Agropyron cristatum is best adapted to dry rangeland conditions and is most frequently found in such circumstances. It prefers from 23 to 38 cm of precipitation per year, but can tolerate more moisture on favourable sites, extending its range into tundra and taiga conditions and elevations up to 2000 m above sea level in the southern portions of its adapted area. It prefers well drained, deep, loamy soils of medium and moderately coarse texture, including Chernozemic, Solonetzic, Regosolic, Brunisolic and Luvisolic soils. A. cristatum can tolerate salinity in the range of 5 to 15 mS/cm and prefers moderately alkaline conditions. It has low to medium fertility requirements. It will not tolerate prolonged flooding.
Agropyron cristatum is the most shade-tolerant of the crested wheatgrasses, but does best in open conditions. A. cristatum is extremely drought tolerant. It achieves this drought tolerance by starting growth very early in the season, then going dormant from seed set until fall when it will exhibit vegetative regrowth if moisture is sufficient.
A recent study has shown that invasive populations of Agropyron cristatum have spread across the upper U.S. as well as southern Canada, and the invading Agropyron cristatum populations have been found to have a higher granivory than native grasslands and they maintain dominance even when native grassland species are reintroduced. This current study indicated that the increased granivory of Agropyron cristatum did not contribute to its competitive success. The study did show that although A. cristatum was found to have higher granivory, after 2 years the difference between A. cristatum's granivory and that of native species lessens, and that there was no apparent preference among the animals for either wheat. Therefore, the factors responsible for Agropyron cristatum's high granivore content are still relatively unknown.
Agropyron cristatum is very tolerant of grazing, although under dry conditions new stands should be protected from grazing for at least two years as the seedling are slow to develop. A. cristatum can be damaged by several fungi, including leaf and stripe rusts, snow mold and some arthropods including black grass bugs (Labops sp.) in pure plantings.


Agropyron cristatum has been bred into many cultivars selected for attributes ranging from forage yield to drought tolerance to turf types that will exhibit more pronounced degrees of rhizomatous growth habit. It has been and continues to be, widely used in both agricultural and industrial reclamation activities.
Agropyron cristatum is known among other grasses and wheats for its relatively high granivory. Granivory describe the interaction between animals and seeds. Agropyron cristatum's high granivory indicates that animals feed on the seeds of the plant as their primary, or even exclusive, food source. Although this raises concerns about the plant's continued ability to reproduce if its seeds are all being consumed, the high granivory of this species does indicate that Agropyron cristatum is an important food source. Studies have been conducted in search of the cause of Agropyron cristatum's increased granivory, but as of yet a high relative granivory has not been proven to be a unique characteristic of A. cristatum, and could actually be attributed to factors other than the plant's genome, such as environmental conditions.
One promising factor that could lead to, and be responsible for, increased granivore in Agropyron cristatum is a certain genetic difference found on chromosome 6 of plants with a higher granivore content. Plants with a translocation on chromosome 6P yield wheat of greater weight and longer spike length than those without the mutation. Agropyron cristatum possesses higher tiller number, higher floret numbers, and greater resistance to various pathogens such as wheat rusts, powdery mildew, and barley yellow dwarf virus than many of its close wheat relatives. It has been used to cross-breed with other species of grass and wheat to transfer a greater disease resistance to them, as well as enhance their properties as a food source. This cross-breeding involves the transferring of the chromosome 6P translocation to the species it is cross-breeding with. Chromosome 6P of A. cristatum has also been proven to play an important role in regulating fertile tiller number and it possesses positive and negative regulators of tiller number. These regulators were specifically found to be on the 6PS and 6PL chromosome arms. High floret numbers and number of kernals per spike is controlled by genes located on chromosome 6P of Agropyron cristatum. Agropyron cristatum’s genes can be used to instill leaf resistance in other species of wheat. Three backcrosses between Agropyron cristatum and Aegilops tauschii produces a number stable, fertile lines of Aegilops tauschii that then has resistance to leaf rust.   Also, multi-spike cultivars of A. cristatum have been found to be more stable agronomically and achieve higher yields than cultivars with large-spike type.
It is an easy grass to establish by seed, having both high germination rates and high seedling vigour. It also establishes rapidly relative to many other grasses. Under non-irrigated conditions in low precipitation areas, Crested Wheatgrasses are consistently some of, if not the, highest yielding and persistent of domestic forage grasses. However, A. cristatum is lower yielding, although it is slightly more palatable, relative to other Crested Wheatgrasses.
Agropyron cristatum shows an immense toughness to a wide range of environmental conditions. Agropyron cristatum can be grown in cold temperatures, drought conditions, and relatively high amounts of salinity. It also has a resistance to barley yellow dwarf, wheat streak mosaic viruses, and leaf rust disease as well as containing high protein content.
Agropyron cristatum is a highly competitive and persistent plant in drier areas, and has a moderate ability to spread by seed. As such, its use in and adjacent to remaining natural grassland communities within its adapted areas in outside its native Eurasian distribution has come under criticism as a factor in natural grassland biodiversity loss, although the subject is still being studied. One such fear is that its seedlings' emergence does not decrease under herbicide treatment.


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The heart and dart (Agrotis exclamationis) is a moth of the family Noctuidae. The species was first described by Carl Linnaeus in his 1758 10th edition of Systema Naturae. A familiar moth to many, it is considered one of the most common of the European region. It occurs throughout the Palearctic realm from Ireland to Japan.
This is a quite variable species with forewings ranging from pale to dark brown but always recognizable by the distinctively shaped dark stigmata which give it its common name. The wingspan is 35–44 mm. The hindwings are whitish (compared with other common Agrotis species, the hindwings of this species are usually paler than in heart and club but darker than in turnip moth). This species usually has a dark area at the front of the thorax, visible as a horizontal bar when viewing the moth head on. The differences are not consistent however; they are highly variable in both colour and markings, and identification of atypical or worn examples may prove impossible without examination of genitalia. See Townsend et al.
This moth flies at night from May to July and is attracted to light, sometimes in large numbers. It also frequently visits nectar-rich flowers such as Buddleia, ragwort and red valerian.
The larva The caterpillars are coloured grey-brown, paler ventrally and have a pale dorsal line and clearly recognizable black point warts.
The pupa is  bright red. On the cremaster sit two short curved thorns.Larvae feed on a variety of plants, both wild and cultivated (see list below). This is one of the notorious cutworms and often severs or fatally damages plants at the base. The species overwinters as a full-grown larva in a chamber in the soil before pupating in the spring.
The flight season refers to the British Isles. This may vary in other parts of the range.


The bogong moth (Agrotis infusa) is a temperate species of night-flying moth, notable for its biannual long-distance seasonal migrations towards and from the Australian Alps, similar to the diurnal monarch butterfly. During the autumn and winter it is found in southern Queensland, western New South Wales, western Victoria, and also in South and Western Australia. Adult bogong moths breed and larvae hatch during this period, consuming winter pasture plants during their growth. During the spring, the moths migrate south or east and reside in mountains such as Mount Bogong, where they gregariously aestivate over the summer until their return towards breeding grounds again in the autumn.
The moth's name, bogong, is derived from an Australian Aboriginal language; the Dhudhuroa word bugung describes the brown colouration of the moth. It is an icon of Australian wildlife due to its historical role as an important food source and because Aboriginal peoples would come to where the moths spend the summer to feast on them and hold intertribal gatherings. In recent years, it has invaded major cities like Canberra, Melbourne, and Sydney due to strong winds during its spring migration.
Starting around 1980 and accelerating rapidly after 2016, the Bogong Moth population has sharply declined as a result of increasingly severe droughts, along with increased temperatures in caves used by the moths for aestivation, both primarily resulting from anthropogenic climate change. In December 2021 the bogong moth was added to the IUCN Red List as an Endangered Species.


Bogong moth populations are primarily located across southern Australia, west of the Great Dividing Range. The regions contain populations of nonmigratory and migratory moths of this species, distinguished by their differing seasonal presences in each region. The adult bogong moth lays eggs across New South Wales, southern Queensland, and northern parts of Victoria, where larvae hatch and grow until adulthood. During the spring season and subsequent summer aestivation, bogong moths migrate south or east towards the Australian Alps, and can be found in the Australian Capital Territory and Bogong Mountains. However, bogong moths can also be found in locations as far as Tasmania and New Zealand due to strong winds that blow them off their path.


Bogong moth eggs and larvae are primarily found in self-mulching soils (soil that mixes itself) and crop pastures, where both wild and agricultural larval food sources are abundant during the autumn and winter seasons. The heavy presence of larvae in these pastures can lead to significant crop damage. During the spring and summer seasons, when grasses overtake these pastures, conditions are unfavourable for larval survival since larvae do not consume these plants. This leads to delayed breeding, as bogong moths are multivoltine and so can raise multiple generations. Instead, adult bogong moths migrate in a southerly direction during the summer and aestivate (remain dormant), until conditions are favourable again.


During the spring migration, adult bogong moths can be found in their ideal aestivation sites, which generally consist of cool, dark caves and crevices but can include spaces underneath tors and even fallen tree trunks. Stable temperatures and humidity make these locations ideal for bogong moth aestivation. Crevices regulate their environment through wind flow, and caves generally have more regular temperatures and greater humidity. This reduces water loss in bogong moths during their inactivity. While temporary sites can be used in lower elevations, these sites undergo massive population fluctuations and movements throughout their use. Permanent aestivation sites are generally found in higher altitudes of 1500 metres (4920 ft) or above, with the largest, most stable aggregations found on the summits of mountains such as Mt. Gingera.


Adult bogong moths lay up to 2000 eggs in the soil or on plants near the soil after returning from aestivation sites in the autumn migration. Incubation times vary depending on temperature, with eggs hatching after a period of 4–7 days in laboratory conditions. The larvae of bogong moths undergo six instars. Caterpillars grow slowly throughout the first three instars, taking until June to develop over the winter. However, the larvae go through fast growth during the spring, reaching the final instar in late August to September, soon before migration. They are active at night, when they feed on plants within breeding grounds. Larvae undergo pupation in soil chambers at a depth of 20–150 mm (0.8-7.9 in). Pupation can last between 3–11 weeks depending on the temperature and environment. Pupae are 20 mm (0.8 in) in length and have a shiny, brown appearance. Adults emerge from the soil chambers and shortly after begin migration. Adult bogong moths are active at night, and have different seasonal behaviors. During the spring, bogong moths feed and migrate south, where they aestivate during the summer. Adult bogong moths are not sexually mature yet, and do not actively seek food during this period of dormancy. In autumn, the moths migrate again and return to their breeding grounds, laying eggs and dying.


Bogong moth larvae subsist on winter pasture crops and wild crop weeds such as cape weeds within bogong moth breeding grounds, primarily depending on annual dicotyledons that grow during the winter. Attacks on a wide variety of cultivated crops have been seen, with plants such as Medicago species, wheat, cabbages, cauliflowers, silver beet, peas, and potatoes all recorded being consumed by bogong moth caterpillars. However, the larvae avoid grasses, which overtake pastures during the summer, making summer unfavourable due to lack of larval food sources. Adult bogong moths feed on the nectar of flowers such as Epacris, Grevillea and Eucalyptus while breeding or migrating, but will not actively feed during aestivation.


During the spring migration, bogong moths gregariously aggregate with densities reaching 17,000 moths per square metre (10.8 square feet) within caves, crevices, and other areas hidden from the sunlight. The lack of light and relatively constant temperature and humidity makes these spots favourable during aestivation. The first moths that arrive occupy the deepest and darkest locations, using their fore tarsi to grip onto the rock faces, and aggregations form around these initial areas, with moths arriving later settling for less ideal areas with more sunlight, higher temperatures, and decreased humidity. To diminish the amount of light that reaches their light-sensitive eyes, later moths push themselves underneath the wings and abdomens of moths that arrived earlier and place their hind legs on top of the moths beneath them. This physical contact and aggregation allow the moths to retain body moisture. When the aestivating moths are disturbed, the moths within the area of disturbance briefly spread out and leave the aggregation, dropping excrement when unsettled before quickly returning to the aggregation and re-positioning themselves.
While the moths mostly remain dormant during aestivation, there are some periods of activity within the aggregation which are correlated to changes in light intensity.  During dawn and dusk, portions of the population become active, first crawling around and spreading out, and then flying out of their shelter into the open. While some water drinking has been observed, no evidence of copulation or active foraging has been found during these periods of activity.


The bogong moth suffers from predation during both its migration and aestivation. During the spring and autumn migrations, several species of birds, mammals, and even fish have been recorded preying on the moth. Little raven, currawong and Richard's pipit congregations form to feast on bogong moths as they travel from aestivation sites during migration. Aboriginal people also travelled to aestivation sites to feast on the dormant moths, and may have searched for these bird congregations to locate these sites. Bats also attack the moths during active flight periods during the dusk, and foxes, bush rats, and dusky antechinus have been recorded eating moths. Of the mammals that prey on the bogong moth, the endangered mountain pygmy-possum is the most reliant on bogong moths as a source of food.


Bogong moths were historically used as a food source by Aboriginal peoples located in South-eastern Australia. Groups would travel to the area towards the summits of mountains to harvest moths, where they also met with other Aboriginal peoples, fostering inter-tribal relations as people gathered and feasted during these harvests. They would go into the caves and scrape aestivating moths off the walls into nets and dishes using sticks. Once gathered, the moths would be roasted to remove the scales and wings and then either eaten immediately or ground into a paste and made into "moth meat" cakes that would last and could be taken home. The moth was said to have a nice nutty flavour that was most similar to walnuts or almonds. The bogong moth is an icon of Australian wildlife due to this historical role as an important food source and as an inter-tribal gathering point in Southeastern Australia.
An excavation of Cloggs Cave, near Buchan in Victoria, revealed microscopic remains of moth on a small grinding stone, estimated to be about 2,000 years old. This is the first confirmed evidence of insect food remains discovered on a stone artefact in the whole world. The Gunaikurnai people were one of the peoples who used to travel to the mountains to obtain the high-fat, energy-rich food, and stories of these travels had been passed down in their oral history. The excavation was done by researchers from Monash University in collaboration with the traditional owners, represented by the Gunaikurnai Land and Waters Corporation (GLaWAC).


Bogong moths have had a role in damaging crop plants since winter pastures serve as breeding grounds and larval food sources. Outbreaks of caterpillars have been recorded across New South Wales, with damage primarily done to heavy black soil flats. Medicago species, wheat, cabbages, cauliflowers, silver beet, peas, and potatoes have been recorded as being attacked by bogong moth larvae.


Millions of bogong moths have been blown off course from their spring migration into major cities like Canberra, Melbourne, and Sydney due to strong winds, with notable instances of Bogong Moth invasions including the 2000 Olympics in Sydney. The large amount of light and noise present can also attract moths towards cities. Parliament House in Canberra, present in the middle of the Bogong Moth flight path during migrations, was notably susceptible to moths. Light pollution from the building traps them during their flight and encourages the moth to find shelter within crevices, shadows, and sometimes even the insides of the buildings during the light and heat of the day. Efforts have been made to try and drive the moths out by turning off lights, covering attractive corners and pathways indoors, and using insecticides. However, insecticide use has ceased in urban areas due to concerns over environmental consequences.


Agrotis ipsilon, the dark sword-grass, black cutworm, greasy cutworm, floodplain cutworm or ipsilon dart, is a small noctuid moth found worldwide. The moth gets its scientific name from black markings on its forewings shaped like the letter "Y" or the Greek letter upsilon. The larvae are known as "cutworms" because they cut plants and other crops. The larvae are serious agricultural pests and feed on nearly all varieties of vegetables and many important grains.
This species is a seasonal migrant that travels north in the spring and south in the fall to escape extreme temperatures in the summer and winter. The migration patterns reflect how reproduction occurs in the spring and ceases in the fall.
Females release sex pheromones to attract males for mating. Pheromone production and release in females and pheromone responsiveness in males is dependent on the juvenile hormone (JH) and pheromone biosynthesis activating neuropeptide (BPAN). In the span of 2 months, the moth progresses through the life cycle stages egg, larvae, pupa, and adult. Throughout this time period, this moth faces the risk of predation and parasitism, such as by Hexamermis arvalis or by the parasite Archytas cirphis.


38–48 mm. Antennae in male bipectinated. Forewings brown, reddish-tinged, mixed with pale greyish-ochreous, costa and sometimes median area suffused with dark fuscous ; first and second lines edged with dark fuscous; spots outlined with black, reniform followed by a short black dash ; subterminal line obscure, usually preceded in middle by two fine black marks. Hindwings whitish-grey or whitish, terminally suffused with fuscous. Larva ochreous brownish or bronzy-grey, sides sometimes greenish ; dorsal, subdorsal, and spiracular lines faintly darker or lighter, usually darker-edged ; head sufFusedly brown-marked.


Larvae feed on weeds such as bluegrass, curled dock, lambsquarters, yellow rocket, and redroot pigweed. They will often eat all the weeds available before moving to attacking crops. Favored crops include most vegetable plants, alfalfa, clover, cotton, rice, sorghum, strawberry, sugarbeet, tobacco, and occasionally grains and grasses.


Adults feed on flower nectar. They are also attracted to deciduous trees and shrubs such as linden, wild plum, crabapple, and lilac. They are a pollinator of fetterbush lyonia.


In a given year, the number of generations differs based on location and weather conditions. In Canada, there are 1 or 2 generations, while in the United States, there are 2 to 4 per year. This species is abundant in warmer temperatures (such as Arkansas, US) during the late spring in May–June and early fall in September and October, while they are more abundant in cooler temperature (such as New York, US) during the summer in June and July. One life cycle lasts between 35–60 days.


A. ipsilon are seasonal migratory insects that travel south in the fall to escape harsh cold temperatures and travel north in the spring to escape extremely warm weather. Therefore, changes in thermoperiod as well as photoperiod may influence the onset of migration patterns in this species. Before migration southward in the fall, the reproductive system in both females and males shuts down to prevent copulation before winter. In the spring and early summer, though, before migration north, females release sex pheromones soon after eclosion. In one study, female moths collected from late April to early May were 100% mated.


Agrotis segetum, sometimes known as the turnip moth, is a moth of the family Noctuidae. The species was first described by Michael Denis and Ignaz Schiffermüller in 1775. It is a common European species and it is found in Africa and across Eurasia except for the northernmost parts.
It is a cutworm in the genus Agrotis, which possibly is the genus that includes the largest number of species of cutworms.


This is a very variable species with the fore-wings ranging from pale buff through to almost black. The paler forms have three dark-bordered stigmata on each fore-wing. Antennae of male bipectinated (comb like on both sides) with moderate length branches. The main feature distinguishing it from other Agrotis species is the shade of the hind-wings, pure white in the males and pearly grey in the females. The wingspan is 32–42 mm.
Edward Meyrick, an English schoolteacher who is especially notable for his study of the microlepidoptera, had this to say about the species:
Wingspan 33–41 mm. Antennae in male bipectinated. Fore-wings brownish, sometimes reddish tinged, with darker fuscous strigulae mixed with black scales, sometimes wholly darker suffused; sub-basal, first, and second lines edged with dark fuscous, second sometimes with dots only; spots outlined with black, orbicular and reniform centred with fuscous; subterminal line faint or whitish sprinkled, followed by darker suffusion. Hind-wings white, termen brownish. Larva pale grey or greyish ochreous, sometimes pinkish tinged; dorsal, subdorsal, and spiracular lines usually indicated by dark edges, subdorsal sometimes grey head pale, with two brown marks; plate of 2 more or less brown.
Agrotis segetum  is difficult to certainly distinguish from its congeners. See Townsend et al.
Agrotis clavis (Hufnagel, 1766)
Agrotis exclamationis (Linnaeus, 1758


As with any other noctuid, assorted diseases, predators, and parasitoids attack Agrotis segetum, mainly in the egg and larval stages.


This snake has a wide distribution in Asia, where it occurs in Bangladesh, Bhutan, Brunei, Burma, Cambodia, China, India, Indonesia, Laos, Malaysia, Philippines, Singapore, Thailand, and Vietnam.


Ailanthus altissima /eɪˈlænθəs ælˈtɪsɪmə/, commonly known as tree of heaven, ailanthus, varnish tree, or in Chinese as chouchun (Chinese: 臭椿; pinyin: chòuchūn), is a deciduous tree in the family Simaroubaceae. It is native to northeast and central China, and Taiwan. Unlike other members of the genus Ailanthus, it is found in temperate climates rather than the tropics.
The tree grows rapidly, and is capable of reaching heights of 15 metres (49 ft) in 25 years. While the species rarely lives more than 50 years, some specimens exceed 100 years of age. Its suckering ability allows this tree to clone itself indefinitely. It is considered a noxious weed and vigorous invasive species, and one of the worst invasive plant species in Europe and North America. In twenty-first century North America, the invasiveness of the species has been  compounded by its harboring of the also destructive and invasive spotted lanternfly.


A. altissima is a medium-sized tree that reaches heights between 17 and 27 m (56 and 89 ft) with a diameter at breast height of about 1 m (3 ft 3 in). The bark is smooth and light grey, often becoming somewhat rougher with light tan fissures as the tree ages. The twigs are stout, smooth to lightly pubescent, and reddish or chestnut in color. They have lenticels and heart-shaped leaf scars (i.e., a scar left on the twig after a leaf falls) with many bundle scars (i.e., small marks where the veins of the leaf once connected to the tree) around the edges. The buds are finely pubescent, dome-shaped, and partially hidden behind the petiole, though they are completely visible in the dormant season at the sinuses of the leaf scars. The branches are light to dark gray in color, smooth, lustrous, and contain raised lenticels that become fissures with age. The ends of the branches become pendulous. All parts of the plant have a distinguishing strong odor that is often likened to peanuts, cashews, or rotting cashews.
The leaves are large, odd- or even-pinnately compound on the stem. They range in size from 30 to 90 centimetres (1 to 3 ft) in length and contain 10–41 leaflets organised in pairs, with the largest leaves found on vigorous young sprouts. When they emerge in the spring, the leaves are bronze, then quickly turn from medium to dark green as they grow. The rachis is light to reddish-green with a swollen base. The leaflets are ovate-lanceolate with entire margins, somewhat asymmetric and occasionally not directly opposite to each other. Each leaflet is 5–18 cm (2–7 in) long and 2.5–5 cm (1–2 in) wide. They have a long, tapering end, while the bases have two to four teeth, each containing one or more glands at the tip. The leaflets' upper sides are dark green in color with light green veins, while the undersides are a more whitish green. The petioles are 5–12 millimetres (0.2–0.5 in) long. The lobed bases and glands distinguish it from similar sumac species.
The flowers are small and appear in large panicles up to 50 cm (20 in) in length at the end of new shoots. The individual flowers are yellowish green to reddish in color, each with five petals and sepals. The sepals are cup-shaped, lobed and united while the petals are valvate (i.e., they meet at the edges without overlapping), white and hairy towards the inside. They appear from mid-April in the south of its range to July in the north. A. altissima is dioecious, with male and female flowers being borne on different individuals. Male trees produce three to four times as many flowers as the females, making the male flowers more conspicuous. Furthermore, the male plants emit a foul-smelling odor while flowering to attract pollinating insects. Female flowers contain 10 (or rarely five through abortion) sterile stamens (stamenoids) with heart-shaped anthers. The pistil is made up of five free carpels (i.e., they are not fused), each containing a single ovule. Their styles are united and slender with star-shaped stigmata. The male flowers are similar in appearance, but they lack a pistil and the stamens do function, each being topped with a globular anther and a glandular green disc. The fruits grow in clusters; similar to the Ash (Fraxinus excelsior), the fruits ripen to a bright reddish-brown color in September. A fruit cluster may contain hundreds of seeds. The seeds borne on the female trees are 5 mm (0.2 in) in diameter and each is encapsulated in a samara that is 2.5 cm (1 in) long and 1 cm (0.4 in) broad, appearing July through August, but can persist on the tree until the next spring. The samara is large and twisted at the tips, making it spin as it falls, assisting wind dispersal, and aiding buoyancy for long-distance dispersal through hydrochory. Primary wind dispersal and secondary water dispersal are usually positively correlated in A. altissima, since most morphological characteristics of samaras affect both dispersal modes in the same way – except for the width of the samaras, which in contrast affects both types of dispersal in opposing ways, allowing differentiation in the dispersal strategies of this tree. The females can produce huge numbers of seeds, normally around 30,000 kilograms (66,000 lb) of tree, and fecundity can be estimated nondestructively through measurements of diameter at chest height.


In China, the tree of heaven has a long and rich history. It was mentioned in the oldest extant Chinese dictionary and listed in many Chinese medical texts for its purported curative ability. The roots, leaves, and bark are used in traditional Chinese medicine, primarily as an astringent. The tree has been grown extensively both in China and abroad as a host plant for the ailanthus silkmoth, a moth involved in silk production. Ailanthus has become a part of western culture, as well, with the tree serving as the central metaphor and subject matter of the best-selling American novel A Tree Grows in Brooklyn by Betty Smith.
The tree was first brought from China to Europe in the 1740s and to the United States in 1784. It was one of the first trees brought west during a time when chinoiserie was dominating European arts and was initially hailed as a beautiful garden specimen. However, enthusiasm soon waned after gardeners became familiar with its suckering habits and its foul odor. Despite this, it was used extensively as a street tree during much of the 19th century. Outside Europe and the United States, the plant has been spread to many other areas beyond its native range, and is considered internationally as a noxious weed. In many countries, it is an invasive species due to its ability both to colonize disturbed areas quickly and to suppress competition with allelopathic chemicals. The tree also resprouts vigorously when cut, making its eradication difficult and time-consuming. This has led to the tree being called "tree of hell" among gardeners and conservationists.


The earliest introductions of A. altissima to countries outside of its native range were to the southern areas of Korea and to Japan. The tree may be native to these areas, but the tree is generally agreed to be a very early introduction. Within China, it has also been naturalized beyond its native range in areas such as Qinghai, Ningxia, and Xinjiang.
In 1784, not long after Jussieu had sent seeds to England, some were forwarded to the United States by William Hamilton, a gardener in Philadelphia. In both Europe and America, it quickly became a favoured ornamental, especially as a street tree, and by 1840, it was available in most nurseries. The tree was separately brought to California in the 1890s by Chinese immigrants who came during the California Gold Rush. It has escaped cultivation in all areas where it was introduced, but most extensively in the United States. It has naturalized across much of Europe, including Germany, Austria, Switzerland, the Czech Republic, the Pannonian region (i.e. southeastern Central Europe around the Danube River basin from Austria, Slovakia, and Hungary south to the Balkan ranges) and most countries of the Mediterranean Basin. In Montenegro and Albania A. altissima is widespread in both rural and urban areas, and while in the first it was introduced as an ornamental plant, it very soon invaded native ecosystems with disastrous results and became an invasive species. Ailanthus has also been introduced to Argentina, Australia (where it is a declared weed in New South Wales and Victoria), New Zealand (where it is listed under the National Pest Plant Accord and is classed an "unwanted organism"), the Middle East, and in some countries in South Asia such as Pakistan. In South Africa, it is listed as an invasive species that must be controlled, or removed and destroyed.
In North America, A. altissima is present from Massachusetts in the east, west to southern Ontario, southwest to Iowa, south to Texas, and east to the north of Florida. In the west, it is found from New Mexico west to California and north to Washington. In the east of its range, it grows most extensively in disturbed areas of cities, where it was long ago present as a planted street tree. It also grows along roads and railways. For example, a 2003 study in North Carolina found the tree of heaven was present on 1.7% of all highway and railroad edges in the state, and had been expanding its range at the rate of 4.76% counties per year. Similarly, another study conducted in southwestern Virginia determined that the tree of heaven is thriving along roughly 30% of the state's interstate highway system length or mileage. It sometimes enters undisturbed areas, as well, and competes with native plants. In western North America, it is most common in mountainous areas around old dwellings and abandoned mining operations. It is classified as a noxious or invasive plant on National Forest System lands and in many states because its prolific seed production, high germination rate, and capacity to regrow from roots and root fragments enable A. altissima to out-compete native species. For this reason, control measures on public lands and private property are advised where A. altissima has naturalized.


Tree of heaven is an opportunistic plant that thrives in full sun and disturbed areas. It spreads aggressively both by seeds and vegetatively by root sprouts, re-sprouting rapidly after being cut. It is considered a shade-intolerant tree and cannot compete in low-light situations, though it is sometimes found competing with hardwoods, but such competition rather indicates it was present at the time the stand was established. On the other hand, a study in an old-growth hemlock-hardwood forest in New York found that Ailanthus was capable of competing successfully with native trees in canopy gaps where only 2 to 15% of full sun was available. The same study characterised the tree as using a "gap-obligate" strategy to reach the forest canopy, meaning it grows rapidly during a very short period rather than growing slowly over a long period. It is a short lived tree in any location and rarely lives more than 50 years. Ailanthus is among the most pollution-tolerant of tree species, including sulfur dioxide, which it absorbs in its leaves. It can withstand cement dust and fumes from coal tar operations, as well as resist ozone exposure relatively well. Furthermore, high concentrations of mercury have been found built up in tissues of the plant.
Ailanthus has been used to re-vegetate areas where acid mine drainage has occurred and it has been shown to tolerate pH levels as low as 4.1 (approximately that of tomato juice). It can withstand very low phosphorus levels and high salinity levels. The drought-tolerance of the tree is strong due to its ability to effectively store water in its root system. It is frequently found in areas where few trees can survive. The roots are also aggressive enough to cause damage to subterranean sewers and pipes. Along highways, it often forms dense thickets in which few other tree species are present, largely due to the toxins it produces to prevent competition. The roots are poisonous to people.
Ailanthus produces an allelopathic chemical called ailanthone, which inhibits the growth of other plants. The inhibitors are strongest in the bark and roots, but are also present in the leaves, wood and seeds of the plant. One study showed that a crude extract of the root bark inhibited 50% of a sample of garden cress (Lepidium sativum) seeds from germinating. The same study tested the extract as an herbicide on garden cress, redroot pigweed (Amaranthus retroflexus), velvetleaf (Abutilon theophrasti), yellow bristlegrass (Setaria pumila), barnyard grass (Echinochloa crusgalli), pea (Pisum sativum cv. Sugar Snap) and maize (Zea mays cv. Silver Queen). It proved able to kill nearly 100% of seedlings with the exception of velvetleaf, which showed some resistance. Another experiment showed a water extract of the chemical was either lethal or highly damaging to 11 North American hardwoods and 34 conifers, with the white ash (Fraxinus americana) being the only plant not adversely affected. The chemical does not, however, affect the tree of heaven's own seedlings, indicating that A. altissima has a defence mechanism to prevent autotoxicity. Resistance in various plant species has been shown to increase with exposure. Populations without prior exposure to the chemicals are most susceptible to them. Seeds produced from exposed plants have also been shown to be more resistant than their unexposed counterparts.
The tree of heaven is a very rapidly growing tree, possibly the fastest growing tree in North America. Growth of one to two metres (3 ft 3 in to 6 ft 7 in) per year for the first four years is considered normal. Shade considerably hampers growth rates. Older trees, while growing much slower, still do so faster than other trees. Studies found that Californian trees grew faster than their East Coast counterparts, and American trees in general grew faster than Chinese ones.
In northern Europe the tree of heaven was not considered naturalised in cities until after the Second World War. This has been attributed to the tree's ability to colonize areas of rubble of destroyed buildings where most other plants would not grow. In addition, the warmer microclimate in cities offers a more suitable habitat than the surrounding rural areas; it is thought that the tree requires a mean annual temperature of 8 °C (46 °F) to grow well, which limits its spread in more northern and higher altitude areas. For example, one study in Germany found the tree of heaven growing in 92% of densely populated areas of Berlin, 25% of its suburbs and only 3% of areas outside the city altogether. In other areas of Europe this is not the case as climates are mild enough for the tree to flourish. It has colonized natural areas in Hungary, for example, and is considered a threat to biodiversity at that country's Aggtelek National Park.
Several species of Lepidoptera use the leaves of Ailanthus as food, including the Indian moon moth (Actias selene) and the common grass yellow (Eurema hecabe). In North America the tree is the host plant for the ailanthus webworm (Atteva aurea), though this ermine moth is native to Central and South America and originally used other members of the mostly tropical Simaroubaceae as its hosts. In the US, it has been found to host the brown marmorated stink bug and the Asiatic shot-hole borer. The spotted lanternfly (L. delicatula), relies on the metabolites of A. altissima for the completion of its life cycle and the pervasiveness of A. altissima is seen as a driving factor in L. delicatula's invasive spread outside of China. In its native range A. altissima is associated with at least 32 species of arthropods and 13 species of fungi.
In North America, the leaves of ailanthus are sometimes attacked by Aculops ailanthii, a mite in the family Eriophyidae. Leaves infested by the mite begin to curl and become glossy, reducing their ability to function. Therefore, this species has been proposed as a possible biocontrol for ailanthus in the Americas. Research from September 2020 indicates a verticillium wilt, caused by Verticillium nonalfalfae, may function as a biological control for A. altissima, with the weevil Eucryptorrhynchus brandti serving as a vector.
Due to the tree of heaven's weedy habit, landowners and other organisations often resort to various methods of control to keep its populations in check. For example, the city of Basel in Switzerland has an eradication program for the tree. It can be very difficult to eradicate, however. Means of eradication can be physical, thermal, managerial, biological or chemical. A combination of several of these can be most effective, though they must of course be compatible. All have some positive and negative aspects, but the most effective regimen is generally a mixture of chemical and physical control. It involves the application of foliar or basal herbicides to kill existing trees, while either hand pulling or mowing seedlings to prevent new growth.


Tree of heaven is a popular ornamental tree in China and valued for its tolerance of difficult growing conditions. It was once very popular in cultivation in both Europe and North America, but this popularity dropped, especially in the United States, due to the disagreeable odor of its blossoms and the weediness of its habit. The problem of odor was previously avoided by only selling pistillate plants since only males produce the smell, but a higher seed production also results. Michael Dirr, a noted American horticulturalist and professor at the University of Georgia, reported meeting, in 1982, a grower who could not find any buyers. He further writes (his emphasis):
For most landscaping conditions, it has no value as there are too many trees of superior quality; for impossible conditions this tree has a place; selection could be made for good habit, strong wood and better foliage which would make the tree more satisfactory; I once talked with an architect who tried to buy Ailanthus for use along polluted highways but could not find an adequate supply [...]
— Michael A. Dirr, Manual of Woody Landscape Plants
In Europe, however, the tree is still used in the garden to some degree as its habit is generally not as invasive as it is in America. In the United Kingdom it is especially common in London squares, streets, and parks, though it is also frequently found in gardens of southern England and East Anglia. It becomes rare in the north, occurring only infrequently in southern Scotland. It is also rare in Ireland. In Germany the tree is commonly planted in gardens. The tree has furthermore become unpopular in cultivation in the west because it is short-lived and that the trunk soon becomes hollow, making trees more than two feet in diameter unstable in high winds.
A few cultivars exist, but they are not often sold outside of China and probably not at all in North America:
'Hongye' – The name is Chinese and means "red leaves". As the name implies it has attractive vivid red foliage
'Thousand Leaders'
'Metro' – A male cultivar with a tighter crown than usual and a less weedy habit
'Erythrocarpa' – The fruits are a striking red
'Pendulifolia' – Leaves are much longer and hang elegantly


Nearly every part of A. altissima has had various uses in Chinese traditional medicine, although there is no high-quality clinical evidence that it has an effect on any disease.
A tincture of the root bark was thought useful by American herbalists in the 19th century. It contains phytochemicals, such as quassin and saponin, and ailanthone. The plant may be mildly toxic. The noxious odours have been associated with nausea and headaches, and with contact dermatitis reported in both humans and sheep, which developed weakness and paralysis. It contains a quinone irritant, 2,6-dimethoxybenzoquinone, as well as quassinoids.


The 1943 book A Tree Grows in Brooklyn by Betty Smith uses the tree of heaven as its central metaphor, using it as an analogy for the ability to thrive in a difficult environment. At that time as well as now, ailanthus was common in neglected urban areas. She writes:
There's a tree that grows in Brooklyn. Some people call it the Tree of Heaven. No matter where its seed falls, it makes a tree which struggles to reach the sky. It grows in boarded up lots and out of neglected rubbish heaps. It grows up out of cellar gratings. It is the only tree that grows out of cement. It grows lushly...survives without sun, water, and seemingly earth. It would be considered beautiful except that there are too many of it.
— A Tree Grows in Brooklyn, Introduction
In William Faulkner's novel, Sanctuary, a "heaven-tree" stands outside the Jefferson jail, where Lee Goodwin and a "negro murderer" are incarcerated. The tree is associated with the black prisoner's despair in the face of his impending execution and the spirituals that he sings in chorus with other black people who keep a sort of vigil in the street below:
...they sang spirituals while white people slowed and stopped in the leafed darkness that was almost summer, to listen to those who were sure to die and him who was already dead singing about heaven and being tired; or perhaps in the interval between songs a rich, sourceless voice coming out of the high darkness where the ragged shadow of the heaven-tree which snooded the street lamp at the corner fretted and mourned: "Fo days mo! Den dey ghy stroy de bes ba'yton singer in nawth Mississippi!"
Upon the barred and slitted wall the splotched shadow of the heaven-tree shuddered and pulsed monstrously in scarce any wind; rich and sad, the singing fell behind.
In the 2013 book Teardown: Memoir of a Vanishing City by journalist Gordon Young, the tree is referenced in a description of the Carriage Town neighborhood in Flint, Michigan.
Festive Victorian-era homes in various stages of restoration battled for supremacy with boarded-up firetraps and overgrown lots landscaped with weeds, garbage, and “ghetto palms,” a particularly hardy invasive species known more formally as Ailanthus altissima, or the tree of heaven, perhaps because only God can kill the things. Around the corner, business was brisk at a drug house where residents and customers alike weren't above casually taking a piss in the driveway.
Ailanthus is also sometimes counter-nicknamed "tree from hell" due to its prolific invasiveness and the difficulty in eradicating it. In certain parts of the United States, the species has been nicknamed the "ghetto palm" because of its propensity for growing in the inhospitable conditions of urban areas, or on abandoned and poorly maintained properties, such as in war-torn Afghanistan.
Until 26 March 2008, a 60-foot (18 m)-tall member of the species was a prominent "centerpiece" of the sculpture garden at the Noguchi Museum in the Astoria section in the borough of Queens in New York City. The tree had been spared by the sculptor Isamu Noguchi when in 1975 he bought the building which would become the museum and cleaned up its back lot. The tree was the only one he left in the yard, and the staff would eat lunch with Noguchi under it. "[I]n a sense, the sculpture garden was designed around the tree", said a former aide to Noguchi, Bonnie Rychlak, who later became the museum curator. By 2008, the old tree was found to be dying and in danger of crashing into the building, which was about to undergo a major renovation. The museum hired the Detroit Tree of Heaven Woodshop, an artists' collective, to use the wood to create benches, sculptures and other amenities in and around the building. The tree's rings were counted, revealing its age to be 75, and museum officials hoped it would regenerate from a sucker.


The rufous-crowned sparrow (Aimophila ruficeps) is a small American sparrow. This passerine is primarily found across the Southwestern United States and much of the interior of Mexico, south to the transverse mountain range, and to the Pacific coast to the southwest of the transverse range. Its distribution is patchy, with populations often being isolated from each other. Twelve subspecies are generally recognized, though up to eighteen have been suggested. This bird has a brown back with darker streaks and gray underparts. The crown is rufous, and the face and supercilium are gray with a brown or rufous streak extending from each eye and a thick black malar streak.
These sparrows feed primarily on seeds in the winter and insects in the spring and summer. The birds are often territorial, with males guarding their territory through song and displays. Flight is awkward for this species, which prefers to hop along the ground for locomotion. They are monogamous and breed during spring. Two to five eggs are laid in the bird's nest, which is cup-shaped and well hidden. Adult sparrows are preyed upon by house cats and small raptors, while young may be taken by a range of mammals and reptiles. They have been known to live for up to three years, two months. Although the species has been classified as least concern, or unthreatened with extinction, some subspecies are threatened by habitat destruction and one may be extinct.


This bird belongs to the family Passerellidae, which consists of the American sparrows. The American sparrows are seed-eating New World birds with conical bills, brown or gray plumage, and distinctive head patterns. Birds in the genus Aimophila tend to be medium-sized at 5 to 8 inches (13 to 20 cm) in length, live in arid scrubland, have long bills and tails in proportion to their body size as well as short, rounded wings, and build cup-shaped nests.
The rufous-crowned sparrow was described in 1852 by American ornithologist John Cassin as Ammodramus ruficeps. It has also been described as belonging to the genus Peucaea, which contains several sparrows in the genus Aimophila that share characteristics, such as a larger bill and a patch of yellow under the bend of the wing, that other members of the genus do not. However, splitting the Peucaea sparrows into a separate genus is not generally recognized. A 2008 phylogenetic analysis of the genus Aimophila divided it into four genera, with the rufous-crowned sparrow and its two closest relatives, the Oaxaca sparrow and rusty sparrow, being maintained as the genus Aimophila. In addition, this study suggested that the rufous-crowned sparrow may be more closely related to the brown towhees of the genus Pipilo than the other members of the historical genus Aimophila.
The derivation of the current genus name, Aimophila, is from the Greek aimos/ἀιμος, meaning "thicket", and -philos/-φιλος, meaning "loving". The specific epithet is a literal derivation of the common name, derived from the Latin rufus, meaning "reddish" or "tawny", and -ceps, from caput, meaning "head". The bird is also occasionally referred to colloquially as the rock sparrow because of its preference for rocky slopes.


Twelve subspecies are generally recognized, although sometimes up to eighteen are named.
A. r. ruficeps, the nominate subspecies, was described by Cassin in 1852. It is found in the coastal ranges of California and on the western slopes of the Sierra Nevada. This subspecies is darker and noticeably smaller than A. r. eremoeca and has distinct rufous-brown streaking on its upperparts.
A. r. canescens was described by American ornithologist W. E. Clyde Todd in 1922, and it is found in southwestern California and northeast Baja California as far east as the base of the San Pedro Mártir. While the species itself is listed as of least concern, this subspecies is listed as a "species of special concern" by the California Department of Fish and Wildlife, signifying that this population is threatened with extinction. It appears to be extremely similar to A. r. ruficeps but is darker.
A. r. obscura, described by Donald R. Dickey and Adriaan van Rossem in 1923, is found in the Channel Islands of California on Santa Cruz, Anacapa, and formerly on Santa Catalina. While the Santa Catalina population has not been observed since 1863, the subspecies seems to have colonized Anacapa Island. No records exist of them before 1940. This subspecies is similar to A. r. canescens but is darker.
A. r. sanctorum was described by van Rossem in 1947. It was found on the Todos Santos Islands off the coast of northwest Baja California. This subspecies is believed to be extinct. This is the darkest of the coastal subspecies, especially on its underbelly.
A. r. sororia was described by Robert Ridgway in 1898, and is found in the mountains of southern Baja California, specifically the Sierra de la Laguna. It is the palest of the coastal subspecies.
A. r. scottii, described by George Sennett in 1888, is found from northern Arizona to New Mexico south to northeastern Sonora and northwestern Coahuila. It appears to be a darker gray than A. r. eremoeca and has narrower and darker rufous streaks on its breast.
A. r. rupicola was described by van Rossem in 1946. It is found in the mountains of southwestern Arizona. It is similar in appearance to A. r. scottii but is darker and grayer on its back.
A. r. simulans was described by van Rossem in 1934, and it is found in northwestern Mexico from southeastern Sonora and southwestern Chihuahua to Nayarit and northern Jalisco. It has more rufous coloration on its back and is paler on its underbelly than A. r. scottii.
A. r. eremoeca was described by N. C. Brown in 1882. It is found from southeastern Colorado to New Mexico, Texas, northern Chihuahua, and central Coahuila. It has grayish upperparts and a dark breast.
A. r. fusca, described by Edward William Nelson in 1897, is found in western Mexico from southern Nayarit to southwestern Jalisco, northern Colima, and Michoacan. It is darker and more rufous on its upperparts than A. r. australis. It also possesses a darker rufous crown which does not show a gray stripe down the middle.
A. r. boucardi was described by Philip Sclater in 1867, and it is found in eastern Mexico from southern Coahuila to San Luis Potosí, northern Puebla, and southern Oaxaca. This subspecies is darker than A. r. eremoeca and has dull brown, not rufous, streaking on the chest.
A. r. australis, described by Edward William Nelson in 1897, occurs in southern Mexico from Guerrero to southern Puebla and Oaxaca. A. r. scottii is similar in appearance, but this subspecies is smaller and has a shorter bill.
The other six subspecies that are occasionally recognized are A. r. extima and A. r. pallidissima, which were described by A. R. Phillips in 1966, A. r. phillipsi, which was described by J.P. Hubbard and Crossin in 1974, and A. r. duponti, A. r. laybournae, and A. r. suttoni, which were described by J.P. Hubbard in 1975.


The rufous-crowned sparrow is a smallish sparrow at 5.25 inches (13.3 cm) in length, with males tending to be larger than females. It ranges from 15 to 23 g (0.53 to 0.81 oz) in weight and averages about 19 g (0.67 oz). It has a brown back with darker streaks and gray underparts. Its wings are short, rounded, and brown and lack wingbars, or a line of feathers of a contrasting color in the middle of the bird's wing. The sparrow's tail is long, brown, and rounded. The face and supercilium (the area above the eye) are gray with a brown or rufous streak extending from each eye and a thick black streak on each cheek. The crown ranges from rufous to chestnut, a feature which gives it its common name, and some subspecies have a gray streak running through the center of the crown. The bill is yellow and cone-shaped. The sparrow's throat is white with a dark stripe. Its legs and feet are pink-gray. Both sexes are similar in appearance, but the juvenile rufous-crowned sparrow has a brown crown and numerous streaks on its breast and flanks during the spring and autumn.
The song is a short, fast, bubbling series of chip notes that can accelerate near the end, and the calls include a nasal chur and a thin tsi. When threatened or separated from its mate, the sparrow makes a dear-dear-dear call.


This bird is found in the southwestern United States and Mexico from sea level up to 9,800 feet (3,000 m), though it tends to be found between 3,000 and 6,000 feet (910 and 1,830 m). It lives in California, southern Arizona, southern New Mexico, Texas, and central Oklahoma south along Baja California and in western Mexico to southern Puebla and Oaxaca. In the midwestern United States, the sparrow is found as far east as a small part of western Arkansas, and also in a small region of northeastern Kansas, its most northeastern habitat. The range of this species is discontinuous and is made up of many small, isolated populations. The rufous-crowned sparrow is a non-migratory species, though the mountain subspecies are known to descend to lower elevations during severe winters. Male sparrows maintain and defend their territories throughout the year.
This sparrow is found in open oak woodlands and dry uplands with grassy vegetation and bushes. It is often found near rocky outcroppings. The species is also known from coastal scrublands and chaparral areas. The rufous-crowned sparrow thrives in open areas cleared by burning.


The average territory size of rufous-crowned sparrows in the chaparral of California ranges from 2 acres (0.81 ha) to 4 acres (1.6 ha). The density of territories varies by habitat, including 2.5 to 5.8 territories per 99 acres (40 ha) of three- to five-year-old burned chaparral to 3.9 to 6.9 territories for the same amount of coastal scrubland. One pair tends to be supported by a territory, although birds without a mate have been seen sharing a territory with a mated pair.
This sparrow is awkward in flight and primarily uses running and hopping to move. The rufous-crowned sparrow will at times forage in pairs during the breeding season, and in family-sized flocks in late summer and early autumn. During the winter they can occasionally be found in loose mixed-species foraging flocks.
Predators of adult sparrows include house cats and small raptors like Cooper's and sharp-shinned hawks, American kestrels, and white-tailed kites. The nests may be raided by a range of species including mammals and reptiles such as snakes, though nest predation has not yet been directly observed, and nesting sparrows have been observed using three kinds of displays to distract potential predators; the rodent run, the broken wing, and the tumbling off the bush. Birds adopt a rodent run display to distract predators. The head, neck and tail are lowered, wings held out, and feathers fluffed as the bird runs rapidly and voices a continuous alarm call. In the broken wing display, the sparrow imitates having a broken wing by dropping one to the ground and hopping away from the nest with one wing dragging, leading the predator away until the bird ceases the act and escapes the predator. The adult rufous-crowned sparrow distracts a nest predator by falling from the top of a bush to attract the predator to itself in the tumbling off the bush display.
The longest lifespan recorded for a rufous-crowned sparrow is three years, two months. Two species of tick, Amblyomma americanum and Ixodes pacificus, are known to parasitize the sparrow.


This sparrow feeds primarily on small grass and forb seeds, fresh grass stems, and tender plant shoots during autumn and winter. During these seasons, insects such as ants, grasshoppers, ground beetles, and scale insects as well as spiders make up a small part of its diet. In the spring and summer, the bird's diet includes a greater quantity and variety of insects.
The rufous-crowned sparrow forages slowly on or near the ground by walking or hopping under shrubs or dense grasses. Though it occasionally forages in weedy areas, it is almost never observed foraging in the open. It has occasionally been observed feeding in branches and low shrubs. During the breeding season, it gleans its food from grasses and low shrubs. However, normally the species obtains its food by either pecking or less frequently scratching at leaf litter. This bird tends to forage in a small family group and in a limited area.
It is unknown whether this species obtains all of the water it needs from its food or if it must also drink; however, it has been observed both drinking and bathing in pools of water after rain storms.


The rufous-crowned sparrow breeds in sparsely vegetated scrubland. Males attract a mate by singing from regular positions at the edge of their territories throughout the breeding season. These birds are monogamous, taking only one mate at a time, and pairs often remain together for several years. If singing males come within contact of each other, they may initially raise their crowns and face the ground to display this feature; if that fails to make the other bird leave, they stiffen their body, droop their wings, raise their tails, and stick their head straight out. Males guard their territories year-round.
While it is not known when precisely the breeding season starts, the earliest that a sparrow has been observed carrying nesting material was on March 2 in southern California. The female bird builds a bulky, thick-walled open-cup nest typically on the ground, though occasionally in a low bush up to 18 in (46 cm) above it, from dried grasses and rootlets, sometimes with strips of bark, small twigs, and weed stems. Nests are well hidden, as they are built near bushes or tall grasses or overhanging rock with concealing vegetation. Once a sparrow chooses a nesting site, it tends to return to the site for many years. It lays between two and five eggs at a time and typically only raises one brood a year, though some birds in California have been observed raising two or even three broods a year. In case of a nesting failure, replacement clutches may be laid. The eggs are an unmarked, pale bluish-white. Broods of the rufous-crowned sparrow have very occasionally been observed to be parasitized by the brown-headed cowbird.
Incubation of the eggs lasts 11 to 13 days and is performed solely by the female. The hatchlings are naked and quills do not begin to show until the third day. Only females brood the nestlings, though both parents may bring whole insects to their young. When a young rufous-crowned sparrow leaves the nest after eight or nine days, it is still incapable of flight, though it can run through the underbrush; during this time it is still fed by the parents. Juveniles tend to leave their parent's territory and move into adjacent habitat in autumn or early winter. Reproductive success varies strongly with annual rainfall and is highest in wet El Niño years, since cool rainy weather reduces the activity of snakes, the main predator of the sparrow's nests.


The rufous-crowned sparrow is treated as a species of least concern, or not threatened with extinction, by BirdLife International due to its large geographical range of about 463,323 sq mi (1,200,000 km²), estimated population of 2.4 million individuals, and lack of a 30% population decline over the last ten years. In years without sufficient rains, many birds fail to breed and those that do produce fewer offspring. Some of the local populations of this bird are threatened and declining in number. The island subspecies and populations have declined in some cases: A. r. sanctorum of the Todos Santos Islands is believed to be extinct, and the populations on Santa Catalina Island and Baja California's Islas de San Martin have not been observed since the early 1900s. Populations of the species in southern California are also becoming more restricted in range because of urbanization and agricultural development in the region. Additionally, the sparrow is known to have been poisoned by the rodenticide warfarin, though more research is needed to determine the effects of pesticides on the rufous-crowned sparrow.


The species was once widespread in East Asia, but large-scale exports and the destruction of its forest habitat have reduced populations in eastern Russia and in China to below 1,000 pairs in each country; Japan, however, is thought to still hold some 5,000 pairs. The Asian populations are migratory, overwintering in lowland eastern China and southern Japan.
Specimens frequently escape from collections, and in the 20th century, a large, feral population was established in Great Britain; more recently, small numbers have bred in Ireland, concentrated in the parks of Dublin. Now, about 7,000 are in Britain with other populations on the European continent, the largest of which is in the region of Berlin. Isolated populations exist in the United States. The town of Black Mountain, North Carolina, has a limited population, and a free-flying feral population of several hundred mandarins exist in Sonoma County, California. This population is the result of several ducks escaping from captivity, then reproducing in the wild. In 2018, a single bird, dubbed Mandarin Patinkin, was seen in New York City's Central Park.
The habitats it prefers in its breeding range are the dense, shrubby forested edges of rivers and lakes. It mostly occurs in low-lying areas, but it may breed in valleys at altitudes of up to 1,500 m (4,900 ft). In winter, it additionally occurs in marshes, flooded fields, and open rivers. While it prefers fresh water, it may also be seen wintering in coastal lagoons and estuaries. In its introduced European range, it lives in more open habitat than in its native range, around the edges of lakes, water meadows, and cultivated areas with woods nearby.


In the wild, mandarin ducks breed in densely wooded areas near shallow lakes, marshes or ponds. They nest in cavities in trees close to water and during the spring. A single clutch of nine to twelve eggs is laid in April or May. Although the male may defend the brooding female and his eggs during incubation, he himself does not incubate the eggs and leaves before they hatch. Shortly after the ducklings hatch, their mother flies to the ground and coaxes the ducklings to leap from the nest. After all of the ducklings are out of the tree, they will follow their mother to a nearby body of water.


Mandarins feed by dabbling or walking on land. They mainly eat plants and seeds, especially beech mast. The species will also add snails, insects and small fish to its diet. The diet of mandarin ducks changes seasonally; in the fall and winter, they mostly eat acorns and grains. In the spring, they mostly eat insects, snails, fish and aquatic plants. In the summer, they eat dew worms, small fish, frogs, mollusks, and small snakes. They feed mainly near dawn or dusk, perching in trees or on the ground during the day.


Predation of the mandarin duck varies between different parts of its range. Mink, raccoon dogs, otters, polecats, Eurasian eagle-owls, and grass snakes are all predators of the mandarin duck. The greatest threat to the mandarin duck is habitat loss due to loggers. Hunters are also a threat to the mandarin duck, because often they are unable to recognize the mandarin in flight and as a result, many are shot by accident. Mandarin ducks are not hunted for food, but are still poached because their extreme beauty is prized.


For Koreans, mandarin ducks represent peace, fidelity, and plentiful offspring. Similar to the Chinese, they believe that these ducks mate for life. For these reasons, pairs of wooden-carved mandarin ducks called wedding ducks are often given as wedding gifts and play a significant role in Korean marriage.


Their breeding habitat is wooded swamps, shallow lakes, marshes, ponds and creeks in the eastern United States, the west coast of the United States, some adjacent parts of southern Canada, and the west coast of Mexico. They usually nest in cavities in trees close to water, although they will take advantage of nesting boxes in wetland locations. Other species may compete with them for nesting cavities, such as birds of prey, as well as mammals such as grey squirrels, and these animals may also occupy nest boxes meant for wood ducks. Wood ducks may end up nesting up to a mile away from their water source as a result.  Females line their nests with feathers and other soft materials, and the elevation provides some protection from predators. Unlike most other ducks, the wood duck has sharp claws for perching in trees and can, in southern regions, produce two broods in a single season—the only North American duck that can do so.
Females typically lay between 7 and 15 white-tan eggs that incubate for an average of 30 days. However, if nesting boxes are placed too close together, females may lay eggs in the nests of their neighbours, which may lead to nests which may contain as many as 30 eggs and unsuccessful incubation, a behaviour known as "nest dumping".
After hatching, the precocial ducklings climb to the opening of the nest cavity, jump down from the nest tree and make their way to water. The mother calls them to her, but does not help them in any way. They prefer nesting over water so the young have a soft landing, but will nest up to 140 m (460 ft) away from the shoreline. The day after they hatch, the young climb to the nest entrance and jump to the ground. The ducklings can swim and find their own food by this time.
Wood ducks feed by dabbling (feeding from the surface rather than diving underwater) or grazing on land. They mainly eat berries, acorns, and seeds, but also insects, making them omnivores.


The birds are year-round residents in parts of its southern range, but the northern populations migrate south for the winter. They overwinter in the southern United States near the Atlantic Coast. 75% of the wood ducks in the Pacific Flyway are non-migratory. Due to their attractive plumage, they are also popular in waterfowl collections and as such are frequently recorded in Great Britain as escapees—populations have become temporarily established in Surrey in the past, but are not considered to be self-sustaining in the fashion of the closely related mandarin duck. Along with the mandarin duck, the wood duck is considered an invasive species in England and Wales, and it is illegal to release them into the wild. Given its native distribution, the species is also a potential natural vagrant to Western Europe and there have been records in areas such as Cornwall, Scotland and the Isles of Scilly, which some observers consider may relate to wild birds; however, given the wood duck's popularity in captivity, it would be extremely difficult to prove their provenance. There is a small feral population in Dublin.


The population of the wood duck was in serious decline in the late 19th century as a result of severe habitat loss and market hunting both for meat and plumage for the ladies' hat market in Europe. By the beginning of the 20th century, wood ducks had virtually disappeared from much of their former range. In response to the Migratory Bird Treaty established in 1916 and enactment of the U.S. Migratory Bird Treaty Act of 1918, wood duck populations began to recover slowly. By ending unregulated hunting and taking measures to protect remaining habitat, wood duck populations began to rebound in the 1920s. The development of the artificial nesting box in the 1930s gave an additional boost to wood duck production. More information on the efficacy of nest boxes can be found in the Conservation Evidence webpage.
Landowners as well as park and refuge managers can encourage wood ducks by building wood duck nest boxes near lakes, ponds, and streams. Fulda, Minnesota has adopted the wood duck as an unofficial mascot, and a large number of nest boxes can be found in the area.
Expanding North American beaver populations throughout the wood duck's range have also helped the population rebound as beavers create an ideal forested wetland habitat for wood ducks.
The population of the wood duck has increased a great deal in the last several years. The increase has been due to the work of many people constructing wood duck boxes and conserving vital habitat for the wood ducks to breed. During the open waterfowl season, U.S. hunters have only been allowed to take two wood ducks per day in the Atlantic and Mississippi Flyways. However, for the 2008–2009 season, the limit was raised to three. The wood duck limit remains at two in the Central Flyway and at seven in the Pacific Flyway. It is the second most commonly hunted duck in North America, after the mallard.


The Eurasian skylark (Alauda arvensis) is a passerine bird in the lark family, Alaudidae. It is a widespread species found across Europe and the Palearctic with introduced populations in New Zealand, Australia and on the Hawaiian Islands. It is a bird of open farmland and heath, known for the song of the male, which is delivered in hovering flight from heights of 50 to 100 metres (160 to 330 ft). The sexes are alike. It is streaked greyish-brown above and on the breast and has a buff-white belly.
The female Eurasian skylark builds an open nest in a shallow depression on open ground well away from trees, bushes and hedges. She lays three to five eggs which she incubates for around 11 days. The chicks are fed by both parents but leave the nest after eight to ten days, well before they can fly. They scatter and hide in the vegetation but continue to be fed by the parents until they can fly at 18 to 20 days of age. Nests are subject to high predation rates by larger birds and small mammals. The parents can have several broods in a single season.


The Eurasian skylark was described by the Swedish naturalist Carl Linnaeus in 1758 in the tenth edition of his Systema Naturae and retains its original binomial name of Alauda arvensis. It is one of the four species placed in the genus Alauda. The genus name is from the Latin alauda, "lark". Pliny thought the word was originally of Celtic origin. The specific arvensis is also Latin, and means "of the field". The results of a molecular phylogenetic study of the lark family Alaudidae published in 2013 suggested that Eurasian skylark is most closely related to the Oriental skylark Alauda gulgula.
Formerly, many authorities considered the Japanese skylark as a separate species. It is now usually considered a subspecies of the Eurasian skylark. Alternate names for the Eurasian skylark include northern skylark and sky lark.


The Eurasian skylark is 18–19 cm (7.1–7.5 in) in length. Like most other larks, the Eurasian skylark is a rather dull-looking species, being mainly brown above and paler below.  It has a short blunt crest on the head, which can be raised and lowered. In flight it shows a short tail and short broad wings. The tail and the rear edge of the wings are edged with white, which are visible when the bird is flying away, but not if it is heading towards the observer. The male has broader wings than the female. This adaptation for more efficient hovering flight may have evolved because of female Eurasian skylarks' preference for males that sing and hover for longer periods and so demonstrate that they are likely to have good overall fitness.
It is known for the song of the male, which is delivered in hovering flight from heights of 50 to 100 m, when the singing bird may appear as just a dot in the sky from the ground. The long, unbroken song is a clear, bubbling warble delivered high in the air while the bird is rising, circling or hovering. The song generally lasts two to three minutes, but it tends to last longer later in the mating season, when songs can last for 20 minutes or more. At wind farm sites, male skylarks have been found to sing at higher frequencies as a result of wind turbine noise.


In the 19th century multiple batches of Eurasian skylarks were released in New Zealand beginning in 1864 in Nelson (in the South Island) and in 1867 in Auckland (in the North Island). The wild population increased rapidly and had spread throughout both the North and South Islands by the 1920s.
In Australia the Eurasian skylark was introduced on multiple occasions beginning in 1850. It is now widespread in the southeast of the continent. In New South Wales it mostly occurs south of 33°S. It is widespread throughout Victoria and Tasmania and also occurs in the south-eastern corner of South Australia around Adelaide.
The Eurasian skylark was introduced to the southeastern Hawaiian Islands beginning in 1865. Although once common, it has declined in abundance on Oahu and is no longer found on Kauai. A study published in 1986 found European skylarks remained only on the islands of Hawaii and Maui and estimated a total population of 10,000 individuals.
The Eurasian skylark was introduced to Vancouver Island off the west coast of Canada in 1903; additional birds were introduced in 1913. The population grew and by 1962 there were around 1000 individuals. The numbers have subsequently declined due to loss of habitat, and in 2007 there were estimated to be only around 100 individuals spread over four small areas of the Saanich Peninsula.


Eurasian skylarks first breed when they are one year of age. Nesting may start in late March or early April. The nest is probably built by the female alone and is a shallow depression in the ground lined with grasses. The clutch is 3 to 5 eggs. The eggs of the nominate subspecies average 23.4 mm × 16.8 mm (0.92 in × 0.66 in) in size and weigh around 3.35 g (0.118 oz). They have a grey-white or greenish background and are covered in brown or olive spots. They are incubated only by the female beginning after the last egg is laid and hatch synchronously after 11 days. The altricial young are cared for by both parents and for the first week are fed almost exclusively on insects. The nestlings fledge after 18 to 20 days but they usually leave the nest after 8 to 10 days. They are independent of their parents after around 25 days. The parents can have up to 4 broods in a season.


In the UK, Eurasian skylark numbers have declined over the last 30 years, as determined by the Common Bird Census started in the early 1960s by the British Trust for Ornithology.  There are now only 10% of the numbers that were present 30 years ago.  The RSPB have shown that this large decline is mainly due to changes in farming practices and only partly due to pesticides.  In the past cereals were planted in the spring, grown through the summer and harvested in the early autumn.  Cereals are now planted in the autumn, grown through the winter and are harvested in the early summer.  The winter grown fields are much too dense in summer for the Eurasian skylark to be able to walk and run between the wheat stems to find its food.
A list of conservation interventions that could impact Alauda arvensis can be found in the Conservation Evidence website. English farmers are now encouraged and paid to maintain and create biodiversity for improving the habitat for Eurasian skylarks.  Natural England's Environmental Stewardship Scheme offers 5 and 10-year grants for various beneficial options.  For example, there is an option where the farmer can opt to grow a spring cereal instead of a winter one, and leave the stubble untreated with pesticide over the winter. The British Trust for Ornithology likens the stubbles to "giant bird tables" – providing spilt grain and weed seed to foraging birds.
The RSPB's research, over a six-year period, of winter-planted wheat fields has shown that suitable nesting areas for Eurasian skylarks can be made by turning the seeding machine off (or lifting the drill) for a 5 to 10 metres stretch as the tractor goes over the ground to briefly stop the seeds being sown.  This is repeated in several areas within the same field to make about two skylark plots per hectare.  Subsequent spraying and fertilising can be continuous over the entire field.  DEFRA suggests that Eurasian skylark plots should not be nearer than 24 m to the perimeter of the field, should not be near to telegraph poles, and should not be enclosed by trees.
When the crop grows, the Eurasian skylark plots (areas without crop seeds) become areas of low vegetation where Eurasian skylarks can easily hunt insects, and can build their well camouflaged ground nests.  These areas of low vegetation are just right for skylarks, but the wheat in the rest of the field becomes too closely packed and too tall for the bird to seek food.  At the RSPB's research farm in Cambridgeshire skylark numbers have increased threefold (from 10 to 30 pairs) over six years.  Fields where Eurasian skylarks were seen the year before (or nearby) would be obvious good sites for skylark plots.  Farmers have reported that skylark plots are easy to make and the RSPB hope that this simple effective technique can be copied nationwide.


Albizia julibrissin is known by a wide variety of common names, such as Persian silk tree and pink siris. It is also called Lankaran acacia or bastard tamarind, though it is not too closely related to either genus. The species is called Chinese silk tree, silk tree or mimosa in the United States, which is misleading—the former can refer to any species of Albizia which is most common in any one locale; and, although once included in Mimosa, neither is it very close to the Mimoseae. To add to the confusion, several species of Acacia, notably Acacia baileyana and Acacia dealbata, are also known as "mimosa" (especially in floristry), and many Fabaceae trees with highly divided leaves are called thus in horticulture.
Its leaves slowly close during the night and during periods of rain, the leaflets bowing downward; thus its modern Persian name shabkhosb (شب‌خسب) means "night sleeper". This tendency also explains the Chinese common name hehuan, which means "shut happy" and symbolizes a happy couple in bed. In Japan its common names are nemunoki, nemurinoki and nenenoki which all mean "sleeping tree". Nemu tree is a partial translation of nemunoki.


Original habitats of the tree include regions from Iran (Persia) and the Republic of Azerbaijan to China and Korea.
A. julibrissin is widely planted as an ornamental plant in parks and gardens, grown for its fine leaf texture, flowers and attractive horizontal canopy. Other positive attributes are a fast growth rate, low water requirements and the ability to thrive planted in full sun in hot summer climates.  It is frequently planted in semi-arid areas like California's Central Valley, central Texas and Oklahoma. Although capable of surviving drought, growth will be stunted and the tree tends to look sickly. As such it should be given infrequent, deep waterings during the summer, which will benefit growth and flowering.
The broad crown of a mature tree makes it useful for providing dappled shade. The flower colour varies from white in A. julibrissin f. alba, to rich red-tipped flowers. Variants with cream or pale yellow flowers are also reported. Other cultivars are becoming available: 'Summer Chocolate' has red foliage ageing to dark bronze, with pale pink flowers; 'Ishii Weeping' (or 'Pendula') has a drooping growth habit.


The common kingfisher (Alcedo atthis), also known as the Eurasian kingfisher and river kingfisher, is a small kingfisher with seven subspecies recognized within its wide distribution across Eurasia and North Africa. It is resident in much of its range, but migrates from areas where rivers freeze in winter.
This sparrow-sized bird has the typical short-tailed, large-headed kingfisher profile; it has blue upperparts, orange underparts and a long bill. It feeds mainly on fish, caught by diving, and has special visual adaptations to enable it to see prey under water. The glossy white eggs are laid in a nest at the end of a burrow in a riverbank.


This species has the typical short-tailed, dumpy-bodied, large-headed, and long-billed kingfisher shape. The adult male of the western European subspecies, A. a. ispida has green-blue upperparts with pale azure-blue back and rump, a rufous patch by the bill base, and a rufous ear-patch. It has a green-blue neck stripe, white neck blaze and throat, rufous underparts, and a black bill with some red at the base. The legs and feet are bright red. It is about 16 centimetres (6+1⁄2 in) long with a wingspan of 25 cm (10 in), and weighs 34–46 grams (1+3⁄16–1+5⁄8 oz). The female is identical in appearance to the male except that her lower mandible is orange-red with a black tip. The juvenile is similar to the adult, but with duller and greener upperparts and paler underparts. Its bill is black, and the legs are also initially black. Feathers are moulted gradually between July and November with the main flight feathers taking 90–100 days to moult and regrow. Some that moult late may suspend their moult during cold winter weather.
The flight of the kingfisher is fast, direct and usually low over water. The short, rounded wings whirr rapidly, and a bird flying away shows an electric-blue "flash" down its back.
In North Africa, Europe and Asia north of the Himalayas, this is the only small blue kingfisher. In south and southeast Asia, it can be confused with six other small blue-and-rufous kingfishers, but the rufous ear patches distinguish it from all but juvenile blue-eared kingfishers; details of the head pattern may be necessary to differentiate the two species where both occur.
The common kingfisher has no song. The flight call is a short, sharp whistle chee repeated two or three times. Anxious birds emit a harsh, shrit-it-it and nestlings call for food with a churring noise.


The common kingfisher is widely distributed over Europe, Asia, and North Africa, mainly south of 60°N. It is a common breeding species over much of its vast Eurasian range, but in North Africa it is mainly a winter visitor, although it is a scarce breeding resident in coastal Morocco and Tunisia. In temperate regions, this kingfisher inhabits clear, slow-flowing streams and rivers, and lakes with well-vegetated banks. It frequents scrubs and bushes with overhanging branches close to shallow open water in which it hunts. In winter it is more coastal, often feeding in estuaries or harbours and along rocky seashores. Tropical populations are found by slow-flowing rivers, in mangrove creeks and in swamps.
Common kingfishers are important members of ecosystems and good indicators of freshwater community health. The highest densities of breeding birds are found in habitats with clear water, which permits optimal prey visibility, and trees or shrubs on the banks. These habitats have also the highest quality of water, so the presence of this bird confirms the standard of the water. Measures to improve water flow can disrupt this habitat, and in particular, the replacement of natural banks by artificial confinement greatly reduces the populations of fish, amphibians and aquatic reptiles, and waterside birds are lost. It can tolerate a certain degree of urbanisation, provided the water remains clean.
This species is resident in areas where the climate is mild year-round, but must migrate after breeding from regions with prolonged freezing conditions in winter. Most birds winter within the southern parts of the breeding range, but smaller numbers cross the Mediterranean into Africa or travel over the mountains of Malaysia into Southeast Asia. Kingfishers migrate mainly at night, and some Siberian breeders must travel at least 3,000 km (1,900 mi) between the breeding sites and the wintering areas.


Like all kingfishers, the common kingfisher is highly territorial; since it must eat around 60% of its body weight each day, it is essential to have control of a suitable stretch of river. It is solitary for most of the year, roosting alone in heavy cover. If another kingfisher enters its territory, both birds display from perches, and fights may occur, in which a bird will grab the other's beak and try to hold it underwater. Pairs form in the autumn but each bird retains a separate territory, generally at least 1 km (1⁄2 mi) long, but up to 3.5 km (2+1⁄4 mi) and territories are not merged until the spring.
The courtship is initiated by the male chasing the female while calling continually, and later by ritual feeding, with copulation usually following.
The nest is in a burrow excavated by both birds of the pair in a low vertical riverbank, or sometimes a quarry or other cutting. The straight, gently inclining burrow is normally 60–90 cm (25–35 in) long and ends in an enlarged chamber. The nest cavity is unlined but soon accumulates a litter of fish remains and cast pellets.
The common kingfisher typically lays two to ten glossy white eggs, which average 1.9 cm (3⁄4 in) in breadth, 2.2 cm (7⁄8 in) in length, and weigh about 4.3 g (5⁄32 oz), of which 5% is shell. One or two eggs in most clutches fail to hatch because the parent cannot cover them. Both sexes incubate by day, but only the female incubates at night. An incubating bird sits trance-like, facing the tunnel; it invariably casts a pellet, breaking it up with the bill. The eggs hatch in 19–20 days, and the altricial young are in the nest for a further 24–25 days, often more. Once large enough, young birds will come to the burrow entrance to be fed. Two broods, sometimes three, may be reared in a season.


The common kingfisher hunts from a perch 1–2 m (3–7 ft) above the water, on a branch, post or riverbank, bill pointing down as it searches for prey. It bobs its head when food is detected to gauge the distance and plunges steeply down to seize its prey usually no deeper than 25 cm (10 in) below the surface. The wings are opened underwater and the open eyes are protected by the transparent third eyelid. The bird rises beak-first from the surface and flies back to its perch. At the perch the fish is adjusted until it is held near its tail and beaten against the perch several times. Once dead, the fish is positioned lengthways and swallowed head-first. A few times each day, a small greyish pellet of fish bones and other indigestible remains is regurgitated.
The food is mainly fish up to 12.5 cm (5 in) long, but the average size is 2.3 cm (7⁄8 in). In Central Europe, 97% of the diet was found to be composed of fish ranging in size from 2 to 10 cm with an average of 6.5 cm (body mass range from <0.1 g to >10 g, average 3 g). Minnows, sticklebacks, small roach and trout are typical prey. About 60% of food items are fish, but this kingfisher also catches aquatic insects such as dragonfly larvae and water beetles, and, in winter, crustaceans including freshwater shrimps. In Central Europe, however, fish represented 99.9% of the diet (data from rivers, streams, and reservoirs from years 1999 to 2013). Common kingfishers have also been observed to catch lamprey. One study found that food provisioning rate increased with brood size, from 1498 g (505 fishes for four nestlings) to 2968 g (894 fishes for eight nestlings). During the fledging period each chick consumed on average 334 g of fish, which resulted in an estimated daily food intake of 37% of the chick's body mass (average over the entire nestling period). The average daily energy intake was 73.5 kJ per chick (i.e., 1837 kJ per 25 days of the fledging period).
A challenge for any diving bird is the change in refraction between air and water. The eyes of many birds have two foveae (the fovea is the area of the retina with the greatest density of light receptors), and a kingfisher can switch from the main central fovea to the auxiliary fovea when it enters water; a retinal streak of high receptor density which connects the two foveae allows the image to swing temporally as the bird drops onto the prey. The egg-shaped lens of the eye points towards the auxiliary fovea, enabling the bird to maintain visual acuity underwater. Because of the positions of the foveae, the kingfisher has monocular vision in air, and binocular vision in water. The underwater vision is not as a sharp as in air, but the ability to judge the distance of moving prey is more important than the sharpness of the image.
Each cone cell of a bird's retina contains an oil droplet that may contain carotenoid pigments. These droplets enhance color vision and reduce glare. Aquatic kingfishers have high numbers of red pigments in their oil droplets; the reason red droplets predominate is not understood, but the droplets may help with the glare or the dispersion of light from particulate matter in the water.