The USDA Forestry Service lists Plumeria rubra as a poisonous plant and warns against touching or eating any part of the plant.
In Cambodia, as with other Plumeria species, P. rubra flowers is used for necklaces, as offerings to deities or as decoration for coffins. The leaves of this species are used in the care of sores and made into soothing infusions.
The flowers and bark of it are also used in traditional Chinese medicine in the treatment of fever, bacillary dysentery, pertussis and so on.
P. rubra possesses fulvoplumierin, an antibiotic that inhibits the growth of Mycobacterium tuberculosis. The plant has also been shown to be an antifungal, antiviral, analgesic, antispasmodic, and hypoglycemic. P. rubra is also reported to contain agoniadin, plumierid, plumeric acid, cerotinic acid, and lupeol, and the stem possesses an alkaloid called triterpinoid. The plant has been known to promote digestion and excretion, along with respiratory and immune functions. The sap of the plant is used as a laxative and is a remedy for bloating and stomachaches. The bark is said to be purgative and is also used for venereal sores. The flowers can be boiled in water or juice and made into a salad to promote bowel movement, urine flow, and to control gas and phlegm. The flowers are also used in the treatment of asthma.
On the Molokai island in the Hawaiian archipelago, P. rubra is cultivated in order to produce neck garlands (leis). They are also used to make a scented oil in many Pacific islands that include Hawaii. The flowers are used to scent coconut oil. The bark contains faint purplish streaks and the wood is hard and compact with a very fine texture. The wood takes a high polish.
Plumeria rubra is the national flower of Nicaragua, where it is known under the local name "sacuanjoche". It is also the village flower of Asan-Maina on Guam.


Plusiodonta compressipalpis, the moonseed moth, is a moth of the family Erebidae. The species was first described by Achille Guenée in 1852. It is found in North America from Minnesota, extreme southern Canada and Connecticut, south to northern Florida and Texas,tambien en el sur de españa.
The wingspan is 25–33 mm. Adults are on wing from May to September. There are two generations over much of east.
The larvae feed on Menispermum species and snailseed vines.


The European golden plover (Pluvialis apricaria), also known as the European golden-plover, Eurasian golden plover, or just the golden plover within Europe, is a largish plover. This species is similar to two other golden plovers: the American golden plover, Pluvialis dominica, and Pacific golden plover, Pluvialis fulva, which are both smaller, slimmer and relatively longer-legged than European golden plover, and both have grey rather than white axillary feathers (only properly visible in flight).


The European golden plover was formally described by the Swedish naturalist Carl Linnaeus in 1758 in the tenth edition of his Systema Naturae. He placed it with the other plovers in the genus Charadrius and coined the binomial name Charadrius apricarius. The species is now placed in the genus Pluvialis that was introduced in 1760 by the French zoologist  Mathurin Jacques Brisson. The genus name is Latin and means "relating to rain", from pluvia, "rain".  It was believed that golden plovers flocked when rain was imminent.  The species name  apricaria is Latin and means "to bask in the sun". The European golden plover is monotypic: no subspecies are recognised.


The European golden plover is quite thickset, with its wings only being slightly longer than its tail. Its most distinct feature is a white "s"-shaped band stretching from its forehead to its flanks.


The European golden plover tends to breed in the Arctic tundra and other palearctic areas, ranging as far west as Iceland, where they are called Heiðlóa, and as far east as central Siberia. It tends to gather in large flocks and winter in open areas, agricultural plains, ploughed land, and short meadows, ranging from Europe to North Africa.
In the United Kingdom, golden plover chicks rely on Tipulidae for feeding, while in Sweden Bibionidae are more important.


The European golden plover's call is a monosyllabic, slightly descending, melancholic "tuu".
Its flight action is rapid and powerful, with regular wingbeats.


The European golden plover spends summers in Iceland, and in Icelandic folklore, the appearance of the first plover in the country means that spring has arrived. The Icelandic media always covers the first plover sighting, which in 2017, took place on March 27. In 2020, the first golden plover was sighted on March 16.


The European golden plover is one of the species to which the Agreement on the Conservation of African-Eurasian Migratory Waterbirds (AEWA) applies.


The American golden plover (Pluvialis dominica), is a medium-sized plover. The genus name is Latin and means relating to rain, from pluvia, "rain".  It was believed that golden plovers flocked when rain was imminent.  The species name dominica refers to Santo Domingo, now Hispaniola, in the West Indies.


Measurements:
Length: 24–28 cm (9+1⁄2–11 in)
Weight: 122–194 g (4+5⁄16–6+13⁄16 oz)
Wingspan: 65–67 cm (25+1⁄2–26+1⁄2 in)
The breeding adult American golden plover has a black face, neck, breast, and belly, with a white crown and nape that extends to the side of the breast. The back is mottled black and white with pale, gold spots. The breeding female is similar, but with less black. When in winter plumage, both sexes have grey-brown upperparts, pale grey-brown underparts, and a whitish eyebrow. The head is small, along with the bill.
It is similar to two other golden plovers, European and Pacific. The American golden plover is smaller, slimmer and relatively longer-legged than European golden plover (Pluvialis apricaria) which also has white axillary (armpit) feathers. It is more similar to Pacific golden plover (Pluvialis fulva) with which it was once considered conspecific under the name "lesser golden plover". The Pacific golden plover is slimmer than the American species, has a shorter primary projection, and longer legs, and is usually yellower on the back.


The breeding habitat of American golden plover is Arctic tundra from northern Canada and Alaska. They nest on the ground in a dry open area. They are migratory and winter in southern South America. They follow an elliptical migration path; northbound birds pass through Central America about January–April and stage in great numbers in places like Illinois before their final push north. In fall, they take a more easterly route, flying mostly over the western Atlantic and Caribbean Sea to the wintering grounds in Patagonia. The bird has one of the longest known migratory routes of over 40,000 km (25,000 mi). Of this, 3,900 km (2,400 mi) is over open ocean where it cannot stop to feed or drink. It does this from body fat stores that it stocks up on prior to the flight. It is a regular vagrant to western Europe.
A comparison of dates and migratory patterns leads to the conclusion that Eskimo curlews and American golden plovers were the most likely shore birds to have attracted the attention of Christopher Columbus to the nearby Americas in early October 1492, after 65 days at sea out of sight of land.


This bird uses scrape nests, lining them with lichens, grass, and leaves. At its breeding grounds, it is very territorial, displaying aggressively to neighbors. Some American plovers are also territorial in their wintering grounds.
The American golden plover lays a clutch of four white to buff eggs that are heavily blotched with both black and brown spots. The eggs generally measure around 48 by 33 mm (1+7⁄8 by 1+5⁄16 in). These eggs are incubated for a period of 26 to 27 days, with the male incubating during the day and the female during the night. The chicks then hatch precocial, leaving the nest within hours and feeding themselves within a day.


These birds forage for food on tundra, fields, beaches and tidal flats, usually by sight. They eat insects and crustaceans.


Large numbers were shot in the late 19th century and the population has never fully recovered.


Poa annua, or annual meadow grass (known in America more commonly as annual bluegrass or simply poa), is a  widespread low-growing turfgrass in temperate climates. Notwithstanding the reference to annual plant in its name, perennial bio-types do exist. This grass may have originated as a hybrid between Poa supina and Poa infirma.


It has a slightly creeping, fibrous, rootstock. The stem grows from 15–25 cm (6-10 in.) high. It is slightly flattened, due to being folded rather than rolled.
The panicle is open and triangular shaped, 5 to 7.5 cm (2 to 3 in.) long. The spikelets are stalked, awnless, 1 to 2 cm (3/8 to 3/4 in.) long when flowering, and loosely arranged on delicate paired or spreading branches.  Sometimes they are tinged purple.
The vivid green leaves are short and blunt at the tips, shaped like the prow of a small canoe.  They are soft and drooping.  Long sheaths clasp the stem.  The leaves are smooth above and below, with finely serrated edges.  Occasionally the leaves are serrated transversely.
The ligule is pointed and silvery. Compared this to Common Meadowgrass Poa pratensis, which has a squared ligule, and Poa trivialis, which has a pointed, but less silvery ligule.
The leaves are smooth above and below, with finely serrated edges.  Occasionally the leaves are serrated transversely.
It is in flower all year around except for severe winters.  The seeds ripen and are deposited 8 months of the year.  The plant grows rapidly from seed, flowering within 6 weeks, seeding and then dying.


It is a common weed of cultivation, known in the Americas as annual bluegrass. It occurs as a common constituent of lawns, where it is also often treated as a weed, and grows on waste ground. Many golf putting greens, including the Oakmont Country Club greens, are annual bluegrass, although many courses have converted to creeping bentgrass (Agrostis stolonifera).
It has appeared on King George Island in the Antarctic South Shetland Islands  as an invasive species, as well as on Australia's subantarctic Heard and Macquarie Islands.


Poa bulbosa is a species of grass known by the common names bulbous bluegrass or bulbous meadow-grass. It is native to Eurasia and North Africa, but it is present practically worldwide as an introduced species. It is widespread in the United States and southern Canada. It was introduced to the eastern United States around 1906 and the western US in 1915 as a contaminant in shipments of alfalfa seed. It was intentionally planted on both the east and west coasts to control weeds and prevent erosion. Today it is a common grass across the continent and is a noxious weed in some areas. It is a sturdy, hardy, persistent, aggressive grass that easily outcompetes many other plants and becomes the dominant species in disturbed habitat types, such as overgrazed fields.
This is an annual or perennial grass forming dense clumps up to about 60 centimeters tall. The stems are smooth and hollow and usually have bulbous sections at their bases about a centimeter in length. The grass is more likely to have bulbous sections if it is growing in a drier area, and study has indicated the bulbous sections are mostly water. If the bulbous bases are detached and replanted they can give rise to new plants. The inflorescence is a wide cluster of branches bearing green leaflike spikelets with darker bases that contain bulbils. Viable seed is rarely produced, and the plant usually reproduces asexually via its basal bulbous sections and via bulbils. Although the plant reproduces vegetatively (asexually) most of the time, it has been shown to possess high genetic variability.
Many types of animals, including wild and domesticated ruminants, small mammals, and birds, readily consume this grass, especially the bulbils in the inflorescences, which contain some starches and fats. The grass is used as a fodder and a pasture grass in parts of its native region.


The red-necked grebe (Podiceps grisegena) is a migratory aquatic bird found in the temperate regions of the northern hemisphere. Its wintering habitat is largely restricted to calm waters just beyond the waves around ocean coasts, although some birds may winter on large lakes. Grebes prefer shallow bodies of fresh water such as lakes, marshes or fish-ponds as breeding sites.
The red-necked grebe is a nondescript dusky-grey bird in winter. During the breeding season, it acquires the distinctive red neck plumage, black cap and contrasting pale grey face from which its name was derived. It also has an elaborate courtship display and a variety of loud mating calls. Once paired, it builds a nest from water plants on top of floating vegetation in a shallow lake or bog.
Like all grebes, the Red-necked is a good swimmer, a particularly swift diver, and responds to danger by diving rather than flying. The feet are positioned far back on the body, near the tail, which makes the bird ungainly on land. It dives for fish or picks insects off vegetation; it also swallows its own feathers, possibly to protect the digestive system. The conservation status of its two subspecies—P. g. grisegena found in Europe and western Asia, and the larger P. g. holboelii (formerly Holbœll grebe), in North America and eastern Siberia—is evaluated as Least Concern, and the global population is stable or growing.


The red-necked grebe was described by the French polymath Georges-Louis Leclerc, Comte de Buffon in  1781 in his Histoire Naturelle des Oiseaux. The bird was also illustrated in a hand-coloured plate engraved by François-Nicolas Martinet in the Planches Enluminées D'Histoire Naturelle which was produced under the supervision of Edme-Louis Daubenton to accompany Buffon's text.  Neither the plate caption nor Buffon's description included a scientific name but in 1783 the Dutch naturalist Pieter Boddaert coined the binomial name Colymbus grisegena in his catalogue of the Planches Enluminées. The type locality was subsequently designated as France. The red-necked grebe is now placed in the genus Podiceps that was erected by the English naturalist John Latham in 1787. The genus name Podiceps comes from Latin podicis, "vent" or "anus", and pes, "foot", and is a reference to the placement of a grebe's legs towards the rear of its body. The species name grisegena is from Latin griseus (grey) and gena (cheek) and refers to the face pattern of the breeding adult.
Grebes are small to medium-large water birds with lobed, rather than webbed, toes. There are several genera, of which the most widespread is Podiceps with nine species, one recently extinct. The red-necked grebe's closest relative is the fish-eating great crested grebe of Europe and western Asia. It is possible that the red-necked grebe originally evolved in North America and later spread to Europe, where a change of diet to include more insects helped to reduce competition with its larger cousin. Fossils of the species dating to the middle Pleistocene have been found in Italy.
The red-necked grebe has two subspecies, the nominate subspecies P. g. grisegena in Europe and western Asia, and P. g. holboelii (Holboell's grebe, named for Danish explorer of Greenlandic birds Carl Peter Holbøll) in North America and eastern Siberia. The east Asian birds have slightly smaller bills than the American form, although the differences are too small to merit separation as a third subspecies.


The red-necked grebe is a medium-large grebe, smaller than the great crested grebe of Eurasia, and the western and Clark's grebes of North America, but noticeably larger than other northern grebe species. The adult of the nominate European subspecies is 40–50 cm (16–20 in) long with a 77–85 cm (30–33 in) average wingspan, and weighs 692–925 g (24.4–32.6 oz). In breeding plumage, it has a black cap that extends below the eye, very pale grey cheeks and throat, a rusty red neck, dark grey back and flanks, and white underparts. The eyes are dark brown and the long, pointed bill is black with a yellow base.
The winter plumage of the red-necked grebe is duskier than that of other grebes; its dark grey cap is less defined, and merges into the grey face, and a pale crescent that curves around the rear of the face contrasts with the rest of the head. The front of the neck is whitish or light grey, the hind neck is darker grey, and the yellow of the bill is less obvious than in summer. Although the red-necked grebe is unmistakable in breeding plumage, it is less distinctive in winter and can be confused with similar species. It is larger than the Slavonian (horned) grebe, with a relatively larger bill and a grey, rather than white face. It is closer in size to the Eurasian great crested grebe, but that species is longer-necked, has a more contrasting head pattern, and always shows white above the eye.
The sexes are similar in appearance, although the male averages heavier than the female. Chicks have a striped head and breast, and older juveniles have a striped face, diffuse blackish cap, pale red neck and extensive yellow on the bill.
The subspecies P. g. holboelii is larger than the nominate race at 43–56 cm (17–22 in) length, with a 61–88 cm (24–35 in) wingspan, and a weight of 750–1,600 g (26–56 oz). The plumages are the same as those of the nominate race, although the adult's bill is more extensively yellow. The difference in size between the sexes is greater than for this subspecies that for P. g. grisegena.
The red-necked grebe flies with its long neck extended and its large feet trailing behind the body, which gives it a stretched-out appearance. The relatively small wings are grey with white secondaries, and beat very rapidly. Its small wing area means that the grebe is unable to take off from land, and needs a lengthy run across water to gain the speed needed for take-off. Like all grebes, the red-necked is an expert swimmer; it uses its feet for propulsion underwater, and steers by rotating its legs, since its tail is too short for this purpose.
This is one of the most vocal grebes during the breeding season, but, like its relatives, it is mainly silent for the rest of the year. It has a loud, wailing or howling display call uooooh, given by a single bird or a pair in duet, by night or during the day, and often from cover. Long sequences of up to 60 consecutive notes may be delivered during singing encounters between rival territorial birds. A great variety of quacking, clucking, hissing, rattling and purring calls are also given, with much individual variation.


Breeding takes place in shallow freshwater lakes, bays of larger lakes, marshes, and other inland bodies of water, often less than 3 ha (7.4 acres) in extent and less than 2 m (6.6 ft) deep. The red-necked grebe shows a preference for waters in forested areas or, further north, in shrub tundra, and favours sites with abundant emergent vegetation, such as reedbeds. The best breeding habitat is fish-ponds, which have an abundance of food in addition to meeting the other requirements. The American subspecies is less tied to a high aquatic plant density, and sometimes breeds on quite open lakes.
All populations are migratory and winter mainly at sea, usually in estuaries and bays, but often well offshore where fish are within diving reach near shallow banks or islands. The preferred passage and wintering habitat is water less than 15 m (49 ft) deep with a sand or gravel bottom, scattered rocks and patches of seaweed. During winter, birds typically feed alone and rarely aggregate into flocks, but on migration, concentrations of over 2000 individuals may occur at favoured staging sites. Migration is usually at night, but may occur during the day, especially when over water. This is particularly noticeable in autumn on the Great Lakes, when up to 18,000 birds may pass Whitefish Point on Lake Superior; these are thought to be Canadian breeders heading for the Atlantic Ocean to winter. This easterly route is longer than that to the Pacific, but avoids the Rockies.
The breeding range of the red-necked grebe overlaps with that of the Slavonian grebe, although the latter species tends to be displaced from sites suited to both. The red-necked grebe prefers an inland temperate climate, and is less successful near coasts and in subarctic and warm temperate zones. It is usually a lowland bird, breeding below 100 m (330 ft), although has nested at up to 1,800 m (5,900 ft) in Turkey.
The nominate subspecies breeds from southern Sweden and Denmark through central and eastern Europe east to western Siberia, and winters mainly in the North and Baltic Seas, with smaller numbers in the Adriatic, Black Sea, Caspian Sea, Mediterranean and on inland lakes. P. g. holboelii breeds in North America in Alaska, western and central Canada, and the northern US east to Minnesota; in Asia it nests in eastern Siberia from Kamchatka south to Hokkaido and west to Mongolia. The Asian birds winter at sea from Japan to the East China Sea, and American breeders winter in the Pacific, mainly from southern Alaska to British Columbia (with smaller numbers south to California), and in the Atlantic from Newfoundland and Labrador to Florida. Some birds remain on the Great Lakes if they are sufficiently ice-free. This species occurs as a rare winter vagrant in Afghanistan, Pakistan and parts of northern and western India.


Red-necked grebes usually nest as isolated pairs with more than 50 m (160 ft) between neighbouring nests, although semi-colonial nesting may occur in suitable sites, where up to 20 pairs each defend a linear territory. Semi-colonial breeding is more likely to occur in prime locations, such as large floating mats of vegetation with no connection to the shoreline. Such sites, safe from most predators and large enough to provide some wind and wave protection, have grebes nesting much closer than shoreline breeders, down to 10 m (33 ft). Pairs nesting in these colonies produce larger clutches of eggs, which hatch earlier in the season and result in larger broods. The territory is defended with various threat displays, including wing-spreading, hunching, and bill-thrusting; pairs breeding in colonies are more aggressive, less likely to leave the nest unguarded and show a greater tendency to move out of sight of the colony when not incubating. Breeding is often in loose association with gulls or other colonial water birds.
The monogamous pair forms in April or May on migration or at the breeding water, and a highly vocal courtship ritual commences. The elaborate breeding performance includes head-shaking, a head-lowered "cat" display, parallel rushes in an upright position and mutual presentations of green weeds, and culminates in a "penguin" dance in which pair members raise the whole body upright, breast to breast.
Like all grebes, the red-necked grebe nests near water into which it can escape, since the position of the legs far back on the body prevents fast movement on land. It often breeds further within reed beds than other grebes. The nest is a floating platform of plant matter anchored to submerged or emergent vegetation, in water 0.5–0.75 m (20–30 in) deep, and with the bulk of nest below the water line. Egg-laying mainly takes place from mid-April to May in Europe, and somewhat later, from mid-May to June, in North America. Parents may leave the nest for significant periods of time during the night, possibly to evade nocturnal predators. It is unclear whether this is for self-protection or to protect the eggs by diverting attention from the nest; the clutch does not appear to suffer from this temporary abandonment, whatever the reason.
The red-necked grebe lays four or five (range one to nine) dull white or pale blue eggs, which average 3.4 cm (1.3 in) in breadth, 5.1 cm (2.0 in) in length, and weigh about 30.5 g (1.08 oz), of which 10% is shell. Parents take turns to incubate the eggs for 21–33 days until the precocial downy chicks hatch; they immediately climb onto the parent's back, where they spend most of their time until they are 10–17 days old. The young may be fed by the parents for up to 54 days after fledging, and can fly at 50–70 days. The parents do not interfere with the feeding of their chicks when they are still being carried. Later they care for the younger chicks for longer, and are aggressive with the older offspring. This equalises the post-fledging survival of all chicks, and encourages their independence. The brood may be split, so that each parent feeds only some of the chicks. This spreads the feeding demand equally between the parents.
After breeding the adults moult their wing feathers and are temporarily flightless; migration commences once the flight feathers have regrown. The red-necked grebe is normally single-brooded, although second broods and re-nesting after a clutch has been lost may extend nesting into July or August.
Eggs may be destroyed and chicks killed by a range of predators, including the raccoon in North America and the carrion crow in Europe. Pike may take swimming chicks. On average, for each adult, 0.65 young birds are still alive by their fourth month, although the mortality rate for the adult is unknown. Red-necked grebes attempt to evade birds of prey by diving; when feeding, dives average less than 30 seconds, although escape dives are more prolonged.


On the breeding grounds, the red-necked grebe feeds mainly on invertebrates including adult and larval aquatic insects, such as water beetles and dragonfly larvae, crayfish and molluscs. Fish (such as smelt) may be important locally or seasonally, especially for the American subspecies, and crustaceans can constitute up to 20% of the grebe's diet. Birds breeding at the coast often make foraging flights to inland lakes or offshore areas to feed.
Aquatic prey is obtained by diving or by swimming on surface with the head submerged, and terrestrial insects and their larvae are picked off vegetation. A line slanting downward from the eye to the tip of the opened lower mandible may be used for sighting on prey before diving or when swimming under water. The grebe probably opens its bill and looks down the eye-line toward its target.
European breeders, which have to compete with the larger great crested grebe for fish, eat a greater proportion of invertebrates than the longer-billed American subspecies, although both races eat mainly fish in winter. Birds of the nominate subspecies from the northernmost breeding populations in Finland and Russia, beyond the range of great crested grebe, have a longer and more slender bill than those further south, reflecting a greater proportion of fish in the diet where their main competitor is absent. If food is scarce, parents may desert unhatched eggs, or allow the smallest chicks to starve, although the latter strategy appears not to be particularly efficient in protecting the older chicks.
Like other grebes, the red-necked grebe ingests large quantities of its own feathers, which remain in the bird's stomach. Feathers are not only swallowed by adults, mainly during self-preening, but are often fed to the young, sometimes within a day of hatching. These feathers soon decompose into a felt-like, amorphous mass. The function of the feathers in the stomach is unknown, although it has been suggested that they help to protect the lower digestive tract from bones and other hard, indigestible material.


The red-necked grebe has a large range, estimated at 1–10 million square kilometres (0.4–3.8 million square miles), and a global population of 150,000–370,000 individuals, with P. g. holboellii about twice as numerous as the nominate race. The population trend has not been quantified, but it is not believed to meet the thresholds for the population decline criterion (declining more than 30% in ten years or three generations) of the IUCN Red List. For these reasons, the species is evaluated as Least Concern.
The red-necked grebe is one of the species to which the Agreement on the Conservation of African-Eurasian Migratory Waterbirds (AEWA) applies. Parties to the Agreement are required to engage in a wide range of conservation strategies which are describes in a detailed action plan. The plan is intended to address key issues such as species and habitat conservation, management of human activities, research, education, and implementation.
The Red-necked Grebe was hunted by humans in northern Europe in the Mesolithic and Paleolithic periods, but there is no evidence that there is any significant level of hunting at the present time. In North America, there are potential threats from pollutants such as polychlorinated biphenyls (PCBs) and pesticides such as DDT which cause reduced reproductive success due to egg sterility and eggshell thinning. Breeding areas may be threatened by the modification and degradation of lakes and by human disturbance from water-based recreational activities. There is no evidence to suggest that these threats could result in a significant risk to the overall population; more than 70% of North American red-necked grebes breed in Canada, where the population is stable or increasing.


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The great grebe (Podiceps major) is the largest species of grebe in the world. A disjunct population exists in northwestern Peru, while the main distribution is from extreme southeastern Brazil to Patagonia and central Chile. The population from southern Chile is considered a separate subspecies, P. m. navasi.


The great grebe was described by the French polymath Georges-Louis Leclerc, Comte de Buffon in 1781 in his Histoire Naturelle des Oiseaux. The bird was also illustrated in a hand-coloured plate engraved by François-Nicolas Martinet in the Planches Enluminées D'Histoire Naturelle which was produced under the supervision of Edme-Louis Daubenton to accompany Buffon's text.  Neither the plate caption nor Buffon's description included a scientific name but in 1783 the Dutch naturalist Pieter Boddaert coined the binomial name Colymbus major in his catalogue of the Planches Enluminées. Buffon mistakenly believed that his specimen had come from Cayenne in French Guiana. The great grebe is now placed in the genus Podiceps that was erected by the English naturalist John Latham in 1787.
Two subspecies are recognised:
P. m. major, (Boddaert, 1783), West Peru, Paraguay, southeastern Brazil, Uruguay to central Chile & southern Argentina
P. m. navasi, (Manghi, 1984), Southern Chile


This species occurs mainly in open waterways. Most birds are seen on low altitude lakes and sluggish rivers (often surrounded by forests), as well as estuarine marshes. While breeding, it frequents the heavily vegetated inlets off of large lakes. Outside of the breeding season, most birds will move to estuaries and bays, usually heavy with kelp (occasionally, they even are seen on the open sea). Non-breeding birds may live along the coasts all year.
These birds are widespread and common through most of their range. Much of their native habitat has not been cleared, especially in the southern portions of their range.


This is a very large grebe, with proportions more like a goose or a cormorant than a typical grebe. They range in length from 67 to 80 cm (26.5 to 31.5 in) and usually weigh about 1,600 grams (3.5 pounds), but can scale to at least 2 kg (4.4 lb). They are buffy-rufous on the neck and chest, blackish on the back and have a whitish belly. The head is sooty gray with a reddish-brown eye. Due to its size and unique coloration, the great grebe is unlikely to be confused with any other bird, including other grebes.


The great grebe lives on a diet mostly of fish, sometimes over 11 cm (4.3 in) long, but usually smaller. Prey competition can occasionally occur with the neotropic cormorant over fish, but that species (in spite of smaller body size) usually takes larger fish. Also insects, crustaceans and mollusks are taken. The diet can switch to almost half crabs during the wintertime along the coasts, and these birds can also take the young of other waterbirds, especially coots.


The black-necked grebe or eared grebe (Podiceps nigricollis) is a member of the grebe family of water birds. It was described in 1831 by Christian Ludwig Brehm. There are currently three accepted subspecies, including the nominate subspecies. Its breeding plumage features a distinctive ochre-coloured plumage which extends behind its eye and over its ear coverts. The rest of the upper parts, including the head, neck, and breast, are coloured black to blackish brown. The flanks are tawny rufous to maroon-chestnut, and the abdomen is white. When in its non-breeding plumage, this bird has greyish-black upper parts, including the top of the head and a vertical stripe on the back of the neck. The flanks are also greyish-black. The rest of the body is a white or whitish colour. The juvenile has more brown in its darker areas. The subspecies californicus can be distinguished from the nominate by the former's usually longer bill. The other subspecies, P. n. gurneyi, can be differentiated by its greyer head and upper parts and by its smaller size. P. n. gurneyi can also be told apart by its lack of a non-breeding plumage. This species is present in parts of Africa, Eurasia, and the Americas.
The black-necked grebe uses multiple foraging techniques. Insects, which make up the majority of this bird's diet, are caught either on the surface of the water or when they are in flight. It occasionally practices foliage gleaning. This grebe dives to catch crustaceans, molluscs, tadpoles, and small frogs and fish. When moulting at saline lakes, this bird feeds mostly on brine shrimp. The black-necked grebe makes a floating cup nest on an open lake. The nest cup is covered with a disc. This nest is located both in colonies and by itself. During the breeding season, which varies depending on location, this species will lay one (sometimes two) clutch of three to four eggs. The number of eggs is sometimes larger due to conspecific brood parasitism. After a 21-day incubation period, the eggs hatch, and then the nest is deserted. After about 10 days, the parents split up the chicks between themselves. After this, the chicks become independent in about 10 days, and fledge in about three weeks.
Although it generally avoids flight, the black-necked grebe travels as far as 6,000 kilometres (3,700 mi) during migration. In addition, it becomes flightless for two months after completing a migration to reach an area where it can safely moult. During this moult, the grebe can double in weight. The migrations to reach these areas are dangerous, sometimes with thousands of grebe deaths. In spite of this, it is classified as a least concern species by the International Union for Conservation of Nature (IUCN). It is likely that this is the most numerous grebe in the world. There are potential threats to it, such as oil spills, but these are not likely to present a major risk to the overall population.


This species was first described by Carl Ludwig Hablitz in 1783 as Colymbus caspicus, from a bird in Bandar-e Anzali. This was originally thought to be a synonym for the horned grebe, until Erwin Stresemann discovered that the description applied more to the black-necked grebe in 1948. Before this, the earliest description was thought to be by Christian Ludwig Brehm in 1831, who gave this bird its current scientific name of Podiceps nigricollis from a German bird. To resolve this, the International Commission on Zoological Nomenclature suppressed the name C. caspicus. The genus name Dytes is sometimes used for this species, a placement which was formalized by Robert Ridgway in 1881.
This bird is closely related to the silvery grebe and the Junin grebe. The extinct Colombian grebe is sometimes considered to be a subspecies of this species, in addition to three other extant subspecies:
P. n. nigricollis – (Brehm, CL, 1831): nominate, is found from western Europe to western temperate Asia (wintering to the south and west), in central and eastern Asia, and in eastern Africa
P. n. gurneyi – (Roberts, 1919): is found in southern Africa
P. n. californicus – (Heermann, 1854): is found from southwestern Canada through the western U.S. It winters as far south as Guatemala
The generic name, Podiceps, comes from two Latin words: podicis, meaning "vent" or "anus" and pes meaning "foot". This is a reference to the attachment point of the bird's legs—at the extreme back end of its body. The specific epithet nigricollis is Latin for "black-necked": niger means "black" and collis means "neck". The subspecies epithet californicus comes from "California", while gurneyi comes from the name of British ornithologist John Henry Gurney Sr.
"Black-necked grebe" has been designated the official name by the International Ornithological Committee (IOC). Both common names for this species refer to features visible when the bird is in its breeding plumage; in such plumage, it has an all-black neck and a spray of golden plumes on each side of its head. The name "eared grebe" is nearly a century older than the name "black-necked grebe". The latter was first used in 1912 by Ernst Hartert, in an effort to bring the common name of the species in line with its scientific name. The name "eared grebe" is still used in North America to refer to this bird.


The black-necked grebe usually measures between 28 and 34 centimetres (11 and 13 in) in length and weighs 265 to 450 grams (9.3 to 15.9 oz). The bird has a wingspan range of 20.5-21.6 in (52-55 cm). The nominate subspecies in breeding plumage has the head, neck, breast, and upper parts coloured black to blackish brown, with the exception of the ochre-coloured fan of feathers extending behind the eye over the eye-coverts and sides of the nape. This eye is mostly red, with a narrow and paler yellow ring on the inner parts of the eye and an orange-yellow to pinkish-red orbital ring. The thin, upturned bill, on the other hand, is black, and is connected to the eye by a blackish line starting at the gape. Sometimes, the foreneck can be found to be mostly tinged brown. The upperwing is blackish to drab brown in colour and has a white patch formed by the secondaries and part of the inner primaries. The flanks are coloured tawny rufous to maroon-chestnut and have the occasional blackish fleck. The underwing and abdomen is white, with an exception to the former being the dark tertials and the mostly pale grey-brown outer primaries. The legs are a dark greenish grey. The sexes are similar.
In non-breeding plumage, the nominate has greyish-black upper parts, cap, nape, and hindneck, with the colour on the upper portion of the latter being contained in a vertical stripe. The dark colour of the cap reaches below the eye and can be seen, diffused, to the ear-coverts. Behind the ear-coverts on the sides of the neck, there are white ovals. The rest of the neck is grey to brownish-grey in colour and has white that varies in amount. The breast is white, and the abdomen is whitish. The flanks are coloured in a mix of blackish-grey with white flecks. The colour of the bill when not breeding differs from that of the breeding plumage, with the former being significantly more grey.
The juvenile black-necked grebe is similar to the non-breeding adult. There are differences, however, including the fact that the dark areas are usually more brownish in the juvenile, with less black. The lores are often tinged pale grey, with whitish marks behind the eye. On the sides of the head and upper neck, there is a buffy or tawny tinge. The chick is downy and has a blackish-grey head with stripes and spots that are white or pale buff-grey. The throat and foreneck are largely pale. The upper parts are mostly dark grey in colour, and the abdomen is white.
The subspecies californicus usually has a longer bill compared to the nominate, and has brown-grey inner primaries during the breeding season. When not breeding, the nominate has diffuse and pale lores less often than Podiceps nigricollis californicus. The other subspecies, P. n. gurneyi, is the smallest of the three subspecies, in addition to having a greyer head and upper parts. The adult of this subspecies also has a rufous-brown tinge on its lesser wing-coverts. It also lacks a non-breeding plumage, in addition to the tufts on the side of its head being paler.


When breeding, the black-necked grebe gives a quiet "ooeek" that ascends in pitch from an already high pitch. This call is also used as a territorial call, in addition to a low and fast trill, which itself is also used during courtship. Another call is a short "puuii" or "wit". This grebe is silent when it is not the breeding season and when it is feeding or resting.


This species breeds in vegetated areas of freshwater lakes across Europe, Asia, Africa, northern South America and the southwest and western United States. After breeding, this bird migrates to saline lakes to moult. Then, after completing the moult and waiting for sometimes several months, it migrates to winter in places such as the south-western Palearctic and the eastern parts of both Africa and Asia. It also winters in southern Africa, another place where it breeds. In the Americas, it winters as far south as Guatemala, although the wintering population there is mainly restricted to islands in the Gulf of California, the Salton Sea, and Baja California. When not breeding, its habitat is primarily saline lakes and coastal estuaries.


This grebe is highly gregarious, usually forming large colonies when breeding and large flocks when not.


This species builds its floating nest in the usually shallow water of open lakes. The nest itself is anchored to the lake by plants. It is built by both the male and the female and made out of plant matter. Most of it is submerged, with the bottom of the shallow cup usually being level with the water. Above the cup, there is a flat disc. This grebe nests both in colonies and by itself. When it does not nest by itself, it will often nest in mixed-species colonies made up of black-headed gulls, ducks, and various other waterbirds. The space between the nests in these colonies is often 1 to 2 metres (3.3 to 6.6 ft). Whether it nests in colonies or not has an effect on the dimensions of the nest. When the bird is not in a colony, the nest has an average diameter of 28 centimetres (11 in), although this can vary, with nests ranging from about 20 centimetres (8 in) to over 30 centimetres (12 in). This is compared to nests in colonies, which have an average diameter of about 25.5 centimetres (10 in). It is suggested that rarely some pairs of this grebe will steward over multiple nests when in colonies.
Pair formation in the black-necked grebe usually starts during pauses in the migration to the breeding grounds, although it occasionally occurs before, in wintering pairs. This pair formation continues after this grebe has arrived to its breeding grounds. Courtship occurs when it arrives at the breeding lake. The displays are performed in the middle of the lake. There is no territory involved in courting; individuals used the whole area of the lake. When advertising for a mate, a black-necked grebe will approach other black-necked grebes with its body fluffed out and its neck erect. It closes its beak to perform a call, poo-eee-chk, with the last note only barely audible. Courtship generally stops at the start of nesting.
In the Northern Hemisphere, this bird breeds from April to August. In east Africa, the breeding season is at least from January to February, while in southern Africa, the breeding season is from October to April.
The black-necked grebe is socially monogamous. Conspecific or intraspecific brood parasitism, where the female lays eggs in the nest of others of their own species is common with nearly 40% of nests being parasitized on average. In terms of territory, this grebe will defend only its nest site.
This grebe lays a clutch, and sometimes two clutches, of three to four chalky greenish or bluish eggs. Nests that have been parasitized, however, will have two more eggs on average, even though the number the host lays is about the same no matter if it has been parasitized or not. The eggs, although initially immaculate, do get stained by plant matter that the nest is built out of. The eggs measure 45 by 30 millimetres (1.8 by 1.2 in) on average and are incubated by both parents for about 21 days. The laying date of the eggs is somewhat synchronized, with birds in small colonies having the laying dates spread out by just a few days, compared to large colonies, where the laying date is spread out over more than 10 days.
After the chicks hatch, the birds will desert their nest. Even though the young can swim and dive during this time, they rarely do, instead staying on the parents' backs for four days after hatching. This behaviour is present in all grebes, and is likely to have evolved because it reduces travel costs, specifically those back to the nest to brood the chicks and give them food. After about 10 days, the parents split the chicks up, with each parent taking care of about half of the brood. After this split, the chicks are independent in about 10 days, and fledge in about three weeks.
When disturbed while incubating, this bird usually (just under 50% of the time) partly covers its eggs with nest material when the disruption is not sudden, but a bird with an incomplete clutch usually does not attempt to cover the eggs. When the disruption is sudden, on the other hand, the black-necked grebe usually (just under 50% of the time) does not cover its eggs. In comparison, other species of grebes cover up their eggs when leaving the nest. Predation is usually not the primary cause of egg loss, with most nesting failures occurring after the chicks have hatched. A major cause of this is the chilling of the young.


The black-necked grebe forages mainly by diving from the water, with dives usually lasting less than 30 seconds. These dives are usually shorter in time when in more shallow water. In between dives, this grebe rests for an average of 15 seconds. When feeding on brine shrimp at hypersaline lakes, it likely uses its large tongue to block the oral cavity. It is hypothesized that it then crushes prey against its palate to remove excess water. It also forages by gleaning foliage, plucking objects off of the surface of water, having its head submerged while swimming, and sometimes capturing flying insects.
This grebe eats mostly insects, of both adult and larval stages, as well as crustaceans, molluscs, tadpoles, and small frogs and fish. When moulting at lakes with high salinity, although, this bird feeds mostly on brine shrimp. The behaviour of black-necked grebes changes in response to the availability of brine shrimp; bodies of water with more shrimp have more grebes, and grebes spend more time foraging when the amount of shrimp and the water temperature decreases.
The young are fed one at a time by the parents, with one bird carrying the young while the other feeds it. The young take food by grabbing it, with their beaks, from their parents, or by grabbing food dropped into the water. When a young bird cannot grab the food, then the adults submerge their bill into the water and shake their bill to break up the food.


When breeding is over, the black-necked grebe usually partakes in a moult migration to saline lakes. It especially prefers lakes with large numbers of invertebrate prey, so it can fatten up while moulting and before going on its winter migration. Some birds, although, moult when on the breeding grounds, but most do not moult until the end of the moult migration. This migration is dangerous, with hundreds and sometimes thousands of birds being killed by snowstorms when traveling to places such as Mono Lake.
When it finishes its moult migration, this bird moults its remiges between August and September, which makes it unable to fly. This moult is preceded by an increase in weight. During the moult, the breast muscles atrophy. When the moult is completed, this grebe continues to gain weight, often more than doubling its original weight. This additional fat is used to power the black-necked grebe's overnight fall migration to its wintering grounds. The fat is most concentrated in the abdomen, second most in the thorax, and least in the chest. This bird usually starts its migration earlier when shrimp is more abundant and when the moulting lake is at a higher than average temperature. It generally leaves on a clear night with lower than average surface temperatures.


This grebe is one of the most inefficient fliers among birds. Generally, it avoids flying at all costs and reserves long-distance flight exclusively for migration. This is combined with the fact that this bird is flightless for two months of the year during its moult. However, when migrating, it travels as much as 6,000 kilometres (3,700 miles) to reach rich feeding areas that are exploited by few other species. In flight, the shape of this grebe is like a loon: straight neck, legs trailing, and wings beating often.
When diving, this bird pulls its head back and then arches it forward into the water, with the body following and a slight springing. The legs start moving only after they are underwater. When swimming on the surface of the water, the body of this grebe is relatively high, although none of the underparts are seen. The neck is held straight up in a relaxed manner, with the bill being held forward and parallel to the water. Each of the feet perform strong alternating strokes.


Large-scaled deaths (such as 150,000 birds on the Salton Sea in 1992) from erysipelas, avian cholera, avian botulism, and West Nile virus have been recorded in the past. In 2013 at the Great Salt Lake, for example, there was an outbreak of West Nile virus that could be the largest recorded in birds in the US. The cause of the outbreak and modes of transmission are unknown, but there has been speculation about the latter. Since West Nile virus is able to survive in brine shrimp and, for a temporary time, the water of the Great Salt Lake around the temperature it was when the die-off happened. Thus, it is likely that grebes could become infected by eating diseased shrimp or being in contaminated water. It is also theorized that West Nile virus could be transmitted among grebes through contact with the excrement of an infected grebe. Avian cholera, another disease that can cause large scale die-offs in this grebe, can be brought on by unknown biotoxins, pathogens, and problems with feather waterproofing.


As of 2016, the black-necked grebe is classified as least concern by the International Union for Conservation of Nature (IUCN). The trend of the population is uncertain, as some populations are decreasing, whereas others are stable, have an uncertain trend, or are increasing. The justification for the current classification of this species is its very large population (estimated around 3.9–4.2 million individuals) combined with a large estimated extent of occurrence (about 155 million km² (60 million sq mi)). This grebe is probably the most numerous grebe in the world.
Unknown biotoxins, pathogens, and the impairment of feather waterproofing can lead to hypothermia and avian cholera. Since this grebe usually winters on the coast, it is also vulnerable to oil pollution. Large-scale disease, such as avian cholera, could threaten the species. These and other factors, such as human disturbance, including collisions with power transmission lines, contribute to declining populations in certain areas. This species used to be threatened in North America by the millinery industry, which helped facilitate the hunting of the birds, and egg collectors. Although this is true, this grebe is hunted in the Gilan Province in Iran, for both commercial and recreational purposes. However, there is no evidence suggesting that these threats could result in a significant risk for the overall population.


The pied-billed grebe (Podilymbus podiceps) is a species of the grebe family of water birds. Because the Atitlán grebe (Podilymbus gigas) has become extinct, the Pied-Billed Grebe is now the sole extant member of the genus Podilymbus. The pied-billed grebe is primarily found in ponds throughout the Americas. Other names of this grebe include American dabchick, rail, dabchick, Carolina grebe, devil-diver, dive-dapper, dipper, hell-diver, pied-billed dabchick, pied-bill, thick-billed grebe, and water witch.


The pied-billed grebe was described by Carl Linnaeus in the 10th edition of his Systema Naturae in 1758 as Colymbus podiceps. The binomial name is derived from Latin Podilymbus, a contraction of podicipes ("feet at the buttocks", from podici-, "rump-" + pes, "foot")—the origin of the name of the grebe order—and Ancient Greek kolymbos, "diver", and podiceps, "rump-headed", from podici- + New Latin ceps.
Outside its own genus, the closest relatives of the pied-billed grebe are the small grebes of the genus Tachybaptus.


Pied-billed grebes are small, stocky, and short-necked. They are 31–38 cm (12–15 in) in length, with a wingspan of 45–62 cm (18–24 in) and weigh 253–568 g (8.9–20.0 oz). They are mainly brown, with a darker crown and back. Their brown color serves as camouflage in the marshes they live in. They do not have white visible under their wings when flying, like other grebes. Their undertail is white and they have a short, blunt chicken-like bill that is a light grey color, which in summer is encircled by a broad black band (hence the name). In the summer, its throat is black. There is no sexual dimorphism. Juveniles have black and white stripes and look more like winter adults. This grebe does not have webbed feet. Its toes have lobes that come out of the side of each toe. These lobes allow for easy paddling. When flying, the feet appear behind the body due to the feet's placement in the far back of the body. Because of the feet placement, they are not able to walk on land.
These grebes may be confused with the least grebe, although that species is much smaller and has a thinner bill. Other similarly sized grebes are very distinct in plumage, i.e. the eared grebe and horned grebe. Both species bear much more colorful breeding plumage, with rufous sides, golden crests along the side of the head against contrasting slaty color (also a rufous neck in the horned); while in winter, both the eared and horned grebes are pied with slaty and cream color and have red eyes. Because of the pied-billed grebe's duck-like habits, some inexperienced observers may confuse it with a duck. However, pied-billed grebes have a very different bill shape (shorter, pointed at the tip, and flattened along the sides), as well as being shorter-necked and shorter-bodied than a duck. Also, unlike ducks, the closest living relatives of the grebe family are flamingos.
Pied-billed grebes are part of the order Saurischia.


Its call is unique, loud and sounds like a "whooping kuk-kuk-cow-cow-cow-cowp-cowp." Its call is similar to the yellow-billed cuckoo.


They are most commonly found throughout North America, Central America, the Caribbean, and South America year round. During the summer breeding season, they are most prevalent in central, northern and northeastern Canada. If they live in an area where the water freezes in the winter they will migrate. Migrating birds generally meet with year-round birds in September and October. They migrate at night. Most migratory birds leave in March or April. They make occasional appearances in Europe and Hawaii. In the United Kingdom, pied-billed grebe visits have numbered 45 sightings as of 2019, appearing generally in October to January. One bird in England bred with a little grebe, producing hybrid young. It is the only grebe on record to have visited the Galapagos Islands.
Pied-billed grebes are found in freshwater wetlands with emergent vegetation, such as cattails. They are occasionally found in salt water. When breeding they are found in emergent vegetation near open water, and in the winter they are primarily found in open water due to the lack of nests to maintain. They may live near rivers, but prefer still water. They may be found in higher elevations when migrating. They will breed in restored and man-made wetlands.
Pied-billed grebes live approximately 10–12 years.


Pied-billed grebes rarely fly. They make a slow dive frequently, especially when in danger, diving to about 20 ft (6.1 m) or less. They dive for about 30 seconds and may move to a more secluded area of the water, allowing only the head to be visible to watch the danger dissipate. This frequency in diving has earned them the description of being reclusive or shy in nature. It has also earned them nicknames like "hell-diver." Pied-billed grebes can trap air in their feathers, which controls their buoyancy. They rarely spend time in flocks. Their courtship include calling and sometimes duets. Males will show territorial behaviour if another male is at the edge of his territory. They face each other and then turn their heads and bills up. Then they turn away and start calling. Then they turn back around to look at one another.


The pied-billed grebe breeds in south-central Canada, throughout the United States, Central America, the Caribbean, and temperate South America. These grebes may lay up to two sets of eggs a year. Their nests sit on top of the water, their eggs sitting in vegetation that resides in the water. Grebes lay between three and ten bluish white smooth elliptical eggs with the female starting the incubation process. They are incubated for around 23 days by both parents, with the female taking over incubation duties towards the end of that time period. They will cover the nest with nesting material if they have to leave it for an extended period of time.
Young grebes may leave the nest within one day of hatching. They are downy at birth. Yellow skin is seen between the lore and top of the head. They do not swim well and stay out of the water. They sleep on their parents' backs. Within four weeks they start swimming. When alerted they will climb on the back of a parent grebe and eventually mature to dive under the water like their parents. Both parents share the role of raising the young – both feeding and carrying them on their backs. Sometimes the parents will dive underwater to get food with the chicks on their backs.


Pied-billed grebes feed mainly on aquatic invertebrates, and also on small fish and amphibians (frogs, tadpoles). They dive to obtain food. Their bills allow them to crush crustaceans, like crawfish. They may also eat plants. They have been shown to eat their own feathers, like other grebes, to aid in digestion (prevent injury from small bones). They will also feed their feathers to their young.


They are extremely sensitive to disturbances, especially by humans. While breeding, if scared, adults may abandon their nests without protecting the eggs. The waves from boats can destroy the nests and their sounds easily frighten the birds.


Pied-billed grebe feathers are thick and soft. Their feathers were formerly used as decorations on hats and earmuffs and they were hunted in the eastern United States, in the 19th century.


The grebes are declining in New England. The reasons are unknown. The states of Connecticut and New Hampshire have declared the pied-billed grebe as endangered. In New Jersey and Massachusetts, they have been declared threatened. In Vermont they are of "special concern." In Rhode Island they are locally extinct. Pied-billed grebes are protected under the Migratory Bird Treaty Act of 1918.
Habitat loss is the grebe's biggest threat. The draining, filling, and general destruction of wetlands causes a loss in their breeding habitats. However, they are still common in the majority of their distribution areas.


Pododesmus macrochisma, common name the green falsejingle or the Alaska jingle, is a species of saltwater clam, a marine bivalve mollusc in the family Anomiidae, the jingle shells.
This species inhabits the northwest Sea of Japan, and more specifically, the coast of the South Primorye at Hokkaido Island, the northern part of Honshu Island, off the southern and eastern Sakhalin in the Kuril Islands, and in the east of Kamchatka in the Commander and Aleutian Islands.  More recently it has been found in the Chukchi Sea near Alaska, potentially due to global warming.


The Carolina chickadee (Poecile carolinensis) is a small passerine bird in the tit family Paridae.


The Carolina chickadee is often placed in the genus Parus with most other tits, but mtDNA cytochrome b sequence data and morphology suggest that separating Poecile more adequately expresses these birds' relationships (Gill et al., 2005). The American Ornithologists' Union has been treating Poecile as distinct genus since 1998. Although it and the black-capped chickadee most likely diverged about 2.5 million years ago, the birds still hybridize in the areas where their ranges overlap. The offspring of mated pairs of hybrid chickadees suffer from lower hatching success, and a male bias sex ratio, consistent with Haldane's rule.


Adults are 11.5–13 cm (4.5–5.1 in) long with a weight of 9–12 g (0.32–0.42 oz), and have a black cap and bib with white sides to the face. Their underparts are white with rusty brown on the flanks; their back is grey. They have a short dark bill, short wings and a moderately long tail. Very similar to the black-capped chickadee, the Carolina chickadee is distinguished by the slightly browner wing with the greater coverts brown (not whitish fringed) and the white fringing on the secondary feathers slightly less conspicuous; the tail is also slightly shorter and more square-ended.
Eggs are about 1.5 cm (0.6 in) long and 1.1 cm (0.4 in) wide. Eggs are white with areas of reddish-brown ranging from dots to small blotches.


The calls and song between the Carolina chickadee and the Black-capped chickadee differ subtly to an experienced ear: the Carolina chickadee's chick-a-dee call is faster and higher pitched than that of the black-capped chickadee, and the Carolina chickadee has a four note fee-bee-fee-bay song, whereas the black-capped omits the high notes. Identification is very difficult even with an excellent view.
The most famous call is the familiar chick-a-dee-dee-dee which gave this bird its name and its song is fee-bee-fee-bay.


Their breeding habitat is mixed or deciduous woods in the United States from New Jersey west to southern Kansas and south to Florida and Texas; there is a gap in the range at high altitudes in the Appalachian Mountains where they are replaced by their otherwise more northern relative, the black-capped chickadee.
They are permanent residents, not usually moving south even in severe winter weather.


These birds hop along tree branches searching for insects, sometimes hanging upside down or hovering; they may make short flights to catch insects in the air. Insects form a large part of their diet, especially in summer; seeds and berries become important in winter. They sometimes hammer seeds on a tree or shrub to open them; they also will store seeds for later use. Carolina chickadees commonly make use of feeders.


During the fall migration and winter, chickadees often flock together in flocks of 8–10 birds. Each of the birds in a flock has a rank; while highly ranked birds will remain on the flock's territory for breeding, lower-ranked birds must find new breeding territory. Many other species of birds, including titmice, nuthatches, and warblers can often be found foraging in these flocks. Mixed flocks occur about 50% of the time. Mixed flocks stay together because the chickadees call out whenever they find a good source of food. This calling out forms cohesion for the group, allowing the other birds to find food more efficiently.
Carolina chickadees actively defend individual spaces of 2–5 ft (0.6–1.5 m) apart; if another bird encroaches on these spaces, the dominant bird may make gargle calls. At feeders, these birds will usually take a seed and fly to a branch isolated from other birds to eat it.
These birds usually sleep in cavities, though they may sleep in branches. Different members of a flock will sleep in the same cavity from night to night. While females usually sleep in the nest cavity, males will normally sleep on a nearby branch.


They nest in a hole in a tree; the pair excavates the nest, using a natural cavity or sometimes an old woodpecker nest. They may interbreed with black-capped chickadees where the ranges overlap, which can make identification difficult. The female will build the nest out of moss and strips of bark; she will then line it with hair or plant fibers. Clutches are usually made up of 3–10 eggs with an incubation period of 12–16 days. The nestling period is usually 16–19 days.


Carolina chickadees are able to lower their body temperatures to induce an intentional state of hypothermia called torpor. They do this to conserve energy during extremely cold winters. In extremely cold weather conditions they look for cavities where they can hide in and spend up to fifteen hours at a time in torpor; during this time they are awake but unresponsive; they should not be picked up and handled at this time, as the stress of being held may cause their death.


Todd M Freeberg (2006) Social Complexity Can Drive Vocal Complexity: Group Size Influences Vocal Information in Carolina Chickadees. DOI 10.1111/j.1467-9280.2006.01743


The willow tit (Poecile montanus) is a passerine bird in the tit family, Paridae. It is a widespread and common resident breeder throughout temperate and subarctic Europe and across the Palearctic. The plumage is grey-brown and off-white with a black cap and bib. It is more of a conifer specialist than the closely related marsh tit, which explains it breeding much further north. It is resident, and most birds do not migrate.


The willow tit was described in 1827 by the Swiss naturalist Thomas Conrad von Baldenstein under the trinomial name Parus cinereus montanus. The type locality is the mountain forests in the Canton of Grisons, Switzerland. The willow tit is now placed in the genus Poecile that was erected by the German naturalist Johann Jakob Kaup in 1829. The genus name, Poecile, is the Ancient Greek name for a now unidentifiable small bird, and the specific montanus is Latin for "of the mountains".
Poecile was at one time treated as a subgenus within the genus Parus but molecular taxonomic analysis, using both nuclear and mitochondrial genes, supports Poecile as a distinct clade. Within Poecile, most of the Old World species (including the willow tit) form a separate clade from the New World chickadees. The taxonomic analysis has shown that the willow tit is sister to the Caspian tit (Poecile hyrcanus).
The willow tit was formerly considered conspecific with the black-capped chickadee of North America due to their very similar appearance. This is seen in an older version of the Peterson Field Guide, Birds of Britain and Europe. Underneath the name it states; "N Am. Black-Capped Chickadee" as an alternate name. In fact the willow tit, black-capped chickadee, marsh tit and Carolina chickadee are all very similar to one another in appearance.
There are 14 recognised subspecies:
P. m. kleinschmidti (Hellmayr, 1900) – Britain
P. m. rhenanus (O. Kleinschmidt, 1900) – northwest France to west Germany, north Switzerland and north Italy
P. m. montanus (Conrad von Baldenstein, 1827) – southeast France to Romania, Bulgaria and Greece
P. m. salicarius (C. L. Brehm, 1831) – Germany and west Poland to northeast Switzerland and Austria
P. m. borealis (de Sélys-Longchamps, 1843) – Scandinavia south to Ukraine
P. m. uralensis (Grote, 1927) – southeast European Russia, west Siberia and Kazakhstan
P. m. baicalensis R. Swinhoe, 1871 east central and east Siberia, north Mongolia, north China and north Korea
P. m. anadyrensis (Belopolski, 1932) northeast Siberia
P. m. kamtschatkensis Bonaparte, 1850 – Kamchatka Peninsula and north Kuril Islands
P. m. sachalinensis (Lönnberg, 1908) – Sakhalin Island and south Kuril Islands
P. m. restrictus (Hellmayr, 1900) – Japan
P. m. songarus (Severtsov, 1873) – southeast Kazakhstan to Kyrgyzstan and northwest China
P. m. affinis Przewalski, 1876 – north central China
P. m. stoetzneri (O. Kleinschmidt, 1921) – northeast China
The Sichuan tit (Poecile weigoldicus) was formerly treated as a subspecies of the willow tit. It was promoted to species status based on a 2002 phylogenetic analysis that compared DNA sequences from the mitochondrial cytochrome-b gene. The single locus results were later confirmed by a larger multi-locus analysis published in 2017.


The willow tit is 11.5 cm (4.5 in) in length, has a wingspan of 17–20.5 cm (6.7–8.1 in) and weighs around 11 g (0.39 oz).
In the east of its range it is much paler than marsh tit, but as one goes west the various races become increasingly similar, so much so that it was not recognised as a breeding bird in Great Britain until the end of the 19th century, despite being widespread.
The willow tit is distinguished from the marsh tit by a sooty brown instead of a glossy blue black cap; the general colour is otherwise similar, though the under parts are more buff and the flanks distinctly more rufous; the pale buff edgings to the secondaries form a light patch on the closed wing. The feathers of the crown and the black bib under the bill are longer, but this is not an easily noticed character. However, the more graduated tail (not square) shows distinctly when spread.
The commonest call is a nasal zee, zee, zee, but the notes of the bird evidently vary considerably. Occasionally a double note, ipsee, ipsee, is repeated four or five times.


The willow tit excavates its own nesting hole, even piercing hard bark; this is usually in a rotten stump or in a tree, more or less decayed. Most nests examined are cups of felted material, such as fur, hair and wood chips, but feathers are sometimes used. The number of eggs varies from six to nine, with reddish spots or blotches.
In a study using ring-recovery data carried out in northern Finland, the survival rate for juveniles for their first year was 0.58, and the subsequent adult annual survival rate was 0.64. For birds that survive the first year the typical lifespan is thus only three years. The maximum recorded age is 11 years; this has been recorded for a bird in Finland and for another near Nottingham in England.


The willow tit has an extremely large range with an estimated population of between 175 and 253 million mature individuals. This large population appears to be slowly decreasing but the decline is not sufficiently rapid to approach the threshold of vulnerability. The species is therefore classed as of least-concern by the International Union for Conservation of Nature. In contrast, the number in the United Kingdom declined by 83% between 1995 and 2017. There was also a contraction in the range. The rapid decline is believed be due to three factors: habitat loss, competition for nest holes by other tits particularly blue tits, and nest predation by the great spotted woodpecker. Over the same period, the number of great spotted woodpeckers increased fourfold.


Cramp, Stanley; Perrins, C.M., eds. (1993). "Parus montanus Willow Tit". Handbook of the Birds of Europe the Middle East and North Africa. The Birds of the Western Palearctic. Vol. VII: Flycatchers to Strikes. Oxford: Oxford University Press. pp. 168–186. ISBN 978-0-19-857510-8.


The marsh tit (Poecile palustris) is a passerine bird in the tit family Paridae and genus Poecile, closely related to the willow, Père David's and Songar tits. It is a small bird, around 12 cm (4.7 in) long and weighing 12 g (0.42 oz), with a black crown and nape, pale cheeks, brown back and greyish-brown wings and tail. Between 8 and 11 subspecies are recognised. Its close resemblance to the willow tit can cause identification problems, especially in the United Kingdom where the local subspecies of the two are very similar: they were not recognised as separate species until 1897.
Globally, the marsh tit is classified as Least Concern, although there is evidence of a decline in numbers (in the UK, numbers have dropped by more than 50% since the 1970s, for example). It can be found throughout temperate Europe and northern Asia and, despite its name, it occurs in a range of habitats including dry woodland. The marsh tit is omnivorous; its food includes caterpillars, spiders and seeds. It nests in tree holes, choosing existing hollows to enlarge, rather than excavating its own. A clutch of 5–9 eggs is laid.


The marsh tit was formally described by the Swedish naturalist Carl Linnaeus in 1758 in the tenth edition of his Systema Naturae under the binomial name Parus palustris. It is now placed in the genus Poecile that was erected by the German naturalist Johann Jakob Kaup in 1829. The genus name, Poecile, is the Ancient Greek name for a now unidentifiable small bird, and is perhaps derived from poikolos, meaning "spotted"; the specific palustris is Latin for "marshy".
An analysis of mitochrondrial DNA sequences has shown that the marsh tit belongs to a group of Eurasian 'typical chickadees' that includes the willow tit, Père David's tit, the black-bibbed tit, and the Songar tit.
Ten subspecies are recognised:
P. p. dresseri (Stejneger, 1886) – central and southern England, Wales] and west France. Still present in the far south-east of Scotland but in small numbers. Slightly smaller than palustris and more brown or olive-coloured on upperparts and dirtier below. Pale brown tertial fringes can sometimes cause confusion with willow tit
P. p. palustris (Linnaeus, 1758) – north and central Europe, from south Scandinavia south to north Iberia, east to central Poland, west Balkans and Greece
P. p. italicus (Tschusi & Hellmayr, 1900) – French Alps, Italy and Sicily. As dresseri, but more rufous on upperparts and paler on underparts In Sicily, it was described the subspecies siculus (De Burg, 1925) considered synonymic with italicus by Vaurie (1959), and subsequently forgotten by most authors. However, in a typical cline pale-dark/north-south all over Europe, and along the Italian peninsula too, where darkest birds are in the Alpine area and palest (as pale as nominate subsp.) in Calabria, the Sicilian very isolated and relict population is composed by the smallest and darkest birds in Europe, therefore rather out of the clinal gradient/intergrade populations. It should deserve better and in depth study before being in synonymized (Priolo, 1969; Corso, 2005; A.Corso, pers. obs.)
P. p. stagnatilis (Brehm, CL, 1855) – east Europe east to south Urals and northwest Turkey
P. p. kabardensis (Buturlin, 1929) – Caucasus and northeast Turkey
P. p. brevirostris Taczanowski, 1872 – south central & southeast Siberia, north Mongolia, northeast China and North Korea
P. p. ernsti (Yamashina, 1933) – Sakhalin Island
P. p. hensoni (Stejneger, 1892) – south Kuril Islands and Hokkaido, Japan
P. p. jeholicus (Kleinschmidt, O & Weigold, 1922) – northeast China and North Korea
P. p. hellmayri Bianchi, 1903 – east China and South Korea


The nominate race has a black cap and nape with a blue sheen visible at close quarters. The black 'bib' below the bill is rather small; the cheeks are white, turning dusky brown on the ear coverts. The upperparts, tail and wings are greyish-brown, with slightly paler fringes to the tertials. The underparts are off-white with a buff or brown tinge strongest on the flanks and undertail coverts. The bill is black and the legs dark grey. Juveniles are very similar to adults, but with a duller black cap and bib, more greyish upperparts and paler underparts; they moult into adult plumage by September.
The marsh tit weighs 12 g (0.42 oz), has a length of 11.5 to 12 cm (4.5 to 4.7 in) (from bill to tail) and a wingspan of 19 cm (7.5 in). The wing length ranges from 60–70 mm (2.4–2.8 in). The oldest recorded marsh tit in Europe reached the age of 11 years, 11 months.
Like other tits it has a large range of call notes; most typical is the explosive "pitchou" note, given when agitated, often leading into "pitchou-bee-bee-bee", which can sound like willow tit when not heard clearly. Unlike many other tits, however, the marsh tit has a well-defined song and a wide song repertoire. Individual birds can have more than five songs, which they use interchangeably. Some of the more common songs include a typical tit-like, ringing, "schip-schip-schip-schip-ship", a more liquid "tu-tu-tu-tu-tu" and sometimes a sweet "tyeu-tyeu-tyeu-tyeu-tyeu". The old Staffordshire name for the species, 'Saw Whetter', refers to the bird's scolding call.
Marsh and willow tits are difficult to distinguish on appearance alone; the races occurring in the UK (P. p. dresseri and P. m. kleinschmidti respectively) are especially hard to separate. When caught for ringing, the pale 'cutting edge' of the marsh tit's bill is a reliable criterion; otherwise, the best way to tell apart the two species is by voice. Plumage characteristics include the lack of a pale wing panel (formed by pale edges to the secondary feathers in the willow tit), the marsh tit's glossier black cap and smaller black 'bib', although none of these is 'completely reliable'; for example, juvenile marsh tits can show a pale wing panel. The marsh tit has a noticeably smaller and shorter head than the willow tit and overall the markings are crisp and neat, with the head in proportion to the rest of the bird (willow tit gives the impression of being 'bull-necked').
A measure of the difficulty in identification is given by the fact that, in the UK, the willow tit was not identified as distinct from marsh tit until 1897. Two German ornithologists, Ernst Hartert and Otto Kleinschmidt, were studying marsh tit skins at the British Museum and found two wrongly-labelled willow tits amongst them (two willow tit specimens were then collected at Coalfall Wood in Finchley, north London, and that species was added to the British list in 1900).


The marsh tit has a worldwide Extent of Occurrence of around 10 million square kilometres. The global population includes between 6.1 million and 12 million birds in Europe alone. The species is classified as Least Concern, though there is some evidence of a decline in numbers. For example, between the 1970s and 2007, marsh tit numbers declined in the United Kingdom by more than 50% and consequently it is on the Red List of species compiled by the UK's Joint Nature Conservation Committee. Research suggests that the cause may be low survival rates from year to year, though the nest failure rate has fallen during the decline. Other studies have shown that reduced diversity of woodland structure and plant species, partly because of the impact of deer browsing, is the cause of the bird's decline.
Data from 157 woodlands covered by the RSPB/BTO Repeat Woodland Bird Survey showed that the abundance of marsh tits in 2003–2004 corresponded with the vegetation present 2–4 metres above the ground, the shrub layer. Data from the 1980s did not show the same results, but marsh tit numbers had increased by 2003–2004 in woods with the most shrub cover. The study concluded that damage to the shrub layer, caused by overgrazing by deer, for example, may make woodland less suitable for marsh tits.
It is a widespread and common resident breeder throughout temperate Europe and northern Asia. It occurs from northern Spain north to south-eastern Scotland and east to western Russia, with a broad gap in western Asia and present again in eastern Asia from the Altai Mountains east to northern Japan and northern and western China.
This species is sedentary, making short post-breeding movements in most of its range, but in northern Europe some move southward in winter. However, marsh tits seem not to perform the occasional irruptions that other members of the tit family do. Most marsh tits stay in their breeding territories year-round; presumably this is related to their food-storing strategy. Analysis of UK ringing data showed that of 108 recoveries (when a ringed bird is found dead or caught by another ringer), 85% were less than 5 km from where the bird was originally caught, and only 1% further than 20 km. Young birds join mixed roaming flocks; adults also join the flocks when they pass through, but do not stray from their territory.
Marsh tits breed mostly in lowland areas, but can reach altitudes of up to 1,300 m. They prefer large areas of moist, broadleaved woodland, often oak or beech, though they can occupy wet alder woodland, riverside trees, parks and gardens or orchards. A study at Monks Wood, Cambridgeshire, England, found that marsh tits required mature trees with a shrub layer below the canopy, but that they avoided parts of the wood with many young trees. Another study in the United Kingdom found that during winter and while foraging, marsh tits spent more time than blue tits in the wood's understorey, and more time lower in the woodland canopy and understorey. Trees and shrubs in 10 breeding territories were also compared. The trees varied significantly between territories, but the shrub characteristics did not, suggesting that the shrubs were more important to the birds. In Wytham Woods, Oxfordshire, marsh tits were "largely absent" from parts of the wood with a dense canopy but poor shrub cover.


In mixed winter tit flocks, seldom more than one or two marsh tits are present, and parties of this species alone are infrequent. Its performances in the bushes and branches are just as neat and agile as those of other tits; it often hangs upside down by one leg.


Marsh tits are monogamous and often pair for life; one pair stayed together for six years. In Europe, hybridisation with the willow tit has been recorded twice.
The nest site is in a hole, usually in a tree but sometimes in a wall or in the ground. Nestboxes may be used. Old willow tit holes may be used and enlarged further. Marsh tits do not usually excavate their own nest holes, though they may enlarge the hollow, carrying the chips to a distance before dropping them. The hole may be within a centimetres or two of the ground or high as 10 m (33 ft). Inside the hole, a nest of moss is made and lined with hair and sometimes a few feathers; 20 cm (7.9 in) of moss is used in damp holes, but much less in dry ones.
Between five and nine white and red-speckled eggs are laid late in April or in May, measuring 16 mm × 12 mm (0.63 in × 0.47 in) and weighing 1.2 g (0.042 oz) each, of which 6% is shell. The eggs are incubated by the female for 14–16 days; incubation begins before the clutch is complete, meaning that the chicks hatch over a period of around two days. She sits closely and gives a typical tit "hissing display" if disturbed. The male helps to feed and care for the young and brings nearly all the food for the first four days after hatching. The altricial, downy chicks fledge after 18–21 days. The fledglings are fed by their parents for a week and become independent after a further 1–7 days. The family stays together for between 11 and 15 days after the first flights of the juveniles. Second broods have been recorded, though they are extremely rare in Britain; most are replacement clutches.
A study of marsh tits in Białowieża Forest, Poland, looked at which factors affected birds' choice to re-occupy a nest hole used the previous year. It found that 35.5% of available holes were used again the following year, and that holes where a brood had failed were less likely to be re-used. A study in Bourton Woods, Gloucestershire, England, found that nestboxes were used by marsh tits in successive years in only 20% of cases.


Mostly spiders and insects are eaten in spring and summer, but seeds – including those of the thistle – nuts and berries are taken in autumn and winter. Beechmast is the preferred food when it can be found. Marsh tits often take seeds and fruit from the plant before taking them to eat elsewhere.
Marsh tits collect and store large numbers of seeds. For a study in Norway, birds were watched for two hours 15 minutes. They ate 43 and cached 83 seeds per hour. In Sweden, storing food is most frequent between September and late February, with the peak in this behaviour occurring from September to October. Hiding places for the seeds include on and in the ground, in leaf litter, in tree stumps, and under moss and lichen in trees. The hidden seeds are prone to being stolen, by other marsh tits or other species, so birds often fly from one site to another before deciding on a hiding place. They tend to retrieve the oldest items first, and memorise their location rather than searching randomly or checking systematically. The marsh tit's hippocampus is 31% bigger than that of the great tit, despite the great tit's larger overall size and larger forebrain; the relative volume of this part of the brain is greater in birds that cache food.


The chestnut-backed chickadee (Poecile rufescens, formerly Parus rufescens) is a small passerine bird in the tit family, Paridae.
It is found in the Pacific Northwest of the United States and western Canada, from southern Alaska to southwestern California. It is a permanent resident within its range, with some seasonal movements as feeding flocks move short distances in search of food. They usually move to lower elevations in the same area upon onset of winter and move back up to higher elevations in late summer.
Chickadees are able to use nocturnal hypothermia to regulate energy expenditure, allowing them to survive harsh winters where other bird species not utilizing thermal regulation would not be able to. Some estimates put the energy conserved while using nocturnal hypothermia all the way up to 32%.
It is a small chickadee, 11.5–12.5 cm (4.5–4.9 in) long with a weight of 8.5–12.6 g (0.30–0.44 oz). The head is dark blackish-brown with white cheeks, the mantle is bright rufous-brown, the wing feathers are dark gray with paler fringes. The underparts are white to pale grayish-white, with rufous or pale gray flanks. It is often considered the most handsome of all chickadees. They often move through the forest in mixed feeding flocks, and are often seen in large groups with bushtits and warblers.
There are three subspecies, with the flanks being grayer and less rufous further south:
Poecile rufescens rufescens (Townsend, 1837). Alaska south to northwest California. Broad rufous band on flanks.
Poecile rufescens neglectus (Ridgway, 1879). Coastal central California (Marin County). Narrow rufous band on flanks.
Poecile rufescens barlowi (Grinnell, 1900). Coastal southwestern California (south of San Francisco Bay). Almost no rufous color on flanks.
Its habitat is low elevation coniferous and mixed coniferous forests. In the San Francisco Bay Area this bird has readily adapted to suburban settings, prompting a range expansion. It is a cavity-nester, usually utilizing an abandoned woodpecker hole, but sometimes excavating on its own. Chestnut-backed chickadees use much fur and hair to make their nests. Their nests are actually 50% fur and hair. The most common hair they use comes from deer, rabbits, and coyotes. The adult chickadees also make a layer of fur about a centimeter thick which is used to cover the eggs on the nest whenever they leave the nest. It lays 5–8 (sometimes 9) eggs per clutch.
Its food is largely insects and other invertebrates gleaned from foliage. Chestnut-backed chickadees take some seeds, especially those of conifers, and fruit. It will visit bird feeders, including hummingbird feeders, and especially loves suet.
Molecular phylogenetic studies have shown that the chestnut-backed chickadee is sister to the boreal chickadee (Poecile hudsonicus).


Pogonomyrmex barbatus is a species of harvester ant from the genus Pogonomyrmex. Its common names include red ant and red harvester ant. These large (5– to 7-mm) ants prefer arid chaparral habitats and are native to the Southwestern United States. Nests are made underground (up to 2.5 m deep) in exposed areas. Their diets consist primarily of seeds, and they consequently participate in myrmecochory, an ant-plant interaction through which the ants gain nutrients and the plants benefit through seed dispersal. Red harvester ants are often mistaken for fire ants, but are not closely related to any fire ant species, native or introduced.


Red harvester ant nests are characterized by a lack of plant growth and small pebbles surrounding the entrance to the tunnel, which usually descends at a pronounced angle. Hulls of seeds may be found scattered around the nest. In grassland areas, such as ranches, the lack of plant life makes red harvester ant colonies very easy to spot, and where they are very plentiful, they may make serious inroads into the grazing available to livestock.
The mounds are typically flat and broad, 0 to 100 mm (0.0 to 3.9 in) high, and 300 to 1,200 mm (12 to 47 in) in diameter. Even larger denuded areas have been reported, on the order of 10 m² (110 sq ft). Three to eight trails typically lead away from the mound, like "arms". These trails are used by ants to collect and bring food back to the mound. "Scout" ants are the first ones out of the mound every morning. They seek food, and mark their path as they return to the mound to alert the worker ants. The worker ants follow the scent trail and collect the food. Other worker ants clean, extend, and generally tend to the mound, the queen, and the brood. All the ants in the colonies are females apart from the winged males produced in the breeding season.


The main food source for red harvester ants usually consists of seeds, which they hoard in great numbers. The food is first ground to a bread-like consistency using the ants' large mandibles, and is then stored in a granary, assuring the colony access to food year-round. Seed collection on behalf of the red harvester ants benefits their ecosystem through the process of myrmecochory, in which ants aid in the dispersal of seeds while foraging for food. Both plants and ants benefit from this relationship: the plants increase their dispersal range and density, while the ants benefit from acquiring nutrients and ensuring a more secure food supply in future harvests. This is typically understood as a mutualistic interaction. Dead insects are also collected during foraging.