors including geography, climate, tribalism, colonialism, the Cold War, neocolonialism, lack of democracy, and corruption. Despite this low concentration of wealth, recent economic expansion and the large and young population make Africa an important economic market in the broader global context.
The continent is surrounded by the Mediterranean Sea to the north, the Isthmus of Suez and the Red Sea to the northeast, the Indian Ocean to the southeast and the Atlantic Ocean to the west. The continent includes Madagascar and various archipelagos. It contains 54 fully recognised sovereign states (countries), eight territories and two de facto independent states with limited or no recognition. Algeria is Africa's largest country by area, and Nigeria is its largest by population. African nations cooperate through the establishment of the African Union, which is headquartered in Addis Ababa.
Africa straddles the equator and the prime meridian. It is the only continent to stretch from the northern temperate to southern temperate zones. The majority of the continent and its countries are in the Northern Hemisphere, with a substantial portion and number of countries in the Southern Hemisphere. Most of the continent lies in the tropics, except for a large part of Western Sahara, Algeria, Libya and Egypt, the northern tip of Mauritania, the entire territories of Morocco, Ceuta, Melilla, and Tunisia which in turn are located above the tropic of Cancer, in the northern temperate zone. In the other extreme of the continent, southern Namibia, southern Botswana, great parts of South Africa, the entire territories of Lesotho and Eswatini and the southern tips of Mozambique and Madagascar are located below the tropic of Capricorn, in the southern temperate zone.
Africa is highly biodiverse; it is the continent with the largest number of megafauna species, as it was least affected by the extinction of the Pleistocene megafauna. However, Africa also is heavily affected by a wide range of environmental issues, including desertification, deforestation, water scarcity, pollution and other issues. These entrenched environmental concerns are expected to worsen as climate change impacts Africa. The UN Intergovernmental Panel on Climate Change has identified Africa as the continent most vulnerable to climate change.
The history of Africa is long, complex, and has often been under-appreciated by the global historical community. Africa, particularly Eastern Africa, is widely accepted as the place of origin of humans and the Hominidae clade (great apes). The earliest hominids and their ancestors have been dated to around 7 million years ago, including Sahelanthropus tchadensis, Australopithecus africanus, A. afarensis, Homo erectus, H. habilis and H. ergaster— the earliest Homo sapiens (modern human) remains, found in Ethiopia, South Africa, and Morocco, date to circa 233,000, 259,000, and 300,000 years ago respectively, and Homo sapiens is believed to have originated in Africa around 350,000–260,000 years ago. Due to being the longest inhabited continent, Africa is also considered by anthropologists to be the most genetically diverse continent on the planet.
Early human civilizations, such as Ancient Egypt and Carthage emerged in North Africa. Following a subsequent long and complex history of civilizations, migration and trade, Africa hosts a large diversity of ethnicities, cultures and languages. The last 400 years have witnessed an increasing European influence on the continent. Starting in the 16th century, this was driven by trade, including the Trans-Atlantic slave trade, which created large African diaspora populations in the Americas. From the late 19th century to  the early 20th century, European nations colonized almost all of Africa, with the exception of Ethiopia and Liberia, though Ethiopia would later be invaded and occupied by Italy from 1936 to 1941. Most present states in Africa emerged from a process of decolonisation following World War II.


Afri was a Latin name used to refer to the inhabitants of then-known northern Africa to the west of the Nile river, and in its widest sense referred to all lands south of the Mediterranean (Ancient Libya). This name seems to have originally referred to a native Libyan tribe, an ancestor of modern Berbers; see Terence for discussion. The name had usually been connected with the Phoenician word ʿafar meaning "dust", but a 1981 hypothesis has asserted that it stems from the Berber word ifri (plural ifran) meaning "cave", in reference to cave dwellers. The same word may be found in the name of the Banu Ifran from Algeria and Tripolitania, a Berber tribe originally from Yafran (also known as Ifrane) in northwestern Libya, as well as the city of Ifrane in Morocco.
Under Roman rule, Carthage became the capital of the province it then named Africa Proconsularis, following its defeat of the Carthaginians in the Third Punic War in 146 BC, which also included the coastal part of modern Libya. The Latin suffix -ica can sometimes be used to denote a land (e.g., in Celtica from Celtae, as used by Julius Caesar). The later Muslim region of Ifriqiya, following its conquest of the Byzantine (Eastern Roman) Empire's Exarchatus Africae, also preserved a form of the name.
According to the Romans, Africa lies to the west of Egypt, while "Asia" was used to refer to Anatolia and lands to the east. A definite line was drawn between the two continents by the geographer Ptolemy (85–165 AD), indicating Alexandria along the Prime Meridian and making the isthmus of Suez and the Red Sea the boundary between Asia and Africa. As Europeans came to understand the real extent of the continent, the idea of "Africa" expanded with their knowledge.
Other etymological hypotheses have been postulated for the ancient name "Africa":
The 1st-century Jewish historian Flavius Josephus (Ant. 1.15) asserted that it was named for Epher, grandson of Abraham according to Gen. 25:4, whose descendants, he claimed, had invaded Libya.
Isidore of Seville in his 7th-century Etymologiae XIV.5.2. suggests "Africa comes from the Latin aprica, meaning "sunny".
Massey, in 1881, stated that Africa is derived from the Egyptian af-rui-ka, meaning "to turn toward the opening of the Ka." The Ka is the energetic double of every person and the "opening of the Ka" refers to a womb or birthplace. Africa would be, for the Egyptians, "the birthplace."
Michèle Fruyt in 1976 proposed linking the Latin word with africus "south wind", which would be of Umbrian origin and mean originally "rainy wind".
Robert R. Stieglitz of Rutgers University in 1984 proposed: "The name Africa, derived from the Latin *Aphir-ic-a, is cognate to Hebrew Ophir ['rich']."
Ibn Khallikan and some other historians claim that the name of Africa came from a Himyarite king called Afrikin ibn Kais ibn Saifi also called "Afrikus son of Abraham" who subdued Ifriqiya.
Arabic afrīqā (feminine noun) and ifrīqiyā, now usually pronounced afrīqiyā (feminine) 'Africa', from ‘afara [‘ = ‘ain, not ’alif] 'to be dusty' from ‘afar 'dust, powder' and ‘afir 'dried, dried up by the sun, withered' and ‘affara 'to dry in the sun on hot sand' or 'to sprinkle with dust'.
Possibly Phoenician faraqa in the sense of 'colony, separation'.


Africa is considered by most paleoanthropologists to be the oldest inhabited territory on Earth, with the Human species originating from the continent. During the mid-20th century, anthropologists discovered many fossils and evidence of human occupation perhaps as early as 7 million years ago (BP=before present). Fossil remains of several species of early apelike humans thought to have evolved into modern man, such as Australopithecus afarensis (radiometrically dated to approximately 3.9–3.0 million years BP, Paranthropus boisei (c. 2.3–1.4 million years BP) and Homo ergaster (c. 1.9 million–600,000 years BP) have been discovered.
After the evolution of Homo sapiens approximately 350,000 to 260,000 years BP in Africa, the continent was mainly populated by groups of hunter-gatherers. These first modern humans left Africa and populated the rest of the globe during the Out of Africa II migration dated to approximately 50,000 years BP, exiting the continent either across Bab-el-Mandeb over the Red Sea, the Strait of Gibraltar in Morocco, or the Isthmus of Suez in Egypt.
Other migrations of modern humans within the African continent have been dated to that time, with evidence of early human settlement found in Southern Africa, Southeast Africa, North Africa, and the Sahara.


At about 3300 BC, the historical record opens in Northern Africa with the rise of literacy in the Pharaonic civilization of Ancient Egypt. One of the world's earliest and longest-lasting civilizations, the Egyptian state continued, with varying levels of influence over other areas, until 343 BC. Egyptian influence reached deep into modern-day Libya and Nubia, and, according to Martin Bernal, as far north as Crete.
An independent centre of civilization with trading links to Phoenicia was established by Phoenicians from Tyre on the north-west African coast at Carthage.
European exploration of Africa began with Ancient Greeks and Romans. In 332 BC, Alexander the Great was welcomed as a liberator in Persian-occupied Egypt. He founded Alexandria in Egypt, which would become the prosperous capital of the Ptolemaic dynasty after his death.
Following the conquest of North Africa's Mediterranean coastline by the Roman Empire, the area was integrated economically and culturally into the Roman system. Roman settlement occurred in modern Tunisia and elsewhere along the coast. The first Roman emperor native to North Africa was Septimius Severus, born in Leptis Magna in present-day Libya—his mother was Italian Roman and his father was Punic.
Christianity spread across these areas at an early date, from Judaea via Egypt and beyond the borders of the Roman world into Nubia; by AD 340 at the latest, it had become the state religion of the Aksumite Empire. Syro-Greek missionaries, who arrived by way of the Red Sea, were responsible for this theological development.
In the early 7th century, the newly formed Arabian Islamic Caliphate expanded into Egypt, and then into North Africa. In a short while, the local Berber elite had been integrated into Muslim Arab tribes. When the Umayyad capital Damascus fell in the 8th century, the Islamic centre of the Mediterranean shifted from Syria to Qayrawan in North Africa. Islamic North Africa had become diverse, and a hub for mystics, scholars, jurists, and philosophers. During the above-mentioned period, Islam spread to sub-Saharan Africa, mainly through trade routes and migration.
In West Africa, Dhar Tichitt and Oualata in present-day Mauritania figure prominently among the early urban centers, dated to 2,000 BC. About 500 stone settlements litter the region in the former savannah of the Sahara. Its inhabitants fished and grew millet. It has been found by Augustin Holl that the Soninke of the Mandé peoples were likely responsible for constructing such settlements. Around 300 BC the region became more desiccated and the settlements began to decline, most likely relocating to Koumbi Saleh. Architectural evidence and the comparison of pottery styles suggest that Dhar Tichitt was related to the subsequent Ghana Empire. Djenné-Djenno (in present-day Mali) was settled around 300 BC, and the town grew to house a sizable Iron Age population, as evidenced by crowded cemeteries. Living structures were made of sun-dried mud. By 250 BC Djenné-Djenno had become a large, thriving market town.
Farther south, in central Nigeria, around 1,500 BC, the Nok culture developed on the Jos Plateau. It was a highly centralized community. The Nok people produced lifelike representations in terracotta, including human heads and human figures, elephants, and other animals. By 500 BC, and possibly earlier, they were smelting iron. By 200 AD the Nok culture had vanished. and vanished under unknown circumstances around 500 AD, having lasted approximately 2,000 years. Based on stylistic similarities with the Nok terracottas, the bronze figurines of the Yoruba kingdom of Ife and those of the Bini kingdom of Benin are suggested to be continuations of the traditions of the earlier Nok culture.


Pre-colonial Africa possessed perhaps as many as 10,000 different states and polities characterized by many different sorts of political organization and rule. These included small family groups of hunter-gatherers such as the San people of southern Africa; larger, more structured groups such as the family clan groupings of the Bantu-speaking peoples of central, southern, and eastern Africa; heavily structured clan groups in the Horn of Africa; the large Sahelian kingdoms; and autonomous city-states and kingdoms such as those of the Akan; Edo, Yoruba, and Igbo people in West Africa; and the Swahili coastal trading towns of Southeast Africa.
By the ninth century AD, a string of dynastic states, including the earliest Hausa states, stretched across the sub-Saharan savannah from the western regions to central Sudan. The most powerful of these states were Ghana, Gao, and the Kanem-Bornu Empire. Ghana declined in the eleventh century, but was succeeded by the Mali Empire which consolidated much of western Sudan in the thirteenth century. Kanem accepted Islam in the eleventh century.
In the forested regions of the West African coast, independent kingdoms grew with little influence from the Muslim north. The Kingdom of Nri was established around the ninth century and was one of the first. It is also one of the oldest kingdoms in present-day Nigeria and was ruled by the Eze Nri. The Nri kingdom is famous for its elaborate bronzes, found at the town of Igbo-Ukwu. The bronzes have been dated from as far back as the ninth century.
The Kingdom of Ife, historically the first of these Yoruba city-states or kingdoms, established government under a priestly oba ('king' or 'ruler' in the Yoruba language), called the Ooni of Ife. Ife was noted as a major religious and cultural centre in West Africa, and for its unique naturalistic tradition of bronze sculpture. The Ife model of government was adapted at the Oyo Empire, where its obas or kings, called the Alaafins of Oyo, once controlled a large number of other Yoruba and non-Yoruba city-states and kingdoms; the Fon Kingdom of Dahomey was one of the non-Yoruba domains under Oyo control.
The Almoravids were a Berber dynasty from the Sahara that spread over a wide area of northwestern Africa and the Iberian peninsula during the eleventh century. The Banu Hilal and Banu Ma'qil were a collection of Arab Bedouin tribes from the Arabian Peninsula who migrated westwards via Egypt between the eleventh and thirteenth centuries. Their migration resulted in the fusion of the Arabs and Berbers, where the locals were Arabized, and Arab culture absorbed elements of the local culture, under the unifying framework of Islam.
Following the breakup of Mali, a local leader named Sonni Ali (1464–1492) founded the Songhai Empire in the region of middle Niger and the western Sudan and took control of the trans-Saharan trade. Sonni Ali seized Timbuktu in 1468 and Jenne in 1473, building his regime on trade revenues and the cooperation of Muslim merchants. His successor Askia Mohammad I (1493–1528) made Islam the official religion, built mosques, and brought to Gao Muslim scholars, including al-Maghili (d.1504), the founder of an important tradition of Sudanic African Muslim scholarship. By the eleventh century, some Hausa states – such as Kano, jigawa, Katsina, and Gobir – had developed into walled towns engaging in trade, servicing caravans, and the manufacture of goods. Until the fifteenth century, these small states were on the periphery of the major Sudanic empires of the era, paying tribute to Songhai to the west and Kanem-Borno to the east.


Inflorescences usually have modified foliage different from the vegetative part of the plant. Considering the broadest meaning of the term, any leaf associated with an inflorescence is called a bract. A bract is usually located at the node where the main stem of the inflorescence forms, joined to the rachis of the plant, but other bracts can exist within the inflorescence itself. They serve a variety of functions which include attracting pollinators and protecting young flowers. According to the presence or absence of bracts and their characteristics we may distinguish the following:
Ebracteate inflorescences: No bracts in the inflorescence.
Bracteate inflorescences: The bracts in the inflorescence are very specialised, sometimes reduced to small scales, divided or dissected.
Leafy inflorescences: Though often reduced in size, the bracts are unspecialised and look like the typical leaves of the plant, so that the term flowering stem is usually applied instead of inflorescence. This use is not technically correct, as, despite their  'normal' appearance, these leaves are considered, in fact, bracts, so that 'leafy inflorescence' is preferable.
Leafy-bracted inflorescences: Intermediate between bracteate and leafy inflorescence.
If many bracts are present and they are strictly connected to the stem, like in the family Asteraceae, the bracts might collectively be called an involucre. If the inflorescence has a second unit of bracts further up the stem, they might be called an involucel.


There is no general consensus in defining the different inflorescences. The following is based on Focko Weberling's Morphologie der Blüten und der Blütenstände (Stuttgart, 1981). The main groups of inflorescences are distinguished by branching. Within these groups, the most important characteristics are the intersection of the axes and different variations of the model. They may contain many flowers (pluriflor) or a few (pauciflor). Inflorescences can be simple or compound.


As with leaves, flowers can be arranged on the stem according to many different patterns. See 'Phyllotaxis' for in-depth descriptions

Similarly arrangement of leaf in bud is called Ptyxis.
When a single or a cluster of flower(s) is located at the axil of a bract, the location of the bract in relation to the stem holding the flower(s) is indicated by the use of different terms and may be a useful diagnostic indicator.
Typical placement of bracts include:
Some plants have bracts that subtend the inflorescence, where the flowers are on branched stalks; the bracts are not connected to the stalks holding the flowers, but are adnate or attached to the main stem (Adnate describes the fusing together of different unrelated parts. When the parts fused together are the same, they are connately joined.)
Other plants have the bracts subtend the pedicel or peduncle of single flowers.
Metatopic placement of bracts include:
When the bract is attached to the stem holding the flower (the pedicel or peduncle), it is said to be recaulescent; sometimes these bracts or bracteoles are highly modified and appear to be appendages of the flower calyx. Recaulescences is the fusion of the subtending leaf with the stem holding the bud or the bud itself, thus the leaf or bract is adnate to the stem of flower.
When the formation of the bud is shifted up the stem distinctly above the subtending leaf, it is described as concaulescent.


An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches.  Morphologically, it is the modified part of the shoot of seed plants where flowers are formed on the axis of a plant. The modifications can involve the length and the nature of the  internodes and the phyllotaxis, as well as variations in the proportions, compressions, swellings, adnations, connations and reduction of main and secondary axes.
One can also define an inflorescence as the reproductive portion of a plant that bears a cluster of flowers in a specific pattern.
The stem holding the whole inflorescence is called a  peduncle. The major axis (incorrectly referred to as the main stem) above the peduncle bearing the flowers or secondary branches is called the rachis.  The stalk of each flower in the inflorescence is called a pedicel. A flower that is not part of an inflorescence is called a solitary flower and its stalk is also referred to as a peduncle. Any flower in an inflorescence may be referred to as a floret, especially when the individual flowers are particularly small and borne in a tight cluster, such as in a pseudanthium.
The fruiting stage of an inflorescence is known as an infructescence.
Inflorescences may be simple (single) or complex (panicle). The rachis may be one of several types, including single, composite, umbel, spike or raceme.