The great blue heron is the largest heron native to North America. Among all extant herons, it is surpassed in size only by the goliath heron (Ardea goliath) and the white-bellied heron (Ardea insignis). It exhibits a minor degree of sexual dimorphism; with males slightly larger than females, but otherwise the sexes are not easily outwardly distinguishable. It has head-to-tail length of 91–137 cm (36–54 in), a wingspan of 167–201 cm (66–79 in), a height of 115–138 cm (45–54 in), and a weight of 1.82–3.6 kg (4.0–7.9 lb). In British Columbia, adult males averaged 2.48 kg (5.5 lb) and adult females 2.11 kg (4.7 lb). In Nova Scotia and New England, adult herons of both sexes averaged 2.23 kg (4.9 lb), while in Oregon, both sexes averaged 2.09 kg (4.6 lb) Thus, great blue herons are roughly twice as heavy as great egrets (Ardea alba), although only slightly taller than them, but they weigh only about half as much as a large goliath heron.
Notable features of great blue herons include slaty (gray with a slight azure blue) flight feathers, red-brown thighs, and a paired red-brown and black stripe up the flanks; the neck is rusty-gray, with black and white streaking down the front; the head is paler, with a nearly white face, and a pair of black or slate plumes runs from just above the eye to the back of the head. The feathers on the lower neck are long and plume-like; it also has plumes on the lower back at the start of the breeding season. The bill is dull yellowish, becoming orange briefly at the start of the breeding season, and the lower legs are gray, also becoming orangey at the start of the breeding season. Immature birds are duller in color, with a dull blackish-gray crown, and the flank pattern is only weakly defined; they have no plumes, and the bill is dull gray-yellow. Among standard measurements, the wing chord is 43–49.2 cm (16.9–19.4 in), the tail is 15.2–19.5 cm (6.0–7.7 in), the culmen is 12.3–15.2 cm (4.8–6.0 in), and the tarsus is 15.7–21 cm (6.2–8.3 in). The heron's stride is around 22 cm (8.7 in), almost in a straight line. Two of the three front toes are generally closer together. In a track, the front toes, as well as the back, often show the small talons.
The subspecies differ only slightly in size and plumage tone, with the exception of A. h. occidentalis, native to South Florida, which also has a distinct white morph, known as the great white heron (not to be confused with the great egret, for which "great white heron" was once a common name). The great white heron differs from other great blues in bill morphology, head plume length, and in having a total lack of pigment in its plumage. It averages somewhat larger than the sympatric race A. h. wardi and may be the largest race in the species. In a survey of A. h. occidentalis in Florida, males were found to average 3.02 kg (6.7 lb) and females average 2.57 kg (5.7 lb), with a range for both sexes of 2 to 3.39 kg (4.4 to 7.5 lb). This is mainly found near salt water, and was long thought to be a separate species. Birds intermediate between the normal morph and the white morph are known as Würdemann's heron; these birds resemble a "normal" great blue with a white head.
The theory that great white herons may be a separate species (A. occidentalis) from the great blue heron has again been given some support by David Sibley.


The "great white heron" could be confused with the great egret (Ardea alba), but is larger, with yellow legs as opposed to the great egret's black legs. The reddish egret (Egretta rufescens) and little blue heron (Egretta caerulea) could be mistaken for the great blue heron, but are much smaller, and lack white on the head and yellow in the bill. In the southern reaches of its range, the great blue sometimes overlaps in range with the closely related and similarly sized cocoi heron (A. cocoi). The cocoi is distinguished by a striking white neck and solid black crown, but the duller juveniles are more easily confused. More superficially similar is the slightly smaller grey heron, which may sometimes vagrate to the northern coasts of North America. The grey heron (which occupies the same ecological niche in Eurasia as the great blue heron) has very similar plumage, but has a solidly soft-gray neck. Erroneously, the great blue heron is sometimes referred to as a "crane".  A heron is differentiated from a crane in flight. The crane's neck is straight and the heron's is always curved.


The great blue heron is found throughout most of North America, as far north as Alaska and the southern Canadian provinces in the summer. In winter, the range extends south through Florida, Mexico, and the Caribbean to South America. Birds east of the Rocky Mountains in the northern part of their range are migratory and winter in the coastal areas of the Southern United States, Central America, or northern South America. From the Southern United States southwards, and on the lower Pacific coast, they are year-round residents. However, their hardiness is such that individuals often remain through cold northern winters, as well, so long as fish-bearing waters remain unfrozen (which may be the case only in flowing water such as streams, creeks, and rivers).
The great blue heron can adapt to almost any wetland habitat in its range. It may be found in numbers in fresh and saltwater marshes, mangrove swamps, flooded meadows, lake edges, or shorelines. It is quite adaptable and may be seen in heavily developed areas as long as they hold bodies of fish-bearing water.
Great blue herons rarely venture far from bodies of water, but are occasionally seen flying over upland areas. They usually nest in trees or bushes near water's edge, often on islands (which minimizes the potential for predation) or partially isolated spots.
It has been recorded as a vagrant in England, Greenland, Hawaii, and the Azores.
The great white heron is unique to South Florida, including Great White Heron National Wildlife Refuge in the Florida Keys.


The primary food for great blue heron is small fish. It is also known to opportunistically feed on larger fish and a wide range of shrimp, crayfish, crabs, aquatic insects (such as dragonflies), other insects (such as grasshoppers), rodents, and other small mammals, amphibians (such as frogs, toads, and salamanders), reptiles (such as lizards and snakes), and birds, especially ducklings.  Primary prey is variable based on availability and abundance. In Nova Scotia, 98% of the diet was flounder. In British Columbia, the primary prey species are sticklebacks, gunnels, sculpins, and perch. California herons were found to live mostly on sculpin, bass, perch, flounder, and top smelt. Non-piscine prey is rarely quantitatively important, though one study in Idaho showed that from 24 to 40% of the diet was made up of voles.
Herons locate their food by sight and usually swallow it whole. They have been known to choke on prey that is too large. It is generally a solitary feeder. Individuals usually forage while standing in water, but also feed in fields or drop from the air, or a perch, into water. Mice are occasionally preyed on in upland areas far from the species' typical aquatic environments. Occasionally, loose feeding flocks form and may be beneficial since they are able to locate schools of fish more easily.
As large wading birds, great blue herons are capable of feeding in deeper waters, thus are able to harvest from niche areas not open to most other heron species. Typically, the great blue heron feeds in shallow waters, usually less than 50 cm (20 in) deep, or at the water's edge during both the night and the day, but especially around dawn and dusk. The most commonly employed hunting technique of the species is wading slowly with its long legs through shallow water and quickly spearing fish or frogs with its long, sharp bill. Although usually ponderous in movements, the great blue heron is adaptable in its fishing methods. Feeding behaviors variably have consisted of standing in one place, probing, pecking, walking at slow speeds, moving quickly, flying short distances and alighting, hovering over water and picking up prey, diving headfirst into the water, alighting on water feet-first, jumping from perches feet-first, and swimming or floating on the surface of the water.


This species usually breeds in colonies, in trees close to lakes or other wetlands. Adults generally return to the colony site after winter from December (in warmer climes such as California and Florida) to March (in cooler areas such as Canada). Usually, colonies include only great blue herons, though sometimes they nest alongside other species of herons. These groups are called a heronry (a more specific term than "rookery"). The size of these colonies may be large, ranging between five and 500 nests per colony, with an average around 160 nests per colony. A heronry is usually relatively close, usually within 4 to 5 km (2.5 to 3.1 mi), to ideal feeding spots. Heronry sites are usually difficult to reach on foot (e.g., islands, trees in swamps, high branches, etc.) to protect from potential mammalian predators. Trees of any type are used when available. When not, herons may nest on the ground, sagebrush, cacti, channel markers, artificial platforms, beaver mounds, and duck blinds. Other waterbirds (especially smaller herons) and, occasionally, even fish and mammal-eating raptors may nest amongst colonies.
Although nests are often reused for many years and herons are socially monogamous within a single breeding season, individuals usually choose new mates each year. Males arrive at colonies first and settle on nests, where they court females; most males choose a different nest each year. Great blue herons build a bulky stick nest. Nests are usually around 50 cm (20 in) across when first constructed, but can grow to more than 120 cm (47 in) in width and 90 cm (35 in) deep with repeated use and additional construction. If the nest is abandoned or destroyed, the female may lay a replacement clutch. Reproduction is negatively affected by human disturbance, particularly during the beginning of nesting. Repeated human intrusion into nesting areas often results in nest failure, with abandonment of eggs or chicks. However, Vancouver B.C. Canada's Stanley Park has had a healthy colony for some years right near its main entrance and tennis courts adjacent to English Bay and not far from Lost Lagoon. The park's colony has had as many as 183 nests.
The female lays 3 to 6 pale blue eggs, which can measure from 50.7 to 76.5 mm (2.00 to 3.01 in) in length and 29 to 50.5 mm (1.14 to 1.99 in) in width, though the smallest eggs in the above sample may have been considered "runt eggs" too small to produce viable young. Egg weights range from 61 to 80 g (2.2 to 2.8 oz). One brood is raised each year. First broods are laid generally from March to April. Eggs are usually laid at two-day intervals, incubated  around 27 days, and hatch asynchronously over a period of several days. Males incubate for about 10.5 hours of each day, while females usually incubate for the remainder of each day and the night, with eggs left without incubation for about 6 minutes of each hour.
The first chick to hatch usually becomes more experienced in food handling and aggressive interactions with siblings, so often grows more quickly than the other chicks. Both parents feed the young at the nest by regurgitating food. Parent birds have been shown to consume up to four times as much food when they are feeding young chicks (about 4300 kJ/day) than when laying or incubating eggs (about 1200 kJ/day). By the time they are 45 days old, the young weigh 86% of the adult's mass. After about 55 days at the northern edge of the range (Alberta) and 80 days at the southern edge of the range (California), young herons take their first flight. They return to the nest to be fed for about another 3 weeks, following adults back from foraging grounds, and are likely to gradually disperse away from their original nest over the course of the ensuing winter. Young herons are not as successful at fish capture as adults, as strike rates are similar, but capture rates are about half that of adults during the first 2 months after fledging.


Predators of eggs and nestlings include turkey vultures (Cathartes aura), common ravens (Corvus corax), and American crows (Corvus brachyrhynchos). Red-tailed hawks (Buteo jamaicensis), American black bears (Ursus americanus), and raccoons (Procyon lotor) are known to take larger nestlings or fledglings, and in the latter predator, many eggs. Adult herons, due to their size, have few natural predators, but a few of the larger avian predators have been known to kill both young and adults, including bald eagles (Haliaeetus leucocephalus) (the only predator known to attack great blue herons at every stage of their lifecycle from in the egg to adulthood), golden eagles (Aquila chrysaetos), and less frequently, great horned owls (Bubo virginianus) and Harris's hawks (Parabuteo unicinctus).
An occasional adult heron, or more likely, an unsteady fledgling, may be snatched by an American alligator (Alligator mississippiensis) or an American crocodile (Crocodylus acutus). Using its considerable size and dagger-like bill, a full-grown heron can be a formidable foe to a predator. In one instance, during an act of attempted predation by a golden eagle, a heron was able to mortally wound the eagle, although it succumbed to injuries sustained in the fight. When predation on an adult or chick occurs at a breeding colony, the colony can sometimes be abandoned by the other birds. The primary source of disturbance and breeding failures at heronries is human activities, mostly through human recreation or habitat destruction, as well as by egg-collectors and hunters.


John James Audubon illustrates the great blue heron in Birds of America, Second Edition (published, London 1827–38) as Plate 161. The image was engraved and colored by Robert Havell's London workshops. The original watercolor by Audubon was purchased by the New-York Historical Society.


The great blue heron (with its color changed to orange) is the basis of logos for the Delmarva Shorebirds minor league baseball team from the team's 1996 inception.
Great white herons feature prominently in the logo for the Major League Soccer club Inter Miami CF.  They were chosen for their local connection, as well as their quickness when hunting.


The intermediate egret, median egret, smaller egret, or yellow-billed egret (Ardea intermedia) is a medium-sized heron.  Some taxonomists put the species in the genus Egretta or Mesophoyx.  It is a resident breeder from east Africa across the Indian subcontinent to Southeast Asia and Australia.


Some authorities classify the intermediate egret in its own monotypic genus, Mesophoyx, while others place it with the smaller egrets in Egretta.
There are three recognised subspecies, and these are sometimes raised in to species:
A. i. brachyrhyncha Brehm, 1854 "yellow-billed egret" - sub-Saharan Africa
A.i. intermedia Wagler, 1827 "intermediate egret" - Asia from the Russian Far East to Japan to India and the Greater Sundas
A. i. plumifera Gould, 1848 "plumed egret" - eastern Indonesia, New Guinea and Australia.
A.i. intermedia differs from A.i. brachyrhyncha and A. i. plumifera by having a black bill when in breeding plumage, while A.i. plumata has a yellow-and-pink bill and A. i. brachyrhyncha has much yellower lores and face. A further difference between the nominate subspecies and the "yellow-billed" subspecies are that the "intermediate" subspecies has black at the top of the legs compared to reddish in the yellow-billed egret.


This species, as its scientific name implies, is intermediate in size between the great egret and smaller white egrets like the little egret and cattle egret, though nearer to little than great.  It is about 56–72 cm (22–28 in) long with a 105–115 cm (41–45 in) wingspan and weighs c. 400 g (14 oz), with all-white plumage, generally dark legs and a thickish yellow bill. Breeding birds may have a reddish or black bill, greenish yellow gape skin, loose filamentous plumes on their breast and back, and dull yellow or pink on their upper legs (regional variations).  The sexes are similar.


The non-breeding colours are similar, but the intermediate is smaller, with neck length a little less than body length, a slightly domed head, and a shorter, thicker bill. The great egret has a noticeable kink near the middle of its neck, and the top of its longer bill nearly aligns with the flat top of its head.  Close up, great egret's gape line extends behind the eye, while the intermediate's is less pointed and ends below the eye.  The intermediate tends to stalk upright with neck extended forward. The great is more patient, often adopting a sideways-leaning "one-eyed" stance.
Little egrets have yellow-soled feet and black bills. They often run after fish in shallow water. Breeding birds have long nuptial plumes on the back of their heads.


The intermediate egret stalks its prey methodically in shallow coastal or fresh water, including flooded fields. It eats fish, frogs, crustaceans and insects. It often nests in colonies with other herons, usually on platforms of sticks in trees or shrubs.  The typical clutch size is 2 or 3 eggs but there can be as many as 6 eggs in a clutch, the colour of the eggs is pale green, with a smooth, slightly pitted shell. Both parents incubate the eggs and they hatch after between 24 and 27 days. The eggs hatch asynchronously, after hatching the adults brood the semialtricial young for 12 days, defending the nests from aerial predators during both incubation and brooding, they crouch over the nest and raise their plumes and point their bill towards the threat. The parenst regurgitate the food to feed the young, initially onto the floor of the nest but later the chick takes the food from the parent's mouth. There may be competition for food within the brood. Pinfeathers may appear on the chicks as early as 4 days old and the young are able to leave the nest at 24 days old, although they return to be fed. Fledging occurs at around 40 sdays old and they leave the colony after 70 days. In Africa and Australia they are reported to be quite successful breeders with 96% and 88% of nests fledging at least a single chick, They are more successful in wet years than in dry years.


The white-necked heron or Pacific heron (Ardea pacifica) is a species of heron that is found on most of the Australian continent wherever freshwater habitats exist. It is also found in parts of Indonesia, New Guinea and New Zealand, but is uncommon in Tasmania. The populations of this species in Australia are known to be nomadic like most water birds in Australia, moving from one water source to another often entering habitats they have not previously occupied, taking advantage of flooding and heavy rain where the surplus of food allows them to breed and raise their young. Population explosions have been known when the environmental conditions are right for this species in places where they have been rare or unknown.


Large water dwelling bird, diurnal animal. Slate-grey to black with distinctive black spots in the centre of the lower fore-neck and throat. During breeding season plum coloured nuptial plumes are present on the back and breast.  Flight: stately, slow steady wing beats. Body size/length 76–106 cm, Wingspan 147–160 cm, Weight 860g.I  have studied the bird, with many photos, estimate wingspan near 2m
The bill is black and the facial skin is commonly blue or yellow. In this species of heron the eyes are green and the legs and feet also being black. Elongated neck and beak designed to reach out and catch fish, frogs, spiders and any other food source available in its habitat. Most of the head and neck is white with the black spots on the neck only visible in non-breeding individuals.
In the downy young, down is longer especially on the crown of the head with the neck and head being white and the upper wing and upperparts being a light grey brown. When juvenile feathers appear they are a dark grey. In Juveniles a broad strip of grey black runs down the front of the neck. Head and neck are commonly a brownish-grey tinge. The adult feathers emerge after wear replaces the juvenile feathers.


The white-necked heron is found throughout most of the Australian continent commonly residing in wetlands, tidal areas, shallow fresh waters, farm dams, clay pans, pastures and run off water in road side ditches.  The species has thrived since modifications were made to the Australian landscape through irrigation, dams and other man made water sources.
Not found in some parts of Western Australia and South Australia. Not found in arid zones, being the Great Sandy Desert, Gibson Desert, and Great Victoria Desert also the Nullarbor Plain.  Information regarding movement of this species is still unknown due to irruptive habits, which are commonly attributed to environmental conditions forcing the white-necked heron to act in a certain way, maintaining genetic adaptions to the harsh Australian environment in order to survive.


It has been suggested that the white-necked heron is a shy bird and very wary of its surrounds only keeping to areas where it has a clear view of what is around it. Commonly found in shallow wetlands usually seen foraging singly, also in pairs. Will defend feeding territories against other species aggressively. Observed harassing species of ibis (Threskiornis moluccus) and raptors, often stealing prey they have caught. When water sources dry up they fly to other water sources at an altitude of 30-100m. Habitat includes terrestrial wetlands and grasslands. Found where permanent water exists. The modified Australian landscape has favoured this species because they also commonly exist in farm dams and flooded pastures searching for prey. Babies start leaving the nest when they are ¾ grown. When young are born the parents alternate shifts to shade the young with their wings to keep them cool. First flight is generally around 6–7 weeks of maturity.


Secure existence in all states and territories of Australia.


The purple heron (Ardea purpurea) is a wide-ranging species of wading bird in the heron family, Ardeidae. The scientific name comes from Latin ardea "heron", and purpureus, "coloured purple". It breeds in Africa, central and southern Europe, and southern and eastern Asia. The Western Palearctic populations migrate between breeding and wintering habitats whereas the African and tropical-Asian populations are primarily sedentary, except for occasional dispersive movements.
It is similar in appearance to the more common grey heron but is slightly smaller, more slender and has darker plumage. It is also a more evasive bird, favouring densely vegetated habitats near water, particularly reed beds. It hunts for a range of prey including fish, rodents, frogs and insects, either stalking them or standing waiting in ambush.
Purple herons are colonial breeders and build a bulky nest out of dead reeds or sticks close to the water' edge among reeds or in dense vegetation. About five bluish-green eggs are laid and are incubated by both birds. The young hatch about four weeks later and fledge six weeks after that. The International Union for Conservation of Nature notes that the global population trend is downwards, largely because of the drainage of wetlands, but assesses the purple heron's conservation status as being of "least concern".


The purple heron is a large bird, 78–97 cm (31–38 in) in length with a standing height from 70 to 94 cm (28 to 37 in) and a 120–152 cm (47–60 in) wingspan. However, it is slender for its size, weighing only 0.5 to 1.35 kg (1.1 to 3.0 lb). It is somewhat smaller than the grey heron, from which it can be distinguished by its darker reddish-brown plumage, and, in adults, its darker grey back. Adults have the forehead and the crown of the head black, with a dark stripe down the back of the neck that terminates in a slender, dangling crest. This is shorter than the crest of the grey heron and does not exceed 140 mm (5.5 in). The sides of the head and the neck are buffish chestnut, with dark streaks and lines down either side of the whole the neck. The mantle is oily brown and the upper scapular feathers are elongated but not the lower ones. The rest of the upper parts and the tail are brownish grey. The front of the neck is paler than the sides and there are some elongated feathers at the base of the neck which are streaked with white, chestnut and black. The breast is chestnut brown, with some blackening at the side, and the belly and under-tail coverts are black. The brownish-yellow beak is long, straight and powerful, and is brighter in colour in breeding adults. The iris is yellow and the legs are brown at the front and yellowish behind.
The call is a harsh "frarnk", but is quieter and more high-pitched than that of the grey heron. It is generally a less noisy bird, though similar guttural sounds are heard emanating from the heronry. It is also less robust, and appears somewhat hollow-chested. Its head and neck are more slender and snake-like than the grey heron and its toes much longer. Unlike that bird, it often adopts a posture with its neck extending obliquely, and even nestlings tend to use this stance.


There are three or four subspecies:
A. p. purpurea – Linnaeus, 1766: nominate, found in Africa, Europe north to the Netherlands, and southwestern Asia east to Kazakhstan.
A. p. bournei – de Naurois, 1966: Bourne's heron. Found in the Cape Verde Islands (included in purpurea by some authors, but treated as a distinct species Ardea bournei by some others).
A. p. madagascariensis – van Oort, 1910: found in Madagascar.
A. p. manilensis – Meyen, 1834: found in Asia from the Indian Subcontinent east to the Philippines and Indonesia, and north to Primorsky Krai, Russia.


The purple heron has a mostly palaearctic distribution and breeds in Europe, Asia and Africa. The range of the western race extends from Portugal eastwards across much of central and southern Europe and parts of North Africa as far as Lake Balkhash in Kazakhstan. In Africa, the bird breeds in Senegal, down the east coast of Africa and in Madagascar. The eastern race extends from the Indian Subcontinent, eastwards to eastern China and the Philippines, and northwards to the basins of the Amur River and the Ussuri River at about 49°N. The southern race is restricted to Madagascar, and a small population of purple herons on the Cape Verde Islands is regarded as a separate race by some authorities.
Between August and October, birds of the western population migrate southwards to tropical Africa, returning northwards in March. Purple herons often overshoot their normal range on their return, and are vagrants to northern Europe including Britain. The eastern population is largely resident, though some birds from the northern part of the range fly southwards to Korea, Thailand and Malaysia. The African birds are resident.
The purple heron inhabits marshes, lagoons and lakes surrounded by dense vegetation. It may frequent mangrove swamps on the coast but it usually chooses freshwater habitats, particularly locations with reed beds (Phragmites). It also visits mudflats, river banks, ditches and canals. In the Cape Verde Islands, it is more likely to be seen in the open, on arid slopes.


Flight is slow, with the neck retracted and the legs extending a long way behind the tail. This is characteristic of herons and bitterns, and distinguishes them from storks, cranes and spoonbills, which extend their necks in flight. It is a secretive bird, spending less time out in the open than the grey heron and tending to skulk in reed beds. Its long toes mean it can walk on floating vegetation, and it sometimes walks over bushes in the same way, not making any attempt to grip the branches. It seldom perches in trees, preferring more terrestrial sites to rest.
It is most active at dawn and dusk, roosting with other birds in the middle of the day and at night, but increasing its diurnal activity while rearing young. It feeds in shallow water, grabbing its prey with its powerful beak. It will often wait motionless for prey, or slowly stalk its victim. The diet consists mainly of fish, small mammals and amphibians, but nestling birds, snakes, lizards, crustaceans, water snails, insects and spiders are also eaten. Terrestrial beetles are the commonest insect prey, followed by water beetles and aquatic larvae, with grasshoppers, dragonflies, bees and flies also being consumed.
The purple heron usually breeds in colonies but sometimes the nests are solitary. It sometimes associates with other species such as the goliath heron (Ardea goliath) or grey heron at multi-species nesting colonies. The site chosen is generally in reed beds, canebrakes or low bushes close to large lakes or other extensive wetlands. It builds a bulky nest of dead reeds, sticks or whatever is available, carelessly pulling the material together. The eggs are bluish-green, averaging 56 by 45 mm (2.20 by 1.77 in). The clutch is usually four or five eggs, with occasionally seven or eight eggs being laid, though these large clutches may have resulted from two females laying in the same nest. The eggs are laid at intervals of three days, and incubation may begin with the first egg, or start when the clutch is complete. Both parents share the incubation, which lasts between 24 and 28 days, and the care of the young. When an adult brings food, its beak is dragged down by the chicks and it regurgitates food from the crop onto the nest, or the young may take food directly from the beak. The young fledge at about six weeks and become independent at two months. They then disperse widely.


There are estimated to be a total of between 270,000 and 570,000 purple herons in the world and the population is probably decreasing slowly. The International Union for Conservation of Nature has assessed its conservation status as being of "least concern" because the rate of decline is insufficient to justify rating it in a more threatened category. The chief threat the bird faces is drainage and disturbance of its wetland habitats, particularly destruction of the reed beds. The purple heron is one of the species to which the Agreement on the Conservation of African-Eurasian Migratory Waterbirds (AEWA) applies.


The sooty shearwater (Ardenna grisea) is a medium-large shearwater in the seabird family Procellariidae. In New Zealand, it is also known by its Māori name tītī, and as muttonbird, like its relatives the wedge-tailed shearwater (A. pacificus) and the Australian short-tailed shearwater (A. tenuirostris).


The sooty shearwater was formally described in 1789 by the German naturalist Johann Friedrich Gmelin under the binomial name Procellaria grisea. The shearwater had been briefly described in 1777 by James Cook in the account of his second voyage to the Pacific, and in 1785 the English ornithologist John Latham had described a museum specimen. The sooty shearwater is now placed in the genus Ardenna, that was introduced in 1853 by Ludwig Reichenbach. The genus name Ardenna was used to refer to a seabird by Italian naturalist Ulisse Aldrovandi in 1603, and the specific epithet grisea is medieval Latin for "grey". The species is considered to be monotypic; no subspecies are recognised.
It appears to be particularly closely related to the great shearwater (A. gravis) and the short-tailed shearwater, all blunt-tailed, black-billed species, but its precise relationships are obscure. In any case, these three species are among the larger species of shearwaters that have been moved into a separate genus Ardenna based on a phylogenetic analysis of mitochondrial DNA.


Sooty shearwaters are 40–51 cm (16–20 in) in length with a 94–110 cm (37–43 in) wingspan. It has the typical "shearing" flight of the genus, dipping from side to side on stiff wings with few wing beats, the wingtips almost touching the water. Its flight is powerful and direct, with wings held stiff and straight, giving the impression of a very small albatross. This shearwater is identifiable by its dark plumage, which is responsible for its name. In poor viewing conditions, it looks all black, but in good light, it shows as dark chocolate-brown with a silvery strip along the center of the underwing.
Usually loud, sooty shearwaters coo and croak while on the breeding grounds.
In the Atlantic, it is the only such bird, whereas in the Pacific part of its range, other all-dark large shearwaters are found. The short-tailed shearwater in particular is almost impossible to tell apart from the present species at a distance.


Sooty shearwaters breed on small islands in the south Pacific and south Atlantic Oceans, mainly around New Zealand, the Falkland Islands, Tierra del Fuego, and in the Auckland Islands and Phillip Island off Norfolk Island. They start breeding in October, and incubate their young for about 54 days. Once the chick hatches, the parents raise their chick for 86 to 109 days.
They are spectacular long-distance migrants, following a circular route, traveling north up the western side of the Pacific and Atlantic Oceans at the end of the nesting season in March–May, reaching subarctic waters in June–July, where they cross from west to east, then return south down the eastern side of the oceans in September–October, reaching to the breeding colonies in November. They do not migrate as a flock, but rather as individuals, associating only opportunistically; in June 1906, for example, two were shot near Guadalupe Island off Baja California, Mexico, several weeks before the bulk of the population would pass by. Likewise, the identity of numerous large, dark shearwaters observed in October 2004 off Kwajalein in the Marshall Islands remain enigmatic; they might have been either sooty or short-tailed shearwaters, but neither species is generally held to pass through this region at that time.
In the Atlantic Ocean, they cover distances in excess of 14,000 km (8,700 mi) from their breeding colony on the Falkland Islands (52°S 60°W) north to 60 to 70°N in the North Atlantic Ocean off north Norway; distances covered in the Pacific are similar or larger; although the Pacific Ocean colonies are not quite so far south, at 35 to 50°S off New Zealand, and moving north to the Aleutian Islands, the longitudinal width of the ocean makes longer migrations necessary. Recent tagging experiments have shown that birds breeding in New Zealand may travel 74,000 km in a year, reaching Japan, Alaska, and California, averaging more than 500 km per day.
In Great Britain, they move south in late August and September; with strong north and north-west winds, they may occasionally become "trapped" in the shallow, largely enclosed North Sea, and heavy passages may be seen flying back north up the British east coast as they retrace their path back to the Atlantic over northern Scotland.


The sooty shearwater feeds on fish and squid. They can dive up to 68 m deep for food, but more commonly take surface food, in particular often following whales to catch fish disturbed by them. They also follow fishing boats to take fish scraps thrown overboard. Isotopic analyses revealed significant niche overlap between sooty shearwaters and great shearwaters.
They breed in huge colonies and the female lays one white egg. These shearwaters nest in burrows lined with plant material, which are visited only at night to avoid predation by large gulls. The architecture of sooty shearwater burrows can vary within and between breeding colonies, and is influenced by competition for breeding space and habitat type, with soil under dense tussac grass being easier to excavate than other substrates.
In New Zealand, about 250,000 muttonbirds are harvested for oils and food each year by the indigenous Māori population. Young birds just about to fledge are collected from the burrows, plucked, and often preserved in salt.  in 2022, climate change was thought to be impacting this cultural harvest by Ngāi Tahu.
Its numbers have been declining in recent decades, and it is presently classified as near threatened by the IUCN. In 2009, the harvest reported record-low catches, on average a trapping cage yielded nearly 500 birds; in 2009, the number was estimated to be closer to 40 per cage.


On August 18, 1961, the Santa Cruz Sentinel reported that thousands of crazed sooty shearwaters were sighted on the shores of North Monterey Bay in California, regurgitating anchovies, flying into objects, and dying on the streets. The incident sparked the interest of local resident Alfred Hitchcock, along with a story about spooky bird behavior by British writer Daphne du Maurier, helping to inspire Hitchcock's 1963 thriller The Birds, a cautionary tale of nature revolting against man. The film is now ranked among the American Film Institute's top-10 thrillers of the last century.
Scientists looking at the stomach contents of turtles and seabirds gathered in 1961 Monterey Bay ship surveys have found toxin-making algae were present in 79% of the plankton the creatures ate. "I am pretty convinced that the birds were poisoned," says ocean environmentalist Sibel Bargu of Louisiana State University. "All the symptoms were extremely similar to later bird poisoning events in the same area."
Plankton expert Raphael Kudela of USC points to leaky septic tanks installed amid a housing boom around Monterey Bay in the early 1960s as the ultimate culprit that may have fed the toxic algae "It is to some extent a natural phenomenon, and the best thing we can do is monitor for the presence of toxins, and treat impacted wildlife."


The wedge-tailed shearwater (Ardenna pacifica) is a medium-large shearwater in the seabird family Procellariidae. It is one of the shearwater species that is sometimes referred to as a muttonbird, like the sooty shearwater of New Zealand and the short-tailed shearwater of Australia. It ranges throughout the tropical Pacific and Indian Oceans, roughly between latitudes 35°N and 35°S. It breeds on islands off Japan, on the Islas Revillagigedo, the Hawaiian Islands, the Seychelles, the Northern Mariana Islands, and off Eastern and Western Australia.


The wedge-tailed shearwater is the largest of the tropical shearwaters. The two colour morphs of the species are dark and pale; the pale morphs predominate in the North Pacific, the dark morph elsewhere. However, both morphs exist in all populations, and bear no relation to sex or breeding condition. The pale morph has grey-brown plumage on the back, head, and upper wing, and whiter plumage below. The darker morph has the same dark grey-brown plumage over the whole body. The species' common name is derived from the large, wedge-shaped tail, which may help the species glide. The bill is dark and legs are salmon pink, with the legs set far back on the body (in common with the other shearwaters) as an adaptation for swimming.
This species is related to the pan-Pacific Buller's shearwater, which differs much in colour pattern, but also has a wedge tail and a thin, black bill. They make up the  Thyellodroma group, a superspecies of the large shearwaters that were for a long time included in the genus Puffinus.


Wedge-tailed shearwaters feed pelagically on fish, squid, and crustaceans. Their diet is 66% fish, of which the most commonly taken is goatfish. The species was thought to mostly take food from surface feeding, but observations of feeding wedge-tails suggested that contact-dipping, where birds flying close to the surface snatch prey from the water was the most commonly used hunting technique. However, a 2001 study which deployed maximum depth recorders found that 83% of wedge-tails dived during foraging trips with a mean maximum depth of 14 m (46 ft) and that they could achieve a depth of 66 m (217 ft).


The wedge-tailed shearwater breeds in colonies on small tropical islands. Breeding seasons vary depending on location, with synchronised breeding seasons more common at higher latitudes. Northern Hemisphere birds begin breeding around February, and Southern Hemisphere birds begin around September. Wedge-tailed shearwaters display natal philopatry, returning to their natal colony to begin breeding at the age of four.
Wedge-tailed shearwaters are monogamous, forming a pair bond that lasts for several years. Divorce between pairs occurs after breeding seasons that end in failure. Nesting occurs  either in burrows or sometimes on the surface under cover. Pairs call frequently as a pair, both to reinforce the pair bond and warn intruders away from their territory. Parents also call to their chicks. The call is long, with an inhaling component (OOO) and exhaling component (err); their Hawaiian name  'ua'u kani means moaning petrel.  Both sexes participate in digging a burrow, or repairing the burrow from last year. Nesting burrows of other species are also used. The breeding season of the Bonin petrel in Hawaii is timed to avoid that of the wedge-tail; in years where Bonin petrel chicks are still in burrows when wedge-tails return to begin breeding, these chicks are killed or evicted. It attends these colonies nocturnally, although nonbreeding wedge-tails are often seen at the surface throughout the day and breeding birds rest outside their burrows before laying.
Both sexes undertake a prelaying exodus  to build up energy reserves; this usually lasts around 28 days. A single egg is laid, if that egg is lost, then the pair will not attempt another that season. After laying, the male usually undertakes the first incubation stint. Both sexes incubate the egg, in stints that can last up to 13 days. Incubation takes around 50 days. After hatching, the chick is brooded for up to 6 days, until it is able to thermoregulate, after which it is left alone in the nest while both parents hunt for food. It is initially fed with stomach oil, an energy-rich, waxy oil of digested prey created in the parent's gut; later, it is fed both solids and stomach oil. Like many procellariids, wedge-tailed shearwater parents alternate long and short trips to provide food, with the parents alternating between short foraging trips (1–4 days) and long trips (about 8 days), the two parents coordinating their feeding effort. Chicks increase in size to 560 g (20 oz) (larger than the adults), then drop to around 430 g (15 oz) before fledging. Fledging occurs after 103–115 days, after which the chick is independent of the adult.
Known breeding colonies include:
Pacific Missile Range Facility, Kauai, Hawaii
Heron Island, Australia
Lady Elliot Island, Australia
Lord Howe Island, Australia
Montague Island, Southern New South Wales, Australia
North West Island, Australia
Muttonbird Island, Coffs Harbour, Northern New South Wales, Australia
Mānana Island, Hawaii, United States
Maui, Hawaii, United States
Mokuauia Island, Hawaii, United States
Kupikipiki’o Pt., Hawaii, United States
Ilot Maitre, Noumea, New Caledonia
Round Island, Mauritius
Ka'ena Point, Oahu, Hawaii
Mañagaha, Saipan, CNMI
San Benedicto Island, Revillagigedo Islands, Mexico
Alphonse Island, Republic of Seychelles
Bijoutier Island, Republic of Seychelles


The Chinese pond heron (Ardeola bacchus) is an East Asian freshwater bird of the heron family, (Ardeidae).
It is one of six species of birds known as "pond herons" (genus Ardeola). It is parapatric (or nearly so) with the Indian pond heron (A. grayii) to the west and the Javan pond heron (A. speciosa) to the south, and these three are presumed to form a superspecies. As a group they are variously affiliated with the squacco heron (A. ralloides) or the Malagasy pond heron (A. idae). As of mid-2011 there are no published molecular analyses of pond heron interrelationships and osteological data is likewise not analyzed for all relevant comparison taxa.


The Chinese pond heron is typically 47 cm (19 in) long with white wings, a yellow bill with a black tip, yellow eyes and legs. Its overall colour is red, blue and white during breeding season, and greyish-brown and flecked with white at other times.
It is found in shallow fresh and salt water wetlands and ponds in China and adjacent temperate and subtropical East Asia. Essentially a lowland bird, its range is delimited by the subarctic regions in the north, and by the mountain ranges in the west and south.
The species is prone to some vagrancy. One individual in breeding plumage was seen by the river at Bonzon near Gangaw – just inside the Chin State of Burma – west of the species' usual range, on April 8, 1995. A stray bird stopping over on Saint Paul Island, Alaska on August 4–9, 1997 was the first recorded occurrence of this species in North America.
Its food consists of insects, fish, and crustaceans. The Chinese pond heron often nests in mixed-species heronries. It lays a clutch of 3–6 blue-green eggs.
It is fairly common and not considered a threatened species by the IUCN.


The Indian pond heron or paddybird (Ardeola grayii) is a small heron. It is of Old World origins, breeding in southern Iran and east to the Indian subcontinent, Burma, and Sri Lanka. They are widespread and common but can be easily missed when they stalk prey at the edge of small water-bodies or even when they roost close to human habitations. They are however distinctive when they take off with bright white wings flashing in contrast to the cryptic streaked olive and brown colours of the body. Their camouflage is so excellent that they can be approached closely before they take to flight, a behaviour which has resulted in folk names and beliefs that the birds are short-sighted or blind.


They appear stocky with a short neck, short thick bill and buff-brown back. In summer, adults have long neck feathers. Its appearance is transformed from their dull colours when they take to flight, when the white of the wings makes them very prominent. It is very similar to the squacco heron, Ardeola ralloides, but is darker-backed. To the east of its range, it is replaced by the Chinese pond heron, Ardeola bacchus.
During the breeding season, there are records of individuals with red legs. The numbers do not suggest that this is a normal change for adults during the breeding season and some have suggested the possibility of it being genetic variants.
Erythristic plumage has been noted. The race phillipsi has been suggested for the populations found in the Maldives, however this is not always recognized. It forms a superspecies with the closely related Chinese pond heron, Javan pond heron and the Madagascar pond heron.
They are usually silent but may make a harsh croak in alarm when flushed or near their nests.
This bird was first described by Colonel W. H. Sykes in 1832 and given its scientific name in honour of John Edward Gray. Karyology studies indicate that pond herons have 68 chromosomes (2N).


The Indian pond heron's feeding habitat is marshy wetlands. They usually feed at the edge of ponds but make extensive use of floating vegetation such as water hyacinth to access deeper water. They may also on occasion swim on water or fish from the air and land in deeper waters. They have also been observed to fly and capture fishes leaping out of water. Sometimes, they fly low over water to drive frogs and fishes towards the shore before settling along the shoreline. They have been noted to pick up crumbs of bread and drop them on the water surface to bait fishes.
The primary food of these birds includes crustaceans, aquatic insects, fishes, tadpoles and sometimes leeches (Herpobdelloides sp.). Outside wetlands, these herons feed on insects (including crickets, dragonflies and bees), fish (Barilius noted as important in a study in Chandigarh) and amphibians.


The breeding season begins with the onset of the monsoons. They nest in small colonies, often with other wading birds, usually on platforms of sticks in trees or shrubs. Most nests are built at a height of about 9 to 10 m in large leafy trees. The nest material is collected by the male while the female builds the nest. Three to five eggs are laid. The eggs hatch asynchronously, taking 18 to 24 days to hatch. Both parents feed the young. Fish are the main diet fed to young. Nest sites that are not disturbed may be reused year after year.


They have few predators but injured birds may be taken by birds of prey.
An arbovirus that causes "Balagodu", trematodes and several other parasites have been isolated from the species. Antibodies to Japanese encephalitis and West Nile virus has been detected in pond herons and cattle egrets from southern India. Traces of heavy metals acquired from feeding in polluted waters may be particularly concentrated in the tail feathers.


Lamba B.S. (1963). "Nidification of some Indian birds. No.6. The Indian Pond Heron or Paddy bird Ardeola grayii (Sykes)". Pavo. 1 (1): 35–43.
de Boer LEM, van Brink JM (1982). "Cytotaxonomy of the Ciconiiformes (Aves), with karyotypes of eight species new to cytology". Cytogenet Cell Genet. 34 (1–2): 19–34. doi:10.1159/000131791. PMID 7151490.
Parasharya BM, Bhat HR (1987). "Unusual feeding strategies of the Little Egret and Pond Heron". Pavo. 25 (1–2): 13–16.


The kori bustard (Ardeotis kori) is the largest flying bird native to Africa. It is a member of the bustard family, which all belong to the order Otidiformes and are restricted in distribution to the Old World. It is one of the four species (ranging from Africa to India to Australia) in the large-bodied genus Ardeotis. In fact, the male kori bustard may be the heaviest living animal capable of flight.
This species, like most bustards, is a ground-dwelling bird and an opportunistic omnivore. Male kori bustards, which can be more than twice as heavy as the female, attempt to breed with as many females as possible and then take no part in the raising of the young. The nest is a shallow hollow in the earth, often disguised by nearby obstructive objects such as trees.


English naturalist William John Burchell described the kori bustard in 1822. The specific epithet kori is derived from the Tswana name for this bird – Kgori.
Two subspecies are currently recognized:
Ardeotis kori kori – the relatively pale nominate race from Botswana, Zimbabwe, Namibia, southern Angola, South Africa and Mozambique.
Ardeotis kori struthiunculus – the "Somali kori" distributed in Ethiopia, Uganda, South Sudan, Kenya and Tanzania. The two races are separated by the miombo woodlands of central Africa. This race has a more boldly patterned head and slightly more black and white patterning on the wings. The two races are similar in size, though A. k. struthiunculus may be slightly larger. This may be a distinct species.


The kori bustard is cryptically coloured, being mostly grey and brown, finely patterned with black and white coloring. The upper parts and neck are a vermiculated black and greyish-buff colour. The ventral plumage is more boldly colored, with white, black and buff. The crest on its head is blackish in coloration, with less black on the female's crest. There is a white eye stripe above the eye. The chin, throat and neck are whitish with thin, fine black barring. A black collar at the base of the hind-neck extends onto the sides of the breast. The feathers around the neck are loose, giving the appearance of a thicker neck than they really have. The belly is white and the tail has broad bands of brownish-gray and white coloration.  Their feathers contain light sensitive porphyrins, which gives their feathers a pinkish tinge at the base- especially noticeable when the feathers are shed suddenly. The head is large and the legs are relatively long. The eye is pale yellow, while the bill is light greenish horn coloured, relatively long, straight and rather flattened at the base. The legs are yellowish. The feet have three forward facing toes. Females are similar in plumage but are much smaller, measuring about 20-30% less in linear measurements and often weighing 2-3 times less than the male. The female is visibly thinner legged and slimmer necked. The juvenile is similar in appearance to the female, but is browner with more spotting on the mantle, with shorter crest and neck plumes. Male juveniles are larger than females and can be the same overall size as the adult male but tends to be less bulky with a thinner neck, shorter head crest, paler eyes and a darker mantle.


The male kori bustard has a length of 105 to 135 cm (3 ft 5 in to 4 ft 5 in) and a wingspan of 230 to 275 cm (7 ft 7 in to 9 ft 0 in). Male birds may typically weigh between 7 and 18 kg (15 and 40 lb). The average weight of adult males of the nominate race in Namibia (20 specimens) was 11.3 kg (25 lb), while A. k. struthiunculus males were found to average 10.9 kg (24 lb). The larger excepted males can scale up to 16 to 19 kg (35 to 42 lb) and a few exceptional specimens may weigh up to at least 20 kg (44 lb). Reports of outsized specimens weighing 23 kg (51 lb), 34 kg (75 lb) and even "almost" 40 kg (88 lb) have been reported, but none of these giant sizes have been verified and some may be from unreliable sources. Among bustards, only male great bustards (Otis tarda) achieve similarly high weights (the mean mass of males in these species is roughly the same) making the male kori and great not only the two largest bustards but also arguably the heaviest living flying animals. As a whole, other species, such as Dalmatian pelican (Pelecanus crispus), Andean condor (Vultur gryphus) and trumpeter swan (Cygnus buccinator), in which the larger males might match the average weight of the largest bustards and may weigh more on average between the sexes as they are less sexually dimorphic in mass than the giant bustards. Other than a 23 kg (51 lb) mute swan (Cygnus olor), the maximum size of the large bustards exceeds that of other flying birds. Other flying African birds (excluding rare vagrant pelicans and vultures to northernmost Africa) rival the average weight between the sexes of Kori bustards, namely great white pelicans (Pelecanus onocrotalus) and Cape vultures (Gyps coprotheres) while wattled cranes (Bugeranus carunculatus) lag slightly behind these on average. The female kori bustard weighs an average of 4.8 to 6.1 kg (11 to 13 lb), with a full range of 3 to 7 kg (6.6 to 15.4 lb). Females of the nominate race (35 specimens) in Namibia weighed a mean of 5.62 kg (12.4 lb), while females from A. k. struthiunculus weighed a mean of 5.9 kg (13 lb). Female length is from 90 to 112 cm (2 ft 11 in to 3 ft 8 in) while their wingspan is around 177 to 220 cm (5 ft 10 in to 7 ft 3 in). The standard measurements of the male include a wing chord of 69.5 to 83 cm (27.4 to 32.7 in), a tail measures from 35.8–44.7 cm (14.1–17.6 in), a culmen from 9.5 to 12.4 cm (3.7 to 4.9 in) and a tarsus from 20 to 24.7 cm (7.9 to 9.7 in). Meanwhile, the female's standard measurements are a wing chord of 58.5 to 66.5 cm (23.0 to 26.2 in), a tail of 30.7 to 39.5 cm (12.1 to 15.6 in), a culmen from 7 to 10.4 cm (2.8 to 4.1 in) and a tarsus from 16 to 19.5 cm (6.3 to 7.7 in). Body mass can vary considerably based upon rain conditions.


The size and dark crest are generally diagnostic amongst the bustards found in the kori bustard's range. However, East Africa holds the greatest diversity of bustards anywhere, including some other quite large species, and these have the potential to cause confusion. Kori bustards are distinguished from Denham's bustard (Neotis denhamii) and Ludwig's bustard (Neotis luwigii), both of which they sometimes forage with, by their greyer appearance and by their lack of a tawny red hind-neck and upper mantle. In flight it can be distinguished from both of these somewhat smaller bustards by not displaying any white markings on the upperwing, which is uniformly grey here. Both Stanley's and Ludwig's bustards lack the kori's dark crest. More similar to, and nearly the same size as, the kori is the closely related Arabian bustard (Ardeotis arabs) (despite its name, the latter species ranges well into East Africa). However, the Arabian species has white-tipped wing coverts, a browner back and very fine neck vermiculations and also lacks the black base to the neck and the black in the wing coverts as seen in the kori.


The kori bustard is found throughout southern Africa, except in densely wooded areas. They are common in Botswana and Namibia, extending into southern Angola and marginally into southwestern Zambia. In Zimbabwe they are generally sparse but locally common, particularly on the central plateau. Their distribution range extends along the Limpopo River valley into southern Mozambique and the eastern lowveld of South Africa. In South Africa they are also infrequent to rare in the Free State, North West and Northern Cape Provinces, extending southwards into the interior of the Western and Eastern Cape Provinces. Kori bustards are absent from the coastal lowlands along the south and east of South Africa and from high mountainous areas. This species is common in Tanzania at Ngorongoro National Park, Kitulo National Park and Serengeti National Park. A geographically disjunct population also occurs in the deserts and savanna of northeastern Africa. Here, the species ranges from extreme southeast South Sudan, north Somalia, Ethiopia through all of Kenya (except coastal regions), Tanzania and Uganda. Kenya may hold the largest population of kori bustards of any country and it can even border on abundant in the North Eastern Province. They are usually residential in their range, with some random, nomadic movement following rainfall.
This species occurs in open grassy areas, often characterized by sandy soil, especially Kalahari sands, and short grass usually near the cover of isolated clumps of trees or bushes. It may be found in plains, arid plateaus, highveld grassland, arid scrub, lightly wooded savanna, open dry bushveld and semi-desert. Where this species occurs, annual rainfall is quite low, between 100 and 600 mm (3.9 and 23.6 in). Breeding habitat is savanna in areas with sparse grass cover and scattered trees and shrubs. When nesting they sometimes use hilly areas. They follow fires or herds of foraging ungulates, in order to pick their various foods out of the short grasses. They may also be found in cultivated areas, especially wheat fields with a few scattered trees. This bustard is not found in well-wooded and forested areas due to the fact that it needs a lot of open space in which to take off. In arid grassland areas it is found along dry watercourses where patches of trees offer shade during the heat of the day.


Kori bustards spend most of their time on the ground, with up to 70% of their time being on foot, although they do occasionally forage in low bushes and trees. This bustard is a watchful and wary bird. Their behavior varies however, and they are usually very shy, running or crouching at the first sign of danger; at other times they can be completely fearless of humans. They have a hesitant, slow manner of walking, and when they detect an intruder they try to escape detection by moving off quietly with the head held at an unusual angle of between 45° and 60°. Being a large and heavy bird, it avoids flying if possible. When alarmed it will first run and, if pushed further, will take to the air on the run with much effort, its wings making heavy wingbeats. Once airborne it flies more easily with slow, measured wingbeats, with the neck extended and the legs folded. It usually remains low and lands again within sight. When they land, kori bustards keep their wings spread and only fold them when the bird has slowed down to a walking speed. Kori bustards have no preen gland, so to keep clean, they produce a powder down. Sun bathing and dust bathing are practiced. This bird has a loud, booming mating call which is often uttered just before dawn and can be heard from far away. Mostly residential, kori bustards may engage in nomadic movements. These migratory movements are probably influenced by rainfall and there is no evidence suggesting any regular pattern. These local migrations take place at night but have not been mapped. In the Etosha National Park these birds have been recorded moving up to 85 km (53 mi) from mopane woodland to open grassland plains and returning again the following season. Trial satellite tagging of one male kori bustard by the National Museums of Kenya demonstrated a migration along the Rift Valley between Tanzania and southeastern South Sudan. Additionally, adult and juvenile males move after the breeding season, whereas females do not appear to do so. Generally the kori bustard feeds during the morning and in the evening, spending the rest of the day standing still in any available shade.


Less vocal than other bustards, the kori bustard is generally silent but, when alarmed, both sexes emit a loud growling bark. This is described as a ca-caa-ca call, repeated several times for up to 10 minutes. This call carries long distances. This call is most often given by females with young and males during agnostic encounters. Chicks as young as two weeks will also emit this alarm call when startled. The male's mating call is a deep, resonant woum-woum-woum-woum or oom-oom-oom or wum, wum, wum, wum, wummm. This call ends with the bill snapping which is only audible at close range. Outside of the breeding display, kori bustards are often silent. A high alarm call, generally uttered by females, is sometimes heard. They may utter a deep vum on takeoff.


During the mating season, these birds are usually solitary but for the breeding pair. Otherwise, they are somewhat gregarious, being found in groups often including 5 to 6 birds but occasionally groups can number up to 40 individuals. Larger groups may be found around an abundant food source or at watering holes. In groups, birds are often fairly far apart from each other, often around a distance of 100 m (330 ft). Foraging groups are often single-sex. Such groups do not last long and often separate after a few days. These groups are believed advantageous both in that they may ensure safety in numbers against predation and may bring the bustards to prime food sources.


Walking slowly and sedately, they forage by picking at the ground with the bills and are most active in the first and last hours of daylight. Kori bustards are quite omnivorous birds. Insects are an important food source, with common species such as locusts, grasshoppers, dung beetles (Scarabaeus ssp.) and caterpillars being most often taken. They may follow large ungulates directly to catch insects flushed out by them or to pick through their dung for edible invertebrates. During outbreaks of locusts and caterpillars, kori bustards are sometimes found feeding on them in numbers. Other insect prey can include bush-crickets (Tettigonia ssp.), termites, hymenopterans and solifuges. Scorpions and molluscs may be taken opportunistically as well.
Small vertebrates may also be taken regularly, including lizards, chameleons, small snakes, small mammals (especially rodents) and bird eggs and nestlings. They may occasionally eat carrion, especially from large animals killed in veld fires. Plant material is also an important food. Grasses and their seeds are perhaps the most prominent plant foods, but they may also eat seeds, berries, roots, bulbs, flowers, wild melons and green leaves. This bustard is very partial to Acacia gum. This liking has given rise to the Afrikaans common name Gompou or, literally translated, "gum peacock". They drink regularly when they can access water but they can be found as far as 40 km (25 mi) from water sources. Unusually, they suck up rather than scoop up water.


The breeding seasons of the two subspecies of Kori bustards are distinguishable. In general, A. k. struthiunculus breeds from December to August and A. k. kori breeds from September to February. Breeding is closely tied with rainfall, and in drought years, may be greatly reduced or not even occur.
Kori bustards engage in lek mating. All bustards have polygynous breeding habits, in which one male displays to attract several females, and mates with them all. Males display at regularly used sites, each male utilizing several dispersed leks or display areas. These displays usually take place in the mornings and evenings. The courtship displays of the males are impressive and elaborate, successfully advertising their presence to potential mates. The males hold their heads backwards, with cheeks bulging, the crest is held erect, the bill open and they inflate their gular pouches, forming a white throat "balloon". During this display the oesophagus inflates to as much as four times its normal size and resembles a balloon. They also puff out their frontal neck feathers which are splayed upwards showing their white underside. The white may be visible up to 1 km (0.62 mi) away during display. Their wings are drooped and their tails are raised upwards and forwards onto their backs like a turkey, the rectrices being held vertically and their undertail coverts fluffed out. They enhance their performance with an exaggerated bouncing gait. When displaying they stride about with their necks puffed out, their tail fanned and their wings planed and pointed downward. They also emit a low-pitched booming noise when the neck is at maximum inflation and snap their bills open and shut. Several males dispersed over a wide area gather to display but usually one is dominant and the others do not display in his presence and move away. The displaying males are visited by the females who presumably select the male with the most impressive display. Occasionally fights between males can be serious during the mating season when display areas are being contested, with the two competitors smashing into each other's bodies and stabbing each other with their bills. They may stand chest-to-chest, tails erect, bills locked and "push" one another for up to 30 minutes.
Following the display, the copulation begins with the female lying down next to the dominant displaying male. He stands over her for 5–10 minutes, stepping from side to side and pecking her head in a slow, deliberate fashion, tail and crest feathers raised. She recoils at each peck. He then lowers himself onto his tarsi and continues pecking her until he shuffles forward and mounts with wings spread. Copulation lasts seconds after which both stand apart and ruffle their plumage. The female then sometimes barks and the male continues with his display.
As with all bustards, the female makes no real nest. The female kori bustard lays her eggs on the ground in a shallow, unlined hollow, rather than the typical scrape. This nest is usually located within 4 m (13 ft) of a tree or shrub, termite mound or an outcrop of rocks. The hollow may measure 300–450 mm (12–18 in) in diameter and be almost completely covered by the female when she's incubating. Due to their ground location, nests are often cryptic and difficult for a human to find, unless stumbled onto by chance. The same site is sometimes reused in successive years. The kori bustard is a solitary nester and there is no evidence of territoriality amongst the females. Usually two eggs are laid, though seldom 1 or 3 may be laid. Clutch size is likely correlated to food supply. They are cryptically colored with the ground color being dark buff, brown or olive and well marked and blotched with shades of brown, grey and pale purple. Eggs are somewhat glossy or waxy and have a pitted-looking surface. Egg size is 81 to 86 mm (3.2 to 3.4 in) in height and 58 to 61 mm (2.3 to 2.4 in) width. The eggs weigh individually about 149 g (5.3 oz), with a range of 121 to 178 g (4.3 to 6.3 oz).
The female, who alone does all the brooding behavior without male help, stays at the nest 98% of the time, rarely eating and never drinking. Occasionally she stretches her legs and raises her wings overhead. The female regularly turns the eggs with her bill. The female's plumage is drab and earth-colored, which makes her well camouflaged. She occasionally picks up pieces of vegetation and drops them on her back to render her camouflage more effective. If they need to feed briefly, the females go to and from the nest with a swift, silent crouching walk. If approached the incubating bird either slips unobtrusively from the nest or sits tight, only flying off at the last moment. The incubation period is 23 to 30 days, though is not known to exceed 25 days in wild specimens. The young are precocial and very well camouflaged. The lores are tawny, the crown tawny mottled black. A broad white supercilium bordered with black meets on the nape, extending down the centre of the nape. The neck is white with irregular black stripes from behind the eye and from the base of the lower mandibles. The upper parts are tawny and black with 3 black lines running along the back. The underparts are whitish. When the chicks hatch, the mother brings them a steady stream of food, most of it soft so the chicks can eat it easily. Captive hatchlings weigh 78 to 116 g (2.8 to 4.1 oz) on their first day but grow quickly. The precocial chicks are able to follow their mother around several hours after hatching. After a few weeks, the young actively forage closely with their mothers. They fledge at 4 to 5 weeks old, but are not self-assured fliers until 3 to 4 months. On average, around 67% of eggs successfully hatch (testimony to the effective camouflage of nests) and around one of the two young survive to adulthood. In Namibia and Tanzania, breeding success has been found to be greatly reduced during times of drought. Most young leave their mothers in their second year of life, but do not start breeding until they are fully mature at three to four years old in both sexes in studies conducted both of wild and captive bustards. The lifespan of wild kori bustards is not known but they may live to at least 26 or possibly 28 years old in captivity.


The kori bustard is often found in areas with a large quantity of antelope and other game. In Tanzania, kori bustards regularly attend blue wildebeest (Connochaetes taurinus) herds and feed on the small mammals and insects disturbed by them. Sometimes kori bustards are found with southern carmine bee-eaters (Merops nubicoides) and northern carmine bee-eaters (Merops nubicus) riding on their backs as they stride through the grass. The bee-eaters make the most of their walking perch by hawking insects from the bustard's back that are disturbed by the bustard's wandering. This is regularly seen in Chobe National Park, Botswana but has only been reported once elsewhere. There is also one record of fork-tailed drongos (Dicrurus adsimilis) perching on their backs in a similar manner. Kori bustards have been observed to behave aggressively to non-threatening animals at watering holes, as they may raise their crests, open their wings and peck aggressively. They have been seen acting aggressively towards red-crested korhaans (Eupodotis ruficrista), springbok (Antidorcas marsupialis), plains zebra (Equus quagga), and gemsbok (Oryx gazella). When kept in captivity, kori bustards have been kept together with numerous other (typically African) species in close quarters. Fifteen other bird species and 12 mammals successfully cohabitated with them (including rhinoceros). However, the bustards sometimes injure or kill the young of everything from waterfowl to dik-diks and may be killed by larger species from ostriches (Struthio camelus) to zebras (Equus spp.).
Being a large, ground-dwelling bird species, the kori bustard is vulnerable to many of Africa's myriad of formidable terrestrial predators. Leopards (Panthera pardus), caracals (Caracal caracal), cheetahs (Acinonyx jubatus), lions (Panthera leo), spotted hyena (Crocuta crocuta), African rock pythons (Python sebae), jackals (Canis spp.), Verreaux's eagle (Aquila verreauxii) and martial eagles (Polemaetus bellicosus) are amongst their potential natural predators. Young bustards are most at risk even to bigger predators, but animals of similar body mass to a bustard, such as jackals or caracals, or larger also capably kill adults. Additionally, warthogs (Phacochoerus spp.), mongoose and baboons (Papio ssp.) may eat eggs and small chicks. Chicks of up to largish size may be vulnerable to raptors such as tawny eagles (Aquila rapax), Verreaux's eagle-owls (Bubo lacteus) and Cape eagle-owls (Bubo capensis) as perhaps rarely an adult female. When alarmed, kori bustards make barking calls and bend forward and spread their tail and wings to appear larger. Adults will growl when their young are threatened by predators. Chicks tend to be the most vulnerable to predators by far. Many, despite their cryptic camouflage and the mother's defenses, are regularly picked off by jackals and leopards at night. Up to 82% of kori bustard chicks die in their first year of life. When found with carmine bee-eaters, the smaller birds may incidentally provide some protection from predators due to their vigilance. The display of the adult male may make it more conspicuous to larger predators, such as hyenas or lions. A shock display may be performed when a bird is alarmed. While too large to be prey for most predatory birds, it is known that the martial eagle is a serious natural enemy even for adult bustards, even adult males of at least twice their own weight. In one documented attack by a martial eagle on an adult kori bustard, both birds ended up wounded, the eagle with a bleeding leg from the bustard's counterattack, but the bustard more seriously injured, with a broken wing and several open wounds. Although it walked away, the injured bustard in the confrontation was found dead the next morning, being fed on by a jackal. At times, successful instances of martial eagle killing adult (presumed adult male) kori bustards have been photographed.


The kori bustard is generally a somewhat scarce bird. Appendix II of CITES and the 2000 Eskom Red Data Book for Birds lists the status of the nominate race as Vulnerable, estimating that in the next three generations, it is expected to decline by 10% in South Africa. In protected areas, they can be locally common. Viable populations exist in unprotected areas as well (e.g. Ethiopia and Sudan, and in Tanzania around Lake Natron and in the foothills of Mount Kilimanjaro) but in these areas, the birds are hunted. They have been much reduced by hunting, having been traditionally snared in Acacia gum baits and traps. Although no longer classified as game birds, they are still sometimes eaten. In Namibia, they are indicated as game as they are called the "Christmas turkey" and in South Africa, the "Kalahari Kentucky". Hunting of bustards is difficult to manage.
The kori bustard is now generally uncommon outside major protected areas. Habitat destruction is a major problem for the species, compounded by bush encroachment due to overgrazing by livestock and agricultural development. Poisons used to control locusts may also effect and collisions with overhead power wires regularly claim kori bustards. One 10 km (6.2 mi) stretch of overhead powerlines in the Karoo killed 22 kori bustards during a five-month period. Kori bustards tend to avoid areas used heavily by humans. Nonetheless, because it has such a large range and its rate of decline is thought to be relatively slow, the kori bustard is not currently listed in a threatened category on the IUCN Red List.
The species is prominent in many native African cultures, variously due to its imposing, impressive size, spectacular displays by adult males or the cryptic nature of the nesting female. The kori bustard features in dances and songs of the San people of Botswana, and paintings of these bustards feature in ancient San rock art. It was associated with royalty in Botswana since they reserved it for their own consumption, and since 2014 it is also the national bird of Botswana.


Ardisia crenata is a species of flowering plant in the primrose family, Primulaceae, that is native to East Asia. It is known by a variety of names such as Christmas berry, Australian holly, coral ardisia, coral bush, coralberry, coralberry tree,  hen's-eyes, and spiceberry. A. crenata is a compact shrub that reaches 1 metre (3.3 ft), often with a single stem.  Leaves are dark green, thick, glossy, and have tightly waved edges The flowers are small, white or reddish, fragrant, and form clusters. The fruit is a glossy, bright red drupe.  The seeds are able to germinate under a dense canopy and are dispersed by birds and humans.
This invasive species was introduced to the United States in the early twentieth century as an ornamental species.  It was observed to have escaped cultivation in 1982.
Preparations made from the root of Ardisia crenata are used as an ingredient in traditional Chinese medicine.


Mechanical control of Christmas berry is a challenge. Useful methods include hand-pulling in the case of small-scale invasions. This is not a very efficient method due to the difficulty of eliminating all the surrounding berries littering the ground that will soon replace the removed material. Another option is discing, which tills the soil up in hopes of destroying the rhizomes. This must be carefully administered to prevent harm of the surrounding local flora and ensuring that the rhizomes are subdued. Cutting as well as burning prove to be ineffective due to the strongly rhizomatous nature of the plant. If a mechanical method is used to control the plant, the site must be regularly monitored for at least a year in order to ensure elimination of Christmas berry.


The leaves of Ardisia crenata are being investigated as a remedy to stop asthmatic contractions and spasms. The plant contains a medicinal substance known as FR900359 that could treat several diseases, including asthma, hypertension, and uveal melanoma.


The ruddy turnstone (Arenaria interpres) is a small wading bird, one of two species of turnstone in the genus Arenaria.
It is now classified in the sandpiper family Scolopacidae but was formerly sometimes placed in the plover family Charadriidae. It is a highly migratory bird, breeding in northern parts of Eurasia and North America and flying south to winter on coastlines almost worldwide. It is the only species of turnstone in much of its range and is often known simply as turnstone.


The ruddy turnstone was formally described by the Swedish naturalist Carl Linnaeus in 1758 in the tenth edition of his Systema Naturae under the binomial name Tringa interpres. The species is now placed together with the black turnstone in the genus Arenaria that was introduced by the French zoologist Mathurin Jacques Brisson in 1760 with the ruddy turnstone as the type species. The genus name arenaria is from Latin arenarius, "inhabiting sand", from arena, "sand". The specific interpres means "messenger"; when visiting Gotland in 1741, Linnaeus thought that the Swedish word Tolk, meaning "interpreter" applied to this species, but in the local dialect the word means "legs" and is used for the redshank.
Two subspecies are recognised:
A. i. interpres (Linnaeus, 1758) – breeds in northeast Canada, Greenland, north Europe to northeast Siberia and west Alaska; winters in west, south Europe, Africa, south, east Asia, Australasia, Pacific islands, west USA and west Mexico
A. i. morinella (Linnaeus, 1766) – breeds in northeast Alaska and north Canada; winters in south USA to South America


It is a fairly small and stocky bird, 22–24 cm (8.7–9.4 in) long with a wingspan of 50–57 cm (20–22 in) and a weight of 85–150 g (3.0–5.3 oz). The dark, wedge-shaped bill is 2–2.5 cm (0.79–0.98 in) long and slightly upturned. The legs are fairly short at 3.5 cm (1.4 in) and are bright orange.
In all seasons, the plumage is dominated by a harlequin-like pattern of black and white. Breeding birds have reddish-brown upper parts with black markings. The head is mainly white with black streaks on the crown and a black pattern on the face. The breast is mainly black apart from a white patch on the sides. The rest of the underparts are white. In flight it reveals a white wingbar, white patch near the base of the wing and white lower back, rump and tail with dark bands on the uppertail-coverts and near the tip of the tail. The female is slightly duller than the male and has a browner head with more streaking.
Non-breeding adults are duller than breeding birds and have dark grey-brown upperparts with black mottling and a dark head with little white. Juvenile birds have a pale brown head and pale fringes to the upperpart feathers creating a scaly impression.
Birds of the subspecies morinella are smaller with darker upperparts and less streaking on the crown.
The ruddy turnstone has a staccato, rattling call and also a chattering alarm-call which is mainly given during the breeding season.


The ruddy turnstone breeds in northern latitudes, usually no more than a few kilometres from the sea. The subspecies A. i. morinella occurs in northern Alaska and in Arctic Canada as far east as Baffin Island. A. i. interpres breeds in western Alaska, Ellesmere Island, Greenland, Norway, Denmark, Sweden, Finland, Estonia and northern Russia. It formerly bred on the Baltic coast of Germany and has possibly bred in Scotland and the Faroe Islands.
In the Americas, the species winters on coastlines from Washington and Massachusetts southwards to the southern tip of South America although it is scarce in southern parts of Chile and Argentina and is only an unconfirmed vagrant in the Falkland Islands. In Europe, it winters in western regions from Iceland, Norway and Denmark southwards. Only small numbers are found on Mediterranean coasts. In Africa, it is common all the way down to South Africa with good numbers on many offshore islands. In Asia, it is widespread in the south with birds wintering as far north as southern China and Japan (mainly in the Ryukyu Islands). It occurs south to Tasmania and New Zealand and is present on many Pacific islands. Some non-breeding birds remain year round in many parts of the wintering range, with some of those birds still taking on breeding plumage in the spring and summer.


The ruddy turnstone has a varied diet including carrion, eggs, fish and plant material but it feeds mainly on invertebrates. Insects and their larvae are particularly important in the breeding season. At other times it also takes crustaceans, spiders, molluscs and worms. It often flips over stones and other objects to get at prey items hiding underneath; this behaviour is the origin of the name "turnstone". It usually forages in flocks.
They have also been observed preying on the eggs of other bird species such as gulls, terns, ducks, and even other turnstones, though this behaviour is uncommon. In the majority of observed cases, turnstones typically go after undefended or unattended nests, puncturing the shells with their beaks to get at the contents within.
Ruddy turnstones engage in a variety of behaviours to locate and capture prey. These behaviours can be placed into six general categories:
Routing: The turnstone manipulates piles of seaweed through flicking, bulldozing, and pecking to expose small crustaceans or gastropod molluscs hidden underneath.
Turning stones: As suggested by its name, the turnstone flicks stones with its bill to uncover hidden littorinids and gammarid amphipods.
Digging: With small flicks of its bill, the turnstone creates holes in the ground substrate (usually sand or mud) and then pecks at the exposed prey – often sandhoppers or seaweed flies.
Probing: The turnstone inserts its bill more than a quarter-length into the ground to get at littorinids and other gastropods.
Hammer–probing: The turnstone cracks open its prey's shell by using its bill as a hammer, and then extracts the animal inside through pecking and probing.
Surface pecking: The turnstone uses short, shallow pecks (less than a quarter bill-length) to get at prey at or just below the ground's surface.
There is evidence that turnstones vary between these feeding behaviours based on individual preference, sex, and even social status with respect to other turnstones. In one studied population, dominant individuals tended to engage in routing while preventing subordinates from doing the same. When these dominant individuals were temporarily removed, some of the subordinates started to rout, while others enacted no change in foraging strategy.


Ruddy turnstones can survive in a wide range of habitats and climatic conditions from Arctic to tropical. The typical breeding habitat is open tundra with water nearby. Outside the breeding season, it is found along coasts, particularly on rocky or stony shores. It is often found on man-made structures such as breakwaters and jetties. It may venture onto open grassy areas near the coast. Small numbers sometimes turn up on inland wetlands, especially during the spring and autumn migrations.
In terms of wintering sites, ruddy turnstones are particularly faithful to specific locations. A study published in 2009 examined turnstones wintering along a stretch of coastline in the Firth of Clyde. It found that 95% of birds resident to the area at the end of winter returned the following autumn. The same study also confirmed ruddy turnstones as one of the longest lived wader species, with annual adult mortality rates of under 15%. Their average lifespan is 9 years with 19 years and 2 months being the longest recorded.


It is a monogamous bird and pairs may remain together for more than one breeding season. The nest is a shallow scrape, often with a lining of leaves. It is about 11 cm (4.3 in) across and 3 cm (1.2 in) deep. It may be built amongst vegetation or on bare stony or rocky ground. Several pairs may nest close together.
A single clutch of two to five eggs is laid with four being most common. The eggs measure about 41 mm × 29 mm (1.6 in × 1.1 in) and weigh around 17.9 g (0.63 oz). They are smooth, slightly glossy and oval to pear-shaped. They are variable in colour but are commonly pale green-brown with dark brown markings, densest at the larger end. Incubation begins when the first egg is laid and lasts for about 22–24 days. The female is mainly responsible for incubating the eggs but the male may help towards the end.
The young birds are precocial and are able to leave the nest soon after hatching. They are buff above with dark grey markings and are white below. They are able to feed themselves but are protected by the parents, particularly the male. They fledge after 19–21 days.


According to the International Union for Conservation of Nature (IUCN), the ruddy turnstone population is currently very stable. Environment Canada surveys suggest that they have in fact decreased in abundance relative to the 1970s, and face a variety of threats during migration and winter. They estimate that the Canadian population is 100,000–500,000 adults. The Canadian Wildlife Service estimates that the worldwide population of ruddy turnstones is 449,000, and that 235,000 are breeding in North America while the rest are breeding throughout the Arctic regions. They are very common and widespread. Their remote breeding range and widespread winter range should help them remain a common species.


The black turnstone (Arenaria melanocephala) is a species of small wading bird. It is one of two species of turnstone in the genus Arenaria the ruddy turnstone (A. interpres) being the other. It is now classified in the sandpiper family, Scolopacidae, but was formerly sometimes placed in the plover family, Charadriidae. It is native to the west coast of North America and breeds only in Alaska.


It is 22–25 centimeters long and weighs 100–170 grams. The black bill is 20–27 millimeters long and slightly upturned. The legs and feet are blackish-brown with a reddish tinge. The bird is largely black and white in appearance. Breeding-plumaged adults have a black head and breast apart from a white spot between the eye and bill, a white stripe over the eye and white flecks on the sides of the breast. The upperparts are blackish-brown with pale fringes to the wing-coverts and scapular feathers. The belly and vent are white. In flight it shows a white wingbar, white shoulder patch and white tail with a broad black band across it. There is white from the lower back to the uppertail-coverts apart from a dark bar across the rump.
In winter the head and breast become largely dark brown with little white. Juveniles are similar to winter adults but browner with buff fringes to the wing-coverts and scapulars and a grey-brown tip to the tail.
The ruddy turnstone is similar but has rufous-brown markings on the upperparts and more white on the head and breast, particularly in breeding plumage. It has narrower wings and a narrower white wingbar. Its legs are orange and brighter than those of the black turnstone though there can be some overlap.
The black turnstone has a variety of calls, especially a rattling trill which can be heard throughout the year. This call is higher and less harsh than the similar call of the ruddy turnstone.  Other calls include a loud, screeching alarm call and a soft, purring call uttered to young chicks. Displaying males produce a long series of staccato notes in flight as well as chirruping trills on the ground.


Arenaria serpyllifolia, commonly known as thyme-leaf sandwort, or thyme-leaved sandwort is an annual or biennial  flowering plant in the pink and carnation family Caryophyllaceae.


Argemone albiflora, the white prickly poppy, also known as the bluestem prickly poppy or the Texas prickly poppy, is a small erect plant with a decorative white flower with a yellow latex. It is deeply rooted with yellow or red stamens. The plant is known for the sharp prickles on its stem and leaves. The sepals fall off as the flower of this plant grows bigger. It grows in the arid regions of the southern Midwest along roadsides and disturbed pieces of land. Native Americans have long revered this plant for its medicinal and other uses.


Argemone albiflora  can be found in the area ranging  from east Texas to northern Arkansas and southern Missouri. Native to areas of Southern California's High Desert such as Twentynine Palms.


White prickly poppy is an annual or biennial plant that often grows in colonies  in sandy or gravelly soils. This plant is often found along fences, roadsides, railroad tracks, on hills and slopes, and in overgrazed pastures.


Individuals of this species are erect, prickly deep-rooted plants with a white flower that contains many yellow and red stamens. The flowers grow solitarily or in loose cymes at the top of the plant.  It contains grayish-green, long (2 to 10 inches)  lanceolate to ovate leaves. The lower leaves are lobed to the midrib while the upper leaves are more shallow. The upper surface of the leaf is smooth although it has a few prickles along the midrib while the lower surface of the leaf is spiny along the midrib and main vein.


Flowers of Argemone albiflora  contain 3 long sepals and 6 petals and is ovate to almost circular with a rugged outer margin. The ovary is single chambered and contains many ovules with 3 to 5 lobed purple stigma. The fruit of this plant is a spiny capsule opening by the terminal splits. The seeds are brownish black.


Argemonine N-CH3
Allocryptopine Protopine Cryptopine Californidine - similar and others found in Papaveraceae


The oil of the white prickly poppy was used as a fine lubricant during WWII.  It was found that the oil content of the seed is 25.8% which is similar to the oil content found in soybeans.
The white prickly poppy is also used for decorative and ornamental purposes.


This plant is usually avoided by cattle and many other animals that roam the southern Midwest area as it is very prickly and has limited nutritional value. The seeds are the most nutritional part of the plant. Quails and doves consume these seeds for its high oil content. Each flower produces an abundance of seeds making it a reliable food source.


The white prickly poppy exudes a yellow latex that Native Americans use for many different ailments. Aztec priests would use the plant in sacrificial rituals. The Comanches held the plant in such high regard that they would make offerings to it during harvesting. They would take the latex and use it to remove warts, treat cold sores and other skin problems. A concoction could also be made from the flower to treat congestion from the cold or the flu. The seeds can also be used as an emetic to induce vomiting or as a laxative or even as a mild sedative.  The entire plant can also be used to treat bladder infection, prostate pain, or to help the throbbing pain of a migraine. A wash made from the tea can also be used to help heal sunburns or scraped skin. The plant is also smoked during some important ceremonies to induce a euphoric and mild sedating effect. However, if not used properly, the plant can be very toxic.


Argemone mexicana (Mexican poppy, Mexican prickly poppy, flowering thistle, cardo or cardosanto) is a species of poppy found in Mexico and now widely naturalized in many parts of the world. An extremely hardy pioneer plant, it is tolerant of drought and poor soil, often being the only cover on new road cuttings or verges. It has bright yellow latex. It is poisonous to grazing animals, and it is rarely eaten, but it has been used medicinally by many peoples, including those in its native area, as well as the Natives of the western US, parts of Mexico and many parts of India. In India, during the colorful festival Holika Dahan, adults and children worship by offering flowers, and this species is in its maximum flowering phase during March when the Holi festival is celebrated. It is also referred to as "kateli ka phool” in India.


Argemone mexicana seeds contain 22–36% of a pale yellow non-edible oil, called argemone oil or katkar oil, which contains the toxic alkaloids sanguinarine and dihydrosanguinarine.
Four quaternary isoquinoline alkaloids, dehydrocorydalmine, jatrorrhizine, columbamine, and oxyberberine, have been isolated from the whole plant of Argemone mexicana. Many other alkaloids such as argemexicaines A and B, coptisine, cryptopine, allocryptopine and chelerythrine have also been found in this plant.
The seed pods secrete a pale yellow latex when cut open. This argemone resin contains berberine and protopine.


The seeds resemble the seeds of Brassica nigra (mustard). As a result, mustard can be adulterated by argemone seeds, rendering it poisonous.  Several significant instances of katkar poisoning have been reported in India, Fiji, South Africa and other countries. The last major outbreak in India occurred in 1998. 1% adulteration of mustard oil by argemone oil has been shown to cause clinical disease. In India, Argemone oil is mixed with sunflower oil and sesame oil to increase the quantity, but this adulteration causes health disorders and renowned brands display "no argemone oil" to qualify purity.
Katkar oil poisoning causes epidemic dropsy, with symptoms including extreme swelling, particularly of the legs.


The Seri of Sonora, Mexico use the entire plant both fresh and dried.  An infusion is made to relieve kidney pain post-natally. When the Spanish arrived in Sonora, they called it cardosanto, taken as a laxative. An Argemone mexicana tea is used in Mali to treat malaria. In the traditional medicine of India, the yellow sap of A. mexicana and the whole plant may be used as a supposed treatment for jaundice.


Biodiesel production from A. mexicana seed oil using crystalline manganese carbonate has been demonstrated.


Argemone munita is a species of prickly poppy known by the common names flatbud prickly poppy and chicalote. "Munita" means "armed", in reference to the many long prickles. This flower is native to California, where it is widespread throughout the western part of the state and its eastern deserts, on slopes to 10,000 feet, and along roadsides. Its range also extends into Baja California, Arizona, and Nevada.
This poppy bears its flowers on erect, spiny stems which can exceed a meter in height. The sap is yellow. The blue-green or mint-green, lobed leaves are also very prickly. The white poppy blooms usually have six crinkly petals, each up to 4 centimeters long, and three sepals. The center of the flower is surrounded by a dense ring of up to 250 small yellow or orange stamens bearing the pollen. The flowers are often compared to sunny-side-up eggs, with the dark pistil in the center as a dot of pepper. The fruits are spiny white capsules 3 to 5 centimeters long, each containing many small seeds.  The leaves and seeds contain alkaloids which are probably toxic.


Argia alberta, the Paiute dancer, is a species of narrow-winged damselfly in the family Coenagrionidae. It is found in North America.
The IUCN conservation status of Argia alberta is "LC", least concern, with no immediate threat to the species' survival. The population is stable.


Argia bipunctulata, the seepage dancer, is a species of narrow-winged damselfly in the family Coenagrionidae. It is found in North America.
The IUCN conservation status of Argia bipunctulata is "LC", least concern, with no immediate threat to the species' survival. The population is stable.


Argia emma, or Emma's dancer, is a species of narrow-winged damselfly in the family Coenagrionidae. It is found in North America.
The IUCN conservation status of Argia emma is "LC", least concern, with no immediate threat to the species' survival. The population is stable.


The variable dancer (Argia fumipennis) is a damselfly of the family Coenagrionidae. It is native to North America, where it is widespread throughout the east and present in the interior western United States.
The male of the subspecies A. f. violacea (the violet dancer) is purple with a blue tip.


Argia fumipennis has three subspecies:
Black dancer A. f. atra
Smoky-winged dancer A. f. fumipennis
Violet dancer A. f. violacea


Argia immunda, the Kiowa dancer, is a species of narrow-winged damselfly in the family Coenagrionidae. It is found in Central America and North America.
The IUCN conservation status of Argia immunda is "LC", least concern, with no immediate threat to the species' survival. The population is stable.


The powdered dancer (Argia moesta) is a damselfly of the family Coenagrionidae. It is native to North America. It may be seen year-round in at least some of its range.


The common name refers directly to the male's pruinosity, appearing to be covered with a powdery blue or grayish substance. Older males are more pruinose, and may even be more ash white than blue. The scientific name moesta, means sorrowful, and may refer to customs (such as those on Ash Wednesday) of dusting oneself with ashes to express sorrow or mourning.


The Aztec dancer (Argia nahuana) damselfly is one of the pond damsels. The dark black stripe on the side of the thorax is forked from front to back. Other field marks include blue postocular spots, pale blue legs with a black stripe, and a blue ring on the seventh segment of the abdomen.


The springwater dancer (Argia plana) is a damselfly of the family Coenagrionidae.


The springwater dancer has a black stripe along the side of its thorax. The male is typically blue, but some can be violet.
The female is pale brown.