The impala is diurnal (active mainly during the day), though activity tends to cease during the hot midday hours; they feed and rest at night. Three distinct social groups can be observed – the territorial males, bachelor herds and female herds. The territorial males hold territories where they may form harems of females; territories are demarcated with urine and faeces and defended against juvenile or male intruders. Bachelor herds tend to be small, with less than 30 members. Individuals maintain distances of 2.5–3 metres (8.2–9.8 ft) from one another; while young and old males may interact, middle-aged males generally avoid one another except to spar. Female herds vary in size from 6 to 100; herds occupy home ranges of 80–180 hectares (200–440 acres; 0.31–0.69 sq mi). The mother–calf bond is weak, and breaks soon after weaning; juveniles leave the herds of their mothers to join other herds. Female herds tend to be loose and have no obvious leadership. Allogrooming is an important means of social interaction in bachelor and female herds; in fact, the impala appears to be the only ungulate to display self-grooming as well as allogrooming. In allogrooming, females typically groom related impalas, while males associate with unrelated ones. Each partner grooms the other six to twelve times.
Social behaviour is influenced by the climate and geography; as such, the impala are territorial at certain times of the year and gregarious at other times, and the length of these periods can vary broadly among populations. For instance, populations in southern Africa display territorial behaviour only during the few months of the rut, whereas in eastern African populations, territoriality is relatively minimal despite a protracted mating season. Moreover, territorial males often tolerate bachelors, and may even alternate between bachelorhood and territoriality at different times of the year. A study of impala in the Serengeti National Park showed that in 94% of the males, territoriality was observed for less than four months.
The impala is an important prey species for several carnivores, such as cheetahs, leopards and lions. The antelope displays two characteristic leaps – it can jump up to 3 metres (9.8 ft), over vegetation and even other impala, covering distances of up to 10 metres (33 ft); the other type of leap involves a series of jumps in which the animal lands on its forelegs, moves its hindlegs mid-air in a kicking fashion, lands on all fours (stotting) and then rebounds. It leaps in either manner in different directions, probably to confuse predators. At times, the impala may also conceal itself in vegetation to escape the eye of the predator. The most prominent vocalisation is the loud roar, delivered through one to three loud snorts with the mouth closed, followed by two to ten deep grunts with the mouth open and the chin and tail raised; a typical roar can be heard up to 2 kilometres (1.2 mi) away. Scent gland secretions identify a territorial male. Impalas are sedentary; adult and middle-aged males, in particular, can hold their territories for years.


Common ixodid ticks collected from impala include Amblyomma hebraeum, Boophilus decoloratus, Hyalomma marginatum, Ixodes cavipalpus, Rhipicephalus appendiculatus and R. evertsi. In Zimbabwe, heavy infestation by ticks such as R. appendiculatus has proved to be a major cause behind the high mortality of ungulates, as they can lead to tick paralysis. Impala have special adaptations for grooming, such as their characteristic dental arrangement, to manage ticks before they engorge; however, the extensive grooming needed to keep the tick load under control involves the risk of dehydration during summer, lower vigilance against predators and gradual wearing out of the teeth. A study showed that impala adjust the time devoted to grooming and the number of grooming bouts according to the seasonal prevalence of ticks.
Impala are symbiotically related to oxpeckers, which feed on ticks from those parts of the antelope's body which the animal cannot access by itself (such as the ears, neck, eyelids, forehead and underbelly). The impala is the smallest ungulate with which oxpeckers are associated. In a study it was observed that oxpeckers selectively attended to impala despite the presence of other animals such as Coke's hartebeest, Grant's gazelle, Thomson's gazelle and topi. A possible explanation for this could be that because the impala inhabits woodlands (which can have a high density of ticks), the impala could have greater mass of ticks per unit area of the body surface. Another study showed that the oxpeckers prefer the ears over other parts of the body, probably because these parts show maximum tick infestation. The bird has also been observed to perch on the udders of a female and pilfer its milk.
Lice recorded from impala include Damalinia aepycerus, D. elongata, Linognathus aepycerus and L. nevilli; in a study, ivermectin (a medication against parasites) was found to have an effect on Boophilus decoloratus and Linognathus species, though not on Damalinia species. In a study of impala in South Africa, the number of worms in juveniles showed an increase with age, reaching a peak when impala turned a year old. This study recorded worms of genera such as Cooperia, Cooperoides, Fasciola, Gongylonema. Haemonchus, Impalaia, Longistrongylus and Trichostrongylus; some of these showed seasonal variations in density.
Imapala show high frequency of defensive behaviours towards flying insects. This is probably the reason for Vale 1977 and Clausen et al 1998 only finding trace levels of feeding by Glossina (tsetse fly) upon impala.
Theileria of impala in Kenya are not cross infectious to cattle: Grootenhuis et al 1975 were not able to induce cattle infection and Fawcett et al 1987 did not find it naturally occurring.


Impala browse as well as graze; either may predominate, depending upon the availability of resources. The diet comprises monocots, dicots, forbs, fruits and acacia pods (whenever available). Impala prefer places close to water sources, and resort to succulent vegetation if water is scarce. An analysis showed that the diet of impala is composed of 45% monocots, 45% dicots and 10% fruits; the proportion of grasses in the diet increases significantly (to as high as 90%) after the first rains, but declines in the dry season. Browsing predominates in the late wet and dry season, and diets are nutritionally poor in the mid-dry season, when impala feed mostly on woody dicots. Another study showed that the dicot proportion in the diet is much higher in bachelors and females than in territorial males.
Impala feed on soft and nutritious grasses such as Digitaria macroblephara; tough, tall grasses, such as Heteropogon contortus and Themeda triandra, are typically avoided. Impala on the periphery of the herds are generally more vigilant against predators than those feeding in the centre; a foraging individual will try to defend the patch it is feeding on by lowering its head. A study revealed that time spent in foraging reaches a maximum of 75.5% of the day in the late dry season, decreases through the rainy season, and is minimal in the early dry season (57.8%).


The impala inhabits woodlands due to its preference for shade; it can also be found on the interface (ecotone) between woodlands and savannahs. Places near water sources are preferred. In southern Africa, populations tend to be associated with Colophospermum mopane and Acacia woodlands. Habitat choices differ seasonally – Acacia senegal woodlands are preferred in the wet season, and A. drepanolobium savannahs in the dry season. Another factor that could influence habitat choice is vulnerability to predators; impala tend to keep away from areas with tall grasses as predators could be concealed there. A study found that the reduction of woodland cover and creation of shrublands by the African bush elephants has favoured impala population by increasing the availability of more dry season browse. Earlier, the Baikiaea woodland, which has now declined due to elephants, provided minimum browsing for impala. The newly formed Capparis shrubland, on the other hand, could be a key browsing habitat. Impala are generally not associated with montane habitats; however, in KwaZulu-Natal, impala have been recorded at altitudes of up to 1,400 metres (4,600 ft) above sea level.
The historical range of the impala – spanning across southern and eastern Africa – has remained intact to a great extent, although it has disappeared from a few places, such as Burundi. The range extends from central and southern Kenya and northeastern Uganda in the east to northern KwaZulu-Natal in the south, and westward up to Namibia and southern Angola. The black-faced impala is confined to southwestern Angola and Kaokoland in northwestern Namibia; the status of this subspecies has not been monitored since the 2000s. The common impala has a wider distribution, and has been introduced in protected areas in Gabon and across southern Africa.


The International Union for Conservation of Nature and Natural Resources (IUCN) classifies the impala as a species of least concern overall. The black-faced impala, however, is classified as a vulnerable species; as of 2008, fewer than 1,000 were estimated in the wild. Though there are no major threats to the survival of the common impala, poaching and natural calamities have significantly contributed to the decline of the black-faced impala. As of 2008, the population of the common impala has been estimated at around two million. According to some studies, translocation of the black-faced impala can be highly beneficial in its conservation.
Around a quarter of the common impala populations occur in protected areas, such as the Okavango Delta (Botswana); Masai Mara and Kajiado (Kenya); Kruger National Park (South Africa); the Ruaha and Serengeti National Parks and Selous Game Reserve (Tanzania); Luangwa Valley (Zambia); Hwange, Sebungwe and Zambezi Valley (Zimbabwe). The rare black-faced impala has been introduced into private farms in Namibia and the Etosha National Park. Population densities vary largely from place to place; from less than one impala per square kilometre in Mkomazi National Park (Tanzania) to as high as 135 per square kilometre near Lake Kariba (Zimbabwe).


The white-throated swift (Aeronautes saxatalis) is a swift of the family Apodidae native to western North America, south to cordilleran western Honduras. Its coastal range extends as far north as Northern California, while inland it has migratory populations found throughout the Great Basin and Rocky Mountain regions, ranging as far north as southern British Columbia. White-throated swifts are found in open areas near cliffs, rock faces, or man-made structures, where they roost. Swifts are social birds, and groups are often seen roosting and foraging for flying insects together.


The white-throated swift is medium-sized bird that is primarily black and white. Its back and wings are blackish-brown or black, and its breast, belly, chin, and throat are white. In newly molted adults, the dark feathers may have a slight greenish sheen, though this is rarely observable in the field. Sexes cannot be distinguished by plumage. Juvenile (first basic) plumage is similar to adult (definitive basic) plumage, but with the feathers on juveniles' heads and necks appearing duller and paler.
Adults are generally 15–18 centimeters in length, and have a tail with a relatively shallow 10 millimeter-deep notch. Adult swifts weigh between 28 and 36 grams, with a mean weight recorded as 32.5 grams, and there is no difference in size between males and females. The white-throated swift has a wingspan of 35.5 cm. White-throated swifts can appear similar to American black swifts and Vaux's swifts, which also occur in western North America, but they can be distinguished by their white underparts which both other species lack.


White-throated swifts are year-round residents of coastal and southern California, southern Arizona, southern New Mexico, western Texas, central Mexico, and the Sierra Madre Occidental and Oriental mountains. Migratory breeding populations can be found from Arizona and New Mexico north to southern British Columbia, and from central California east to eastern Colorado and Wyoming. In the southern part of its migratory range, birds arrive in mid-March and depart in mid-October, while in the northern portion, they do not arrive until mid-April to early-May, and depart in late-August to mid-September. There are two main areas in which non-breeding wintering populations can be found: the first includes southwestern California, eastern Arizona, the Texas Panhandle, while the second is in El Salvador, Guatemala, Honduras, and the Mexican state of Chiapas.


Swifts roost in cliffs, bluffs, and canyons, as well as man-made structures like bridges, overpasses, and walls of quarries. In natural settings, swifts form colonies of as many as 400–500 birds in protected cliff crevices that are generally 6–50 meters above the ground. White-throated swifts will sometimes roost and nest in amongst colonies of swallows, including cliff, violet-green, and northern rough-winged swallows. While foraging, white-throated swifts are found flying over meadows, agricultural fields, and open areas along the edges of ridges and hills.


Swifts are aerial insectivores who frequently forage in areas of rising air along the edges of canyons, foothills, or mountains to capture insects. White-throated swifts have also been observed foraging over agricultural lands, and are known to follow harvesting machinery to capture insects disturbed by the equipment. Stomach content analysis has found that beetles, flies, bees, and true bugs (Hemiptera) constitute a majority of the white-throated swift's diet. Adult swifts nourish their nestlings by carrying a bolus of arthropods in their buccal cavity, and feeding it to the young. Swifts derive most of their water from their diet, but can also be skimming the surface of ponds to drink.
Known predators of White-throated swifts include peregrine falcons, prairie falcons, and American kestrels. When predators are detected, swifts are known to quickly lower their altitude and fly just above the vegetation. There is no data available on levels of nest predation.


During the courtship season, white-throated swifts may perform gliding displays and plummeting courtship falls prior to forming monogamous pairs. Once a pair is established, nests made of plant material and feathers held together with saliva are built on rocky cliff faces or human-made structures. These nests are usually 8–10 centimeters across, and 2.5–2.8 centimeters deep, and may be used for several years in a row. An average of 4–5 white matte eggs are laid over the course of 4–6 days, with incubation generally beginning only after the last egg is laid. In southern parts of their range, egg-laying occurs in April, but in more northerly areas, clutches are usually laid throughout May, with a median first-egg date of May 13. The incubation period lasts on average 24 days, with swifts weighing approximately 2 grams upon hatching. Both parents feed the nestlings, and by the time of fledging, swifts may weigh up to 46 grams. There are no documented cases of swifts attempting a second brood if the first nest fails.


Across North America, white-throated swift populations have declined by 2.8% from 1966–1998, which is considered to be a significant decline. Such declines could be related to loss of roosting and nesting sites from quarrying, mining, and the demolition of older buildings, as well as reduction in food supply due to pesticide use. However, in certain areas swift populations appear to have increasing trends, and most populations appear stable. The patchy distribution of this species makes interpreting these data difficult, but the apparent overall decline is of concern given the concurrent declines of other aerial insectivores.


The Canada darner was first scientifically described in 1908 by Canadian entomologist Edmund Murton Walker, in the same publication as the first descriptions of black-tipped darner, shadow darner, subarctic darner, and variable darner, .


Canada darners spend most of their lives as an aquatic nymph. Larvae are typically found in lakes and ponds, especially bog ponds and beaver ponds. Breeding ponds typically have abundant emergent vegetation. An Ontario study found that the species was restricted to ponds with a pH of no more than 6.2. One study found that larvae may spend either one or two winters underwater before emerging.
Adult males fly slowly along the shores of breeding ponds, sometimes dropping to the surface to search for females. Mating occurs in shrubs nearby. Females lay eggs at water level, usually in marshy vegetation away from shore. Breeding adults are most active in sunny weather.
Away from breeding sites, adult Canada darners are often found with other darner species in feeding swarms in clearings and along roadways in early evening. Adult Canada darners may be seen from June to October.
Canada darners are often considered to be a common to abundant species where they are found. Although the species has not generally been recognised as migratory, a study in Manitoba and Minnesota found that at least some populations of Canada darners undertake north-south migratory movements within their range.


Flight period of these insects lasts from June to October, with some specimen visible in May and November. The adults of the southern hawker  feed on  various insects, caught on the wing.  This is an inquisitive species and will approach people.
These dragonflies breed in still or slow-flowing water. The males are often seen patrolling by a ponds edge or river, where they fight away intruders, crashing into rival males and spiralling through the air. The females are quite inconspicuous when they lay their eggs, but they sometimes give away their spot by clattering up from the reeds.
The eggs are laid by jabbing the abdomen into rotting vegetation or wood. The eggs hatch in the spring, after being laid in the previous summer or autumn. The nymphs feed on aquatic insects, small tadpoles, invertebrates and small fish. They emerge as adults in July and August after two-three years’ development.


The variable darner (Aeshna interrupta) is a dragonfly of the family Aeshnidae, native from Alaska through the Northwest Territories to Newfoundland, south to New Hampshire and Michigan in the eastern United States, and to the mountains of New Mexico, Arizona, and California in the west. It's named after the distinctive broken stripes on the sides of the thorax of many male specimens. It lives in many habitats from northern and mountain peatlands to cattail marshes and temporary pools. It is the characteristic species of grassland ponds.


The common hawker, moorland hawker or sedge darner (Aeshna juncea) is one of the larger species of hawker dragonflies. It is native to Palearctic (from Ireland to Japan) and northern North America. The flight period is from June to early October.
It is 74 millimetres (2.9 in) long with a brown body. The male has a black abdomen with paired blue and yellow spots on each abdominal segment, and narrow stripes along the dorsal surface of the thorax. In the female, the abdomen is brown with yellow or sometimes green or blue spots. The wings of both sexes display a yellow costa (the major vein running along the leading edge of the wings). This species lacks the green thorax stripes of the southern hawker.
Female common hawkers will sometimes dive out of the sky and feign death in order to avoid copulating with males.


The shadow darner patrols along small marshy streams. It is often found feeding along woodland edges or even in deep shadow in full forest. Shadow darners can also be found near ditches, slow streams, and ponds. This darner has a long flight season of late April to November.
The shadow darner naiad feeds on a wide variety of aquatic insects, such as mosquito larvae, other aquatic fly larvae, mayfly larvae, and freshwater shrimp. They also feed on small fish and tadpoles. This adult will eat almost any soft-bodied flying insect, including mosquitoes, flies, butterflies, moths, mayflies, and stoneflies.


Aeshna verticalis, the green-striped darner,  is a species of dragonfly in the family Aeshnidae. It is typically found in northeastern United States and southern Ontario, Quebec, and New Brunswick. The green-striped darner has an IUCN conservation status of "Least Concern" with a stable population.


The crimson sunbird is a resident breeder in tropical southern Asia from India, through Nepal, Bangladesh and Myanmar to Indonesia and Brunei Darussalam. Two or three eggs are laid in a suspended nest in a tree. This species occurs in forest and cultivated areas.


The spotted eagle ray (Aetobatus narinari) is a cartilaginous fish of the eagle ray family, Myliobatidae. As traditionally recognized, it is found globally in tropical regions, including the Atlantic, Pacific and Indian Oceans. Recent authorities have restricted it to the Atlantic (including the Caribbean and Gulf of Mexico) with other populations recognized as the ocellated eagle ray (A. ocellatus) and Pacific white-spotted eagle ray (A. laticeps). Spotted eagle rays are most commonly seen alone, but occasionally swim in groups. They are ovoviviparous, the female retaining the eggs then releasing the young as miniature versions of the parent.
This ray can be identified by its dark dorsal surface covered in white spots or rings. Near the base of the ray's relatively long tail, just behind the pelvic fins, are several venomous, barbed stingers. Spotted eagle rays commonly feed on small fish and crustaceans, and will sometimes dig with their snouts to look for food buried in the sand of the sea bed. These rays are commonly observed leaping out of the water, and on at least two occasions have been reported as having jumped into boats, in one incident resulting in the death of a woman in the Florida Keys. The spotted eagle ray is hunted by a wide variety of sharks. The rays are considered near threatened on the IUCN Red List. They are fished mainly in Southeast Asia and Africa, the most common market being in commercial trade and aquariums. They are protected in the Great Barrier Reef.


Spotted eagle rays have flat disk-shaped bodies, deep blue or black with white spots on top with a white underbelly, and distinctive flat snouts similar to a duck's bill. Their tails are longer than those of other rays and may have 2–6 venomous spines, just behind the pelvic fins. The front half of the long and wing-like pectoral disk has five small gills in its underside.
Mature spotted eagle rays can be up to 5 meters (16 ft) in length; the largest have a wingspan of up to 3 meters (10 ft) and a mass of 230 kilograms (507 lb).


Spotted eagle ray preys mainly upon bivalves, crabs, whelks, benthic infauna they also feed on mollusks, crustaceans, particularly malacostracans. and also upon hermit crabs, shrimp, octopuses, and some small fish.
The spotted eagle ray's specialized chevron-shaped tooth structure helps it to crush the mollusks' hard shells. The jaws of these rays have developed calcified struts to help them break through the shells of mollusks, by supporting the jaws and preventing dents from hard prey. These rays have the unique behavior of digging with their snouts in the sand of the ocean.  While doing this, a cloud of sand surrounds the ray and sand spews from its gills. One study has shown that there are no differences in the feeding habits of males and females or in rays from different regions of Australia and Taiwan.


Spotted eagle rays prefer to swim in waters of 24 to 27 °C (75 to 81 °F). Their daily movement is influenced by the tides; one tracking study showed that they are more active during high tides. Uniquely among rays they dig with their snouts in the sand, surrounding themselves in a cloud of sand that spews from their gills. They also exhibit two motions in which the abdomen and the pectoral fins are moved rapidly up and down: the pelvic thrust and the extreme pelvic thrust. The pelvic thrust is usually performed by a solitary ray, and repeated four to five times rapidly. The extreme pelvic thrust is most commonly observed when the ray is swimming in a group, from which it will separate itself before vigorously thrusting with its pectoral fins. The rays also performs dips and jumps; in a dip the ray will dive and then come back up rapidly, perhaps as many as five times consecutively. There are two main types of jump: in one, the ray propels itself vertically out of the water, to which it returns along the same line; the other is when the ray leaps at a 45 degree angle, often repeated multiple times at high speeds. When in shallow waters or outside their normal swimming areas the rays are most commonly seen alone, but they do also congregate in schools. One form of travelling is called loose aggregation, which is when three to sixteen rays are swimming in a loose group, with occasional interactions between them. A school commonly consists of six or more rays swimming in the same direction at exactly the same speed.


As traditionally defined, spotted eagle rays are found globally in tropical regions from the Indo-Pacific region from the western Pacific Ocean, the Indian Ocean, and the western Atlantic Ocean.
They are found in shallow coastal water by coral reefs and bays, in depths down to 80 meters (262 ft).
Spotted eagle rays are found in warm and temperate waters worldwide. In the western Atlantic Ocean it is found off the eastern coast of United States of America, the Gulf Stream, the Caribbean, and down past the southern part of Brazil. In the Indian Ocean, it is found from the Red Sea down to South Africa and eastward to the Andaman Sea. In the Western Pacific Ocean it can be found near Japan and north of Australia. In the Central Pacific Ocean, it can be found throughout the Hawaiian Islands. In the Eastern-Pacific Ocean, it is found in the Gulf of California down through Puerto Pizarro, an area that includes the Galapagos Islands. Spotted eagle rays are most commonly seen in bays and reefs. They spend much of their time swimming freely in open waters, generally in schools close to the surface, and can travel long distances in a day.
Within these regions, there are significant variations in genetics and morphology. As a consequence, recent authorities have split it into three: This restricts the true spotted eagle ray (A. narinari) to the Atlantic, while the Indo-Pacific population is the ocellated eagle ray (A. ocellatus) and the East Pacific is the Pacific white-spotted eagle ray (A. laticeps).


The spotted eagle ray is included in the IUCN's Red List as "near threatened". The rays are caught mainly in Southeast Asia and Africa.  They are also common in commercial marine life trade and are displayed in aquariums. Among the many efforts to help protect this species, South Africa's decision to deploy fewer protective shark nets has reduced the number of deaths caused by entanglement. South Africa has also placed restrictions on the number of rays that can be bought per person per day. In the U.S. state of Florida, the fishing, landing, purchasing and trading of spotted eagle ray are outlawed. This ray is also protected in the Great Barrier Reef on the eastern coast of Australia.
In Europe there is a breeding program managed by the EAZA for spotted eagle rays to reduce the amount of wild caught individuals needed by public aquaria. From the start until 2018 Burgers' Zoo in the Netherlands kept the studbook. Since 2018, Wroclaw Zoo in Poland is the new studbook keeper. Burgers' Zoo was also the first place in Europe to breed with the species and in 2018 was the most successful breeder worldwide with over 55 births.


The adult males of these spiders reach 4.6–6.2 millimetres (0.18–0.24 in) of length, while females are 6.5–10.3 millimetres (0.26–0.41 in) long. The coloration of the body is very variable. The basic color varies from light or dark brown to yellowish-orange. They usually show a dark longitudinal band in the middle of the abdomen, with some trasversal bands. Sometimes the opisthosoma has two large white spots or one dark brown light-edged spot on the back. Chelicerae and legs are brown,The whole body is densely hairy, especially in the anterior body section (prosoma). The thorax is relatively broad, pear-shaped. The abdomen (opisthosoma) is quite flat and wider than long.


It is native to western North America from British Columbia to California to Colorado, where it grows in many habitat types. This is an aromatic perennial herb growing an erect stem with widely spaced leaves, each lance-shaped to nearly triangular and toothed.


The leaves are up to 8 centimeters long and 7 wide. The inflorescence is a dense spike of many flowers. Each flower has long sepals tipped with bright purple and tubular corollas in shades of pink and purple. The fruit is a light brown, fuzzy nutlet about 2 millimeters long.


The red-winged blackbird (Agelaius phoeniceus) is a passerine bird of the family Icteridae found in most of North America and much of Central America. It breeds from Alaska and Newfoundland south to Florida, the Gulf of Mexico, Mexico, and Guatemala, with isolated populations in western El Salvador, northwestern Honduras, and northwestern Costa Rica. It may winter as far north as Pennsylvania and British Columbia, but northern populations are generally migratory, moving south to Mexico and the southern United States. Claims have been made that it is the most abundant living land bird in North America, as bird-counting censuses of wintering red-winged blackbirds sometimes show that loose flocks can number in excess of a million birds per flock and the full number of breeding pairs across North and Central America may exceed 250 million in peak years. It also ranks among the best-studied wild bird species in the world. The red-winged blackbird is sexually dimorphic; the male is all black with a red shoulder and yellow wing bar, while the female is a nondescript dark brown. Seeds and insects make up the bulk of the red-winged blackbird's diet.


The red-winged blackbird is one of five species in the genus Agelaius and is included in the family Icteridae, which is made up of passerine birds found in North and South America. The red-winged blackbird was formally described as Oriolus phoeniceus by Carl Linnaeus in 1766 in the twelfth edition of his Systema Naturae, but was later moved with the other American blackbirds to the genus Agelaius by Louis Jean Pierre Vieillot in 1816. Linnaeus specified the type location as "America" but this was restricted to Charleston, South Carolina in 1928. The genus name is derived from Ancient Greek agelaios, meaning "gregarious". The specific epithet, phoeniceus, is Latin meaning "crimson" or "red". The red-winged blackbird is a sister species to the red-shouldered blackbird (Agelaius assimilis) that is endemic to Cuba. These two species are together sister to the tricolored blackbird (Agelaius tricolor) that is found on the Pacific coast region of the California and upper Baja California in Mexico.
Depending on the authority, between 20 and 24 subspecies are recognised which are mostly quite similar in appearance. However, there are two isolated populations of bicolored blackbirds that are quite distinctive: A. p. californicus of California and A. p. gubernator of central Mexico. The taxonomy and relationships between these two populations and with red-winged blackbirds is still unclear. Despite the similarities in most forms of the red-winged blackbird, in the subspecies of Mexican Plateau, A. p. gubernator, the female's veining is greatly reduced and restricted to the throat; the rest of the plumage is very dark brown, and also in a different family from the European redwing and the Old World common blackbird, which are thrushes (Turdidae). In the California subspecies, A. p. californicus and A. p. mailliardorum, the veining of the female specimens also covers a smaller surface and the plumage is dark brown, although not in the gubernator grade; and also its superciliary list is absent or poorly developed. The male subspecies mailliardorum, californicus, aciculatus, neutralis, and gubernator lack the yellow band on the wing that is present in most male members of the other subspecies. The red-shouldered blackbird was formerly considered as a subspecies of red-winged blackbird. They were split by the American Ornithologists' Union in 1997.
Listed below are the red-winged blackbird subspecies and groups of subspecies recognized as of January 2014 with their respective distribution areas and the location of their wintering quarters:
Group phoeniceus:
A. p. arctolegus – from southeastern Alaska and Yukon to the north-central United States; migrates to the south-central United States.
A. p. fortis – from Montana to southeastern New Mexico (east of the Rocky Mountains); migrates to Texas.
A. p. nevadensis – from southeastern British Columbia to Idaho, southeastern California, and southern Nevada; migrates to southern Arizona.
A. p. caurinus – from the southwest coast of British Columbia to northwest California; migrates to central California.
A. p. aciculatus – mountains of central-southern California (central-eastern Kern County).
A. p. neutralis – from the coast of southern California (San Luis Obispo County) to the northwest of Baja California.
A. p. sonoriensis – from southeastern California to northeast Baja California, southern Nevada, central Arizona, and northwest Mexico.
A. p. nyaritensis – coastal plains of southwestern Mexico (Nayarit).
A. p. grinnelli – Pacific Slope from western Guatemala to northwest Costa Rica (Guanacaste).
A. p. phoeniceus – from southeastern Canada to Texas and the southeastern United States.
A. p. littoralis – Gulf Coast from Southeast Texas to Northwest Florida.
A. p. mearnsi – extreme southeast Georgia and northern Florida.
A. p. floridanus – South Florida (Everglades to Key West).
A. p. megapotamus – from central Texas and the lower Rio Bravo valley to eastern Mexico (northern Veracruz).
A. p. richmondi – Caribbean slope from Mexico (southern Veracruz) to Belize and northern Guatemala.
A. p. pallidulus – southeast Mexico (north of the Yucatan peninsula).
A. p. nelsoni – south-central Mexico (from Morelos and the adjacent Guerrero west of Puebla and Chiapas).
A. p. matudae – tropical southeast Mexico (not always recognised).
A. p. arthuralleni – northern Guatemala.
A. p. brevirostris – Caribbean slope of Honduras and southeastern Nicaragua (sometimes included in richmondi).
A. p. bryanti – northwest Bahamas.
Group californicus/ mailliardorum:
A. p. mailliardorum – Central California Coast.
A. p. californicus – Central Valley of California.
Group gubernator :
A. p. gubernator – Mexican Plateau (Durango to Zacatecas, Mexico and Tlaxcala).


The common name for the red-winged blackbird is taken from the mainly black adult male's distinctive red shoulder patches, or epaulets, which are visible when the bird is flying or displaying. At rest, the male also shows a pale yellow wingbar. The spots of males less than one year old, generally subordinate, are smaller and more orange than those of adults. The female is blackish-brown and paler below. The female is smaller than the male, at 17–18 cm (6.7–7.1 in) long and weighing 41.5 g (1.46 oz), against his length of 22–24 cm (8.7–9.4 in) and weight of 64 g (2.3 oz). The smallest females may weigh as little as 29 g (1.0 oz) whereas the largest males can weigh up to 82 g (2.9 oz). Each wing can range from 8.1–14.4 cm (3.2–5.7 in), the tail measures 6.1–10.9 cm (2.4–4.3 in), the culmen measures 1.3–3.2 cm (0.51–1.26 in) and the tarsus measures 2.1 cm (0.83 in). The upper parts of the female are brown, while the lower parts are covered by an intense white and dark veining; also presents a whitish superciliary list. Females exhibit a year or salmon pink stain on the shoulders and a clear pink color on the face and below this, while older show a stain usually more crimson on the shoulders and dark pink hue on and under the face. Observations in females in captivity indicate that small amounts of yellow pigment are present on the shoulders of these after leaving the nest, that the concentration of the pigment increases with the first winter plumage after the change of the feathers and that the passage from yellow to orange generally takes place in the second summer with the acquisition of the second winter plumage, after which no further changes in feather color occur. The colored area on the wing increases in surface with the age of the female, and varies in intensity from brown to a bright red-orange similar to that of the males in their first year.
Young birds resemble the female, but are paler below and have buff feather fringes. Both sexes have a sharply pointed bill. The tail is of medium length and is rounded. The eyes, bill, and feet are all black. Unlike most North American passerines which develop their adult plumage in their first year of life so that the one-year-old and the oldest individual are indistinguishable in the breeding season, the red-winged blackbird does not. It acquires its adult plumage only after the breeding season of the year following its birth when it is between thirteen and fifteen months of age. Young males go through a transition stage in which the wing spots have an orange coloration before acquiring the most intense tone typical of adults.
The male measures between 22 and 24 cm (8.7 and 9.4 in) in length, while the female measures 17 or 18 cm (6.7 or 7.1 in). Its wingspan is between 31 and 40 cm (12 and 16 in) approximately. Both the peak male and the legs, the claws and the eyes are black; in the female beak is dark brown and clear in the upper half at the bottom, and the tail is medium in length and rounded. As in other species polygyny exists, the red-winged blackbird exhibit considerable sexual dimorphism both in plumage and size, males weighing between 65 and 80g the females about 35g. Males are 50% heavier than females, 20% larger in its linear dimensions, and 20% larger compared to the length of their wings. The trend towards greater dimorphism in the size of non-monogamous ichterid species indicates that the larger size of males has evolved due to sexual selection.
The male is unmistakable except in the far west of the US, where the tricolored blackbird occurs. Males of that species have a darker red epaulet edged with white, not yellow. Females of tricolored, bicolored, red-shouldered and red-winged blackbirds can be difficult to identify in areas where more than one form occurs. In flight, when the field marks are not easily seen, the red-winged blackbird can be distinguished from less closely related icterids such as common grackle and brown-headed cowbird by its different silhouette and undulating flight.


The most critical period of feather molting runs from late August to early September. When viewed in flight, they have a misaligned or "moth-eaten" appearance and generally slower and more laborious travel. Their mobility is reduced due to the lack of several remiges or rectrices or these are not entirely renewed. Most of the red-winged blackbirds have moved almost entirely by October. By then, some birds have not completed the molt of the feathers of the capital region and the helmsmen of the center of the tail and the internal secondary sprouts have only partially emerged from the pod. Virtually all individuals have completed their molts by mid-October.
Birds do not begin their migration to wintering quarters until the two outer primary sprouts and the two inner or central rectrices have completed at least two-thirds of their development. Therefore, there is a correlation between molting, particularly replacement of the remiges and rectrices, and fall migration in red-winged blackbirds.


The red-winged blackbird is widely spread throughout North America, except in the arid desert, high mountain ranges, and arctic or dense afforestation regions. It breeds from central-eastern Alaska and Yukon in the northwest, and Newfoundland in the northeast, to northern Costa Rica in the south, and from the Atlantic to the Pacific. Northern populations migrate to the southern United States, but those that breed there, in Mexico, and in Central America are sedentary. Red-winged blackbirds in the northern reaches of the range are migratory, spending winters in the southern United States and Central America. Migration begins in September or October, but occasionally as early as August. In western and Central America, populations are generally non-migratory.
The red-winged blackbird inhabits open grassy areas. It generally prefers wetlands, and inhabits both freshwater and saltwater marshes, particularly if cattail is present. It is also found in dry upland areas, where it inhabits meadows, prairies, and old fields. In a large part of its distribution area, it constitutes the most abundant passerine bird in the swamps in which it nests. It is also present in areas without much water, where it inhabits open fields – often agricultural areas – and sparse deciduous forests.
In the winter of 1975–1976, near Milan (western Tennessee), red-winged blackbirds were observed resting in a mixed roost that came to house 11 million individuals in January and early February in a plantation of 4.5 hectares of yellow pine (Pinus taeda) with little undergrowth was seen in soybean fields during the day, being that these constituted only 21% of the habitat in the area and that the other bird species present in the roost were not commonly observed in these fields; they were also common in cornfields. the presences of the bird in feedlots has increased as winter progressed, but, accounting for less than 5% of the ichterids and starlings recorded in both feedlots of cows and pigs, they were much rarer there than the brown-headed cowbird, common grackle and common starling.


In that same study, reproductive individuals in upland habitats demonstrated a slight preference for old and new grasses such as Phleum pratense, Dactylis glomerata, Poa spp., Festuca spp., and Bromus spp. In the early part of the breeding season and new non-graminoid herbaceous plants in the mid and late season. In wetlands, they consistently preferred old and new broadleaf monocotyledons, primarily Carex spp, broadleaf, and Typha spp, and consistently rejected old and new narrow-leaved monocots, primarily narrow-leaved Phalaris arundinacea and Calamagrostis canadensis, and non-graminoid herbaceous plants. Red-winged blackbirds breeding in the highlands favored older tall vegetation only in the early part of the breeding season on April to May, and higher new vegetation and dense vegetation throughout the breeding season. For their part, those who settled in wetlands seemed to have a slight predilection for taller old vegetation. During the breeding season, this species is attracted to tall vegetation that restricts visibility.
Early season preferences for old grassland in the highlands and old broadleaf monocotyledons in wetlands point to the importance of upright residual vegetation. Highland grasses and broadleaf monocots in wetlands stand partially vertically and are easily visible in early spring, unlike clovers, narrow-leaved monocotyledons in wetlands and most non-graminoid herbaceous plants. Old alfalfa plants are also partially upright in early spring, but they are not as consistently chosen a species as old grasses. The start of the territorial activity is earlier when the amount of residual vegetation is large. Vegetation structural strength also appears to be important for nesting as females tend to broadleaf monocotyledons in wetlands throughout the breeding season and new non-graminoid herbaceous plants in the mid to late season.
In southwestern Michigan, the density of cattail stems in different ponds was positively related to the concentration of reproductive adults. However, other studies detected a predilection for more scattered vegetation. One found that red-winged blackbirds avoided the marsh wren (Pantaneros chivirenes), a predator common in nests of this species, reproducing among more dispersed vegetation, which was more easily defended from the chivirines; conversely, the chivirines seemed to prefer the denser vegetation, where they were more likely to avoid the aggressiveness of the red-winged blackbird; These differences in habitat selection between one species and another resulted in spatial segregation of their breeding areas. On the other hand, red-winged blackbirds tend to opt for small lots of vegetation and plants with thick stems.


The red-winged blackbird is territorial, polygynous, gregarious and a short-distance migratory bird. Its way of flying is characteristic, with rapid wing flaps punctuated by brief periods of gliding flight. The behavior of males makes their presence easily perceived: they perch in high places such as trees, bushes, fences, telephone lines, etc. Females tend to stay low, prowling through the vegetation and building their nests. They can be found in home gardens, particularly during their migration, if seeds have been scattered on the ground. The forest curtains serve as a resting place during the day.
For several weeks after their first appearance in early spring, red-winged blackbirds are generally seen in flocks made up entirely of males. During those days, they are seldom seen at their breeding sites, except early morning and late afternoon. In most of the remaining hours of the day, they frequent open and often elevated agricultural land, where they feed mainly on grain stubble and grassy fields. When disturbed while eating, they fly to the nearest deciduous trees and immediately after landing they begin to sing.


The red-winged blackbird is omnivorous. It feeds primarily on plant materials, including seeds from weeds and waste grain such as corn and rice, but about a quarter of its diet consists of insects and other small animals, and considerably more so during breeding season. It prefers insects, such as dragonflies, damselflies, butterflies, moths, and flies, but also consumes snails, frogs, eggs, carrion, worms, spiders, mollusks. The red-winged blackbird forages for insects by picking them from plants, or by catching them in flight. Sometimes insects are obtained by exploring the bases of aquatic plants with their small beaks, opening holes to reach the insects hidden inside. Aquatic insects, particularly odonates emerging, are of great importance in the diet of red-winged blackbirds that breed in swamps. These birds typically capture the odonates when the larvae climb up the stem of a plant from the water, get rid of their exuviae, and cling to the vegetation while their exoskeletons harden. The years of emergence of Periodical cicadas, it provides an overabundant amount of food. In season, they also eat blueberries, blackberries and other fruits. According to Edward Howe Forbush, when they arrive north in the spring, they feed in the fields and meadows. Then, they follow the plows, collecting larvae, earthworms and caterpillars left exposed, and in case there is a plague of Paleacrita vernata caterpillars in a fruit orchard, these birds will fly a kilometer to get them for their chicks.
In season, it eats blueberries, blackberries, and other fruit. These birds can be lured to backyard bird feeders by bread and seed mixtures and suet. In late summer and in autumn, the red-winged blackbird will feed in open fields, mixed with grackles, cowbirds, and starlings in flocks which can number in the thousands. It feeds on corn while it is maturing; once the grain has hardened it is relatively safe from this bird, since its beak and digestive system are not adapted for the consumption of hard and whole corn grains, unlike the Common grackle, which has a longer and stronger beak. Studies of the stomachs of individuals of both sexes reveal that males consume higher proportions of crop grains, while females ingest a relatively larger amount of herb seeds and animal matter. In the winter of 1975–1976, near Milan, western Tennessee, corn and herb seeds were the main foods consumed by red-winged blackbird. Herbs whose seeds were commonly consumed were Sorghum halepense, Xanthium strumarium, Digitaria ischaemum, Sporobolus spp., Polygonum spp. and Amaranthus spp.


The red-winged blackbird nests in loose colonies. The nest is built in cattails, rushes, grasses, sedge, or alder or willow bushes. The nest is constructed entirely by the female over the course of three to six days. It is a basket of grasses, sedge, and mosses, lined with mud, and bound to surrounding grasses or branches. It is located 7.6 cm (3.0 in) to 4.3 m (14 ft) above water.
A clutch consists of three or four, rarely five, eggs. Eggs are oval, smooth and slightly glossy, and measure 24.8 mm × 17.55 mm (0.976 in × 0.691 in). They are pale bluish green, marked with brown, purple, and/or black, with most markings around the larger end of the egg. These are incubated by the female alone, and hatch in 11 to 12 days. Red-winged blackbirds are hatched blind and naked, but are ready to leave the nest 11 to 14 days after hatching.
Red-winged blackbirds are polygynous, with territorial males defending up to 10 females. However, females frequently copulate with males other than their social mate and often lay clutches of mixed paternity. Pairs raise two or three clutches per season, in a new nest for each clutch.
The reproductive season of the red-winged blackbird extends approximately from the end of April to the end of July. On the other hand, in different states has been estimated that the period in which the active nests contained eggs lay between beginning in late April and early late August; and in northern Louisiana nests were found to harbor chicks from late April to late July. The peak of the nesting season (the time with the highest number of active nests) has been recorded between the first half of May and the beginning of June in different places. A study in eastern Ontario found that although red-winged blackbirds began nesting earlier in years with warm springs, associated with low winter values in the North Atlantic Oscillation Index, egg laying dates remained unchanged. Male testosterone levels peak in the early part of the breeding season, but remain high throughout the season. Females reproduce for up to ten years.
Many aspects of territorialism (for example, the number of male songs and displays, and the number of intrusions into foreign territories) peak before mating. After this, the frequency of many of the territorial behaviors decreases and the males are mainly concerned with defending the females, eggs, and chicks against predation. Experiments in the systematic removal of birds from their territories suggest that the extra population of males that is present in swamps before copulations disappears after copulation.
Predation of eggs and nestlings is quite common. Nest predators include snakes, mink, raccoons, and other birds, even as small as marsh wrens. The red-winged blackbird is occasionally a victim of brood parasites, particularly brown-headed cowbirds. Since nest predation is common, several adaptations have evolved in this species. Group nesting is one such trait which reduces the risk of individual predation by increasing the number of alert parents. Nesting over water reduces the likelihood of predation, as do alarm calls. Nests, in particular, offer a strategic advantage over predators in that they are often well concealed in thick, waterside reeds and positioned at a height of one to two meters. Males often act as sentinels, employing a variety of calls to denote the kind and severity of danger. Mobbing, especially by males, is also used to scare off unwanted predators, although mobbing often targets large animals and man-made devices by mistake. The brownish coloration of the female may also serve as an anti-predator trait in that it may provide camouflage for her and her nest while she is incubating.


Predators of red-winged blackbirds include such species as raccoons, American mink, long-tailed weasels, Eurasian magpies, common grackles, owls, red-tailed hawks, short-tailed hawks, and snakes (such as the northern water snake and the plains garter snake). Ravens and grazers such as marsh wrens feed on eggs (and even small chicks), if the nest is left unattended, destroying the eggs, occasionally drinking from them, and pecking the nestlings to death.
The relative importance of different nest predators varies by geographic region: the top predators in different regions include the marsh wren in British Columbia, the magpies in Washington, and the raccoons in Ontario. The incidence of avian predation in red-winged blackbird nests is higher in western populations than in eastern populations.
Due to high predation rates, especially of eggs and chicks, the red-winged blackbird has developed various adaptations to protect its nests. One of them consists of nesting in groups, which reduces the danger since there is a greater number of alert parents. Nesting over water also lowers the chances of an attack. Nests in particular offer a strategic advantage as they are often hidden among dense riparian reeds, at a height of one or two meters. males often act as sentinels, using a repertoire of calls. Males in particular hunt down potential predators to scare them away, even when dealing with much larger animals. Aggressiveness of the red-winged blackbird towards the marsh wren, which also nests in swamps, causes a partial interspecies territorialism. On the other hand, nocturnal predators such as raccoons and American mink are not attacked by adults. Coloration of the female could serve to camouflage it, protecting it and its nest when it is incubated.
The red-winged blackbird can accommodate ectoparasites such as various Phthiraptera, the Ischnocera Philopterus agelaii and Brueelia ornatissima, and hematophagous mites, like Ornithonyssus sylviarum, and endoparasites like Haemoproteus quiscalus, Leucocytozoon icteris, Plasmodium vaughani, nematodes, flukes and tapeworms.


Red-winged blackbirds that breed in the northern part of their range, i.e., Canada and border states in the United States, migrate south for the winter. However, populations near the Pacific and Gulf coasts of North America and those of Middle America are year-round resident. Red-winged blackbirds live in both Northern U.S. and Canada, ranging from Yucatan Peninsula in the south to the southern part of Alaska. These extensions account for the majority of the continent stretching from California's Pacific coast and Canada to the eastern seaboard. Much of the populations within Middle America are non-migratory. During the fall, populations begin migrating towards Southern U.S. Movement of red-winged blackbirds can begin as early as August through October. Spring migration begins anywhere between mid-February to mid-May. Numerous birds from northern parts of the U.S., particularly the Great lakes, migrate nearly 1,200 km (750 mi) between their breeding season and winter Winter territorial areas differ based on geographic location. Other populations that migrate year-round include those located in Middle America or in the western U.S. and Gulf Coast. Females typically migrate longer distances than males. These female populations located near the Great Lakes migrate nearly 230 km (140 mi) farther. Yearly-traveled females also migrate further than adult males, while also moving roughly the same distance as other adult females. Red-winged blackbirds migrate primarily during daytime. In general, males’ migration flocks arrive prior to females in the spring and after females in the fall.


The red-winged blackbirds can devastate farm fields. Despite the fact that they consume weed seeds, they are known to cause great damage to agriculture due to their habits of resting in massive groups and their taste for agricultural products. may be also causes harm to plantings corn, rice, sunflower and sorghum, particularly important near roosts. Red-winged blackbird is the largest species of Icterid in North America and the most damaging to crops. From 215 birds Neotropical migrants have been identified as causing, by a wide margin, the greatest economic loss. In North America, the damage to corn crops by this species has increased since the late 1960s to early 1980s, perhaps because of the increase in the area for grain production, and due to the reduction of small areas with stubble, hayfields and uncultivated land, which, in turn, accentuated the bird's dependence on corn to ensure its livelihood.
Apparently, the male does more damage to this grain than the female. In some areas of Ohio, corn can account for up to 75% of the diet of males and only 6% of that of females in August and September. In South Dakota, in the late summer, the Gizzards of the males studied contained 29% corn, while in the case of the female that number was limited to 9%.


The red-winged blackbird is the dominant species in the large concentrations of blackbirds that feed on the fields of sunflower, corn and small grains maturing in late summer or early fall in the Dakotas. In the 1970s, losses to sunflower and corn crops caused by blackbirds in the Dakotas exceeded $3 million annually for each case. In the northern part of the Great Plains, an area known as the Prairie Pothole Region, in red-winged blackbirds are very abundant in summer, they congregate in post-reproductive flocks that significantly harm crops, particularly sunflower plantations near their home sites. Most sunflower damage occurs between mid-August and early September, when the calorie content of immature seeds is low and birds must consume more of them to satiate themselves. During this initial stage of predation on sunflower crops in which more than 75% of the total damage is caused, the red-winged blackbird represent 80% of the blackbirds observed in the fields of this seed. This period predates the massive migration of birds and most of them are of local origin. Most remain within 200 km (120 mi) of their native sites until the molting of their feathers is complete or nearly complete in late August or early September. Damage can be quite serious in the center and southeast of North Dakota and Northeast South Dakota, areas of high concentration of sunflower production and abundant wetlands that attract red-winged blackbird during the breeding season.
Investigations conducted between 1968 and 1979 revealed that blackbirds, notably the black-winged blackbird and the Common grackle, annually destroyed less than 1% of corn crops in Ohio, amounting to a loss between 4 and 6 million dollars according to 1979 prices. Insects, weeds, diseases and fungi. They are likely to cause losses of more than 20% of the total potential maize harvest in the Midwest states, and harvesting procedures often leave 5% or more of the maize in the fields, revealing that comparative damage by Ichteridae in Ohio is not great. However, the average damage caused by these birds is not of as much economic importance as its distribution among producers. While 97.5% of cornfields studied in Ohio suffered a loss of less than 5%, which constituted about 60% of the total damage in the state, the remaining 2.5% often incurred losses of more than 5% and represented approximately 40% of total injury in Ohio.
All Ohio counties experience some degree of predation on their maize crops from blackbirds, but those most affected are a few counties where marshes that roost them still abound. The counties of Ottawa, Sandusky and Lucas, on the waters of Sandusky Bay and Lake Erie, were the hardest hit. These three counties, among the 19 studied between 1968 and 1976, contained 62% of the fields in which the losses exceeded 5% and 77% of those in which they exceeded 10%. Other counties with extensive localized damage were Erie, Ashtabula also located on the coasts of the mentioned water courses and Hamilton. Almost all the plantations with damages greater than 5% were within 8 km (5.0 mi) of some important roosting of blackbirds. In the 1968–1976 period, in northeast Sandusky County and northwest Ottawa, where large roosts of up to a million birds were discovered in late summer and fall, average losses exceeded 9% in fields 3 to 5 km (1.9 to 3.1 mi) from the roosts, but they were less than 5% at 8 km (5.0 mi) and less than 2% at 16 km (9.9 mi). In southwestern Ontario, in the summer of 1964, it was found that the greatest damage to the cornfields by red-winged blackbird also occurred near roosts in swamps.


As early as 1667, Massachusetts Bay settlers had enacted laws to try to reduce blackbird populations and mitigate damage to corn. According to Henry David Thoreau, a law provided that each single man in a town must kill six of those birds and, as a punishment for not doing so, he could not marry until he had complied with the aforementioned design. Obviously, since blackbirds reproduce at a much higher speed than humans marry, this control strategy was a failure. Pioneers traveling west to the Great Lakes region faced similar problems. By 1749, blackbirds were so abundant around western Lake Erie that people took turns watching over the maturing grain crops.
At the beginning of the 20th century, in some places, when the reeds dried up, these circumstances were used to kill these birds in the following way. A crew approached a roost in silence, hidden in the darkness of the night, and simultaneously lit the reeds at various points, which were quickly enveloped by a single great flame. This caused a huge tumult among the red-winged blackbird, which, lit by fire, were shot down in large numbers as they hovered in midair and screamed all over the place. Sometimes straw was used for the same purpose, which was previously scattered near reeds and alder bushes (Alnus spp.) in which they gathered to rest, the burning of which caused great consternation among the birds. The gang returned the next day to collect the hunted prey.
Arthur Cleveland Bent says that, before it was banned the sale of prey hunting in the market, red-winged blackbirds were massacred in large numbers in autumn and sold in markets. When they had put on weight on a diet of grains or rice, their small bodies were served as delicious snacks on the gourmet tables . Few could distinguish them from charlatans (Dolichonyx oryzivorus).
In 1926, when the US Biological Survey – predecessor of the United States Fish and Wildlife Service carried out its first compilation of roosts of undesirable ichterids, it was recorded in Ohio, with its large populations of these birds and the fifth largest area allocated to the cultivation of corn among the American states, a number of complaints higher than in any other state. During the 1950s, bird control committees were organized in some counties to deal with the damage to maize caused by blackbirds and the Ohio Agricultural Experiment Station now the Ohio Agricultural Research and Development Center and the Department of Zoology and Entomology of the state university began investigating the problem.


Crop predation has led to the use of traps, poison and surfactants by farmers in an attempt to control populations of red-winged blackbirds; these last properties suppress waterproof feathers, making them extremely vulnerable to the cold, but their effectiveness depends on certain atmospheric conditions, namely low temperatures and rainfall. Programs in which baits were used poisoned to reduce icteride concentrations in late summer have been unsuccessful. While thousands of birds have occasionally died, the effect on large roosting-associated flocks that sometimes contain more than a million individuals is small; In addition, specimens of other species frequently die. The use of large lure traps, which often catch hundreds of birds per day, is also ineffective against these large flocks.


It is a species of least-concern. Being one of the largest and most widely distributed birds in North America, little has been done to protect it from the effects of habitat destruction and urbanization. It can survive in a wide range of environments, many populations manage to overcome the loss of natural habitats. However, red-winged blackbirds thrive in wetland areas and with the destruction of natural wetlands their population is likely to shrink. The species is protected under the Migratory Bird Treaty Act 1918, a formal treaty between the United States and Canada that was later expanded to include Mexico. This law gives them legal protection in the United States, but they can be killed "when they are found preying or about to prey on ornamental trees or trees planted for shade, crops, livestock or wildlife."
A study in Illinois indicated that red-winged blackbird populations doubled between 1908 and 1958. It had traditionally reproduced in wetlands, with Ohio primarily inhabiting swamps associated with lakes and rivers. During the 20th century, however, it adapted to man-made habitat changes and now often nests in hayfields, along roads and ditches, and elsewhere in the highlands.
Despite its successful adaptation to changes in practices related to land use, populations of red-winged blackbird have reduced the width of its range during the second half of the twentieth century, and Changes in the abundance and adequacy of grasslands have been implicated in this. In Ohio, the red-winged blackbird was negatively affected between 1966 and 1996 by the decrease in hay production, the earlier harvest of hay and the increase in crops planted in furrows, a situation homologous to that suffered by other species of birds from less numerous grasslands. Although the negative effect of the increased efficiency and the diminishing diversity of modern agricultural practices on the populations of red-winged blackbird in this state may be perceived as a positive event by the producers of corn and sunflower, the agricultural practices that have precipitated the numerical decline of this species may have caused more severe repercussions for birds from less common grasslands, such as the upland sandpiper and the Grasshopper sparrow.
In the late 1970s, Ohio hosted the highest density of blackbirds during the breeding season among all US states and Canadian provinces, However, between 1966 and 1996, the reproductive populations in this state showed a marked decrease. Decreasing the area under hay other than alfalfa between those same years was likely to reduce the availability of quality nesting habitats. Likewise, corn even though it attracts insects that red-winged blackbirds consume and despite the fact that their grains represent in themselves an energy source and soybeans which do not constitute a food source for the species, which together made up 1966 to 1996, an average of 70% of the cultivated area in Ohio, do not provide adequate nesting habitat. In turn, the large annual fluctuations in the area cultivated with hay, both from one year to the next and within the same year of a certain reproductive season, Similarly, in Ontario between 1974 and 1995, the average size of harems in a given year was positively related to hay production from the previous year, although the percentage variation in the size of harems from one season to the next the following was not linearly related to the annual percentage change in hay production; Hay production declined during this period and the size of the harems decreased from approximately three females per male to 1.6 females.
In addition in Ontario, a negative relationship was found between the North Atlantic Oscillation Index (NAO) in the six months prior to a certain reproductive season and the size of the harems. If winter mortality contributes to the decrease in the size of the harems, the annual variations in their size should be related to the annual changes in the rates of return of marked territorial males. Variations in the size of harems and male return rates per year were indeed found to be positively correlated. Although not as pronounced, the relationship between male return rates and winter NAO values was similar to that between changes in harem size and NAO.
On the other hand, in southwestern Quebec, red-winged blackbird populations doubled between 1966 and 1981, apparently in response to the development of maize production. The increased availability of residual grains in spring and summer. the reproductive season will probably play a fundamental role in the growth of this bird's population. Likewise, in the mid-1990s, the reproductive population of North Dakota which at least between 1994 and 2002 was at the forefront among all the American states and Canadian provinces in terms of population density of this species began to increase rapidly.


The tricolored blackbird (Agelaius tricolor) is a passerine bird of the family Icteridae. Its range is limited to the coastal areas of the Pacific coast of North America, from Northern California in the U.S. (with occasional strays into Oregon), to upper Baja California in Mexico.
This highly social and gregarious bird forms the largest colonies of any North American landbird, with a single breeding colony often consisting of tens of thousands of birds.
The common name is taken from the male bird's distinctive white stripes underneath their red shoulder patches, or "epaulets", which are visible when the bird is flying or displaying.
Despite the similar names, this bird is not related to the Old World common blackbird, which is a thrush (Turdidae).
The species' call sounds slightly more nasal than that of the red-wing's - a nasal kip and a sharp check.  The male's song is a garbled on-ke-kaaangh.
The bird migrates south during the colder seasons to Mexico and back to northern California during the warmer seasons.


The tricolored blackbird nests in colonies, but scholars disagree on whether the costs outweigh the benefits of these breeding habits. Nonetheless, nesting in colonies makes the tricolor susceptible to environmental changes. Although the tricolor has been able to adapt to some of the landscape changes, habitat loss played a major role in the reduction of its population.
The tricolor originally could be found in the marshes of California, nesting in wetland cattails and bulrushes. Scholars have noted a reduction in freshwater marshes as breeding grounds for the tricolored blackbird. Largely, this is due to human activity, and it’s estimated that between the 1930s and 1980s alone, over 95% of wetlands were disappeared. Also in this period, the observed tricolor population saw an 89% reduction, while the average colony size saw a 63% reduction. Altogether, the tricolor population dropped from several million to only a few hundred thousand during the twentieth century. Nevertheless, the tricolor was able to adapt in response to these severe landscape reductions. It began to use both native and non-native vegetation as well as agricultural fields as their breeding and foraging grounds.
In the 1930s, over 93% of the tricolor’s colonies were nested in freshwater marshes, but by the early twenty-first century, only 35% of colonies could be found in wetlands. In 2008, over a fourth of colonies were using non-native vegetation to nest, most notably Himalayan blackberry brambles. Although population decline was a consequence of agricultural intensification over the twentieth century, the birds were able to use these environmental changes to their advantage. The 2008 tricolored blackbird census found that nearly half of the total population nested in colonies inside of the grain fields of dairy farms.
As the twentieth century progressed, the tricolor began to increase in concentration within certain colonies. Specifically, in 2000, 59% birds were housed in one of California’s ten largest colonies, and this number increased to 81% by 2011. Mega-colonies of the tricolor have begun to form in the San Joaquin Valley’s numerous agricultural fields. Although the tricolor population consistently decreased during the twentieth century and into the twenty-first century, the population nested in San Joaquin Valley grew exponentially during this time. The number of tricolors in California’s San Joaquin Valley in 1994 was 230% of what it was in 1937. By 2008, over 86% of California’s population were located here.
Conservation efforts by the National Audubon Society in collaboration with the U.S. Fish and Wildlife Service have seen an impact on population numbers. The California population of tricolors saw an increase in the 2017 state survey from pervious years. Nonetheless, most of this population gain was within the San Joaquin Valley or in San Benito County. Other regions of California saw a reduction in population in the 2017 census, including Sacramento Valley, with observations down by about 33% from the prior census in 2014. There has been a steady decline of the tricolors observed in this region over the twentieth and twenty-first centuries, with a greater proportion of birds moving to the San Joaquin Valley. There has also been an overall population decline in Southern California as well, following the dairy industry’s move out of Southern California in the 1980s, in favor of the San Joaquin Valley. The other California region that saw a population increase in 2017 was the Central Coast. Historically, the Central Coast and Central Valley housed millions of tricolors before the population decline associated with human agricultural activity. Still, the population increase seen in 2017—from 627 to 17,576 birds—likely was not due to a growing number of tricolors coming to the Central Coast. Instead, this drastic population increase likely was a result of better survey efforts. Three large new colonies were observed in 2017, two of which were in previously unknown or un-surveyed locations. Expanded knowledge of the tricolor’s breeding habits led researchers to increase their survey efforts in locations they suspected housed bird colonies.
Although most regions of California either remained constant or saw a decrease in the tricolor population in 2017, there was an increase in the state overall. The 2017 census, however, did prove that human conservation efforts can be successful. In Southern California, the population remained relatively constant from 2014 to 2017, but the distribution in this region changed. Riverside County saw a significant increase in population, and 57% of the tricolors observed in Southern California were in one single colony. The San Jacinto Wildlife Area in Riverside County had been the site of successful conservation efforts to provide the tricolor with nesting and foraging habitats.


In 1990 the Department of Fish and Game (DFG) of California, based on significant decline in population numbers documented in the 1980s, added the tricolored blackbird to the published list of "Bird Species of Special Concern".  This classification is an "administrative designation intended to alert biologists, land managers and others to a species declining status and encourages them to provide additional management considerations".  At this time the tricolored was added to the U.S. Fish and Wildlife Service (USFWS) list of Birds of Conservation Concern.
By 1991, the tricolor blackbird's breeding population had fallen to approximately thirty-five thousand adults.  This prompted a petition submitted by the Yolo chapter of the National Audubon Society to the California Fish and Game Commission.  The petition brought the tricolored blackbird under consideration for endangered classification.  The classification was granted until the breeding season in 1992.  It was then that researchers discovered a population exceeding three hundred thousand adults.  A petition to withdraw the endangered classification was submitted and accepted.  Conservation measures were supposed to be developed and implemented to avoid a future decline as seen before.  These measures were developed, but only very limited progress was made in preventing the future decline of the tricolored blackbird.  Interested groups started reconvening shortly after the year 2000.  Managing groups found startling results when conducting population research.
In 2006 the tricolored blackbird was classified as Endangered by BirdLife International. Like the extinct passenger pigeon, the colonial nature of the tricolored blackbird makes it particularly vulnerable to extinction. Native grasslands once used for nesting and feeding have been lost to urban and agricultural development. Birds adapted to nesting in agricultural fields have been disturbed by harvesting during the breeding season.  Once the tricolored blackbird was placed on the BirdLife Endangered Species list, it officially became a concern both regionally and nationally.
The U.S. Fish and Wildlife Service maintained the following "Primary conservation priorities for Tricolor habitat conservation and management" (The Tricolored Blackbird Working Group, 2007):
Maintain, enhance, and protect existing habitat suitable for nesting, foraging, and wintering activities;
Create and restore additional protected breeding habitats to support nesting and foraging;
Identify mechanisms for protecting nesting and foraging habitats;
To the extent allowable by law, survey private lands and identify largest and most vulnerable colonies;
Encourage private landowners to protect active breeding colonies; and
Encourage and enhance active breeding colonies on public lands.
In 2018, the tricolored blackbird was listed as a threatened species under the California Endangered Species Act, years after petitioning by the Center for Biological Diversity.


A. conyzoides is prone to becoming a rampant environmental weed when grown outside of its natural range. It is an invasive weed in Africa, Australia, Southeast Asia, Hawaii, and the USA. It is considered a moderate weed of rice cultivation in Asia.


Agkistrodon piscivorus is a species of pit viper in the subfamily Crotalinae of the family Viperidae. It is one of the world's few semiaquatic vipers (along with the Florida cottonmouth), and is native to the southeastern United States. As an adult, it is large and capable of delivering a painful and potentially fatal bite. When threatened, it may respond by coiling its body and displaying its fangs. Individuals may bite when feeling threatened or being handled in any way. It tends to be found in or near water, particularly in slow-moving and shallow lakes, streams, and marshes. It is a capable swimmer and, like several species of snakes, is known to occasionally enter bays and estuaries and swim between barrier islands and the mainland.
The generic name is derived from the Greek words ἄγκιστρον ankistron "fish-hook, hook" and  ὀδών odon  "tooth", and the specific name comes from the Latin  piscis "fish" and voro "(I) eat greedily, devour"; thus, the scientific name translates to "hook-toothed fish-eater". Common names include variants on cottonmouth, water moccasin, swamp moccasin, black moccasin, and simply viper. Many of the common names refer to the threat display, in which this species will often stand its ground and gape at an intruder, exposing the white lining of its mouth. Many scientists dislike the use of the term water moccasin since it can lead to confusion between the venomous cottonmouth and non-venomous water snakes.


In tests designed to measure the various behavioral responses by wild specimens to encounters with people, 23 of 45 (51%) tried to escape, while 28 of 36 (78%) resorted to threat displays and other defensive tactics. Only when they were picked up with a mechanical hand were they likely to bite.
When sufficiently stressed or threatened, this species engages in a characteristic threat display that includes vibrating its tail and throwing its head back with its mouth open to display the startlingly white interior, often making a loud hiss while the neck and front part of the body are pulled into an S-shaped position. Many of its common names, including "cottonmouth" and "gaper", refer to this behavior, while its habit of snapping its jaws shut when anything touches its mouth has earned it the name "trap jaw" in some areas. Other defensive responses can include flattening the body and emitting a strong, pungent secretion from the anal glands located at the base of the tail. This musk may be ejected in thin jets if the snake is sufficiently agitated or restrained. The smell has been likened to that of a billy goat, as well as to a genus of common flood-plain weeds, Pluchea, that also have a penetrating odor.
Harmless watersnakes of the genus Nerodia are often mistaken for it. These are also semiaquatic, thick-bodied snakes with large heads that can be aggressive when provoked, but they behave differently. For example, watersnakes usually flee quickly into the water, while A. piscivorus often stands its ground with its threat display. In addition, watersnakes do not vibrate their tails when excited. A. piscivorus usually holds its head at an angle around 45° when swimming or crawling.
Brown (1973) considered their heavy muscular bodies to be a striking characteristic, stating this made it difficult to hold them for venom extraction owing to their strength.
This species may be active during the day and at night. However, on bright, sunny days, they are usually found coiled or stretched out somewhere in the shade. In the morning and on cool days, they can often be seen basking in the sunlight. They often emerge at sunset to warm themselves on warm ground (i.e., sidewalks, roads) and then become very active throughout the night, when they are usually found swimming or crawling. Contrary to popular belief, they are capable of biting while under water.
In the north, they hibernate during the winter. Niell (1947, 1948) made observations in Georgia, and noted they were one of the last species to seek shelter, often being found active until the first heavy frosts. At this point, they moved to higher ground and could be found in rotting pine stumps by tearing away the bark. These snakes could be quite active upon discovery and would then attempt to burrow more deeply into the soft wood or escape to the nearest water. In southeastern Virginia, Wood (1954) reported seeing migratory behavior in late October and early November. During a period of three or four days, as many as 50 individuals could be seen swimming across Back Bay from the bayside swamps of the barrier islands to the mainland. He suggested this might have something to do with hibernating habits. In the southern parts of its range, hibernation may be short or omitted altogether.


Adult beetles are typically bright metallic green and about 8.5 millimeters (0.33 in) long and 1.6 millimeters (0.063 in) wide. Elytra are typically a darker green, but can also have copper hues. Emerald ash borer is the only North American species of Agrilus with a bright red upper abdomen when viewed with the wings and elytra spread. The species also has a small spine found at the tip of the abdomen and serrate antennae that begin at the fourth antennal segment. They leave tracks in the trees they damage below the bark that are sometimes visible. Adults beetles of other species can often be misidentified by the public.


The emerald ash borer life cycle can occur over one or two years depending on the time of year of oviposition, the health of the tree, and temperature.
After 400–500 accumulated degree-days above 10 °C (50 °F), adults begin to emerge from trees in late spring, and peak emergence occurs around 1,000 degree-days. After emergence, adults feed for one week on ash leaves in the canopy before mating, but cause little defoliation in the process. Males hover around trees, locate females by visual cues, and drop directly onto the female to mate. Mating can last 50 minutes, and females may mate with multiple males over their lifespan. A typical female can live around six weeks and lay approximately 40–70 eggs, but females that live longer can lay up to 200 eggs.
Eggs are deposited between bark crevices, flakes, or cracks and hatch about two weeks later. Eggs are approximately 0.6 to 1.0 millimeter (0.02 to 0.04 in) in diameter, and are initially white, but later turn reddish-brown if fertile. After hatching, larvae chew through the bark to the inner phloem, cambium, and outer xylem where they feed and develop. Emerald ash borer has four larval instars. By feeding, larvae create long serpentine galleries. Fully mature fourth-instar larvae are 26 to 32 millimeters (1.0 to 1.3 in) long. In fall, mature fourth-instars excavate chambers about 1.25 centimeters (0.49 in) into the sapwood or outer bark where they fold into a J-shape. These J-shaped larvae shorten into prepupae and develop into pupae and adults the following spring. To exit the tree, adults chew holes from their chamber through the bark, which leaves a characteristic D-shaped exit hole. Immature larvae can overwinter in their larval gallery, but can require an additional summer of feeding before overwintering again and emerging as adults the following spring. This two-year life cycle is more common in cool climates, such as European Russia.


The native range of the emerald ash borer is temperate north-eastern Asia, which includes Russia, Mongolia, northern China, Japan, and Korea.
The beetle is invasive in North America where it has a core population in Michigan and surrounding states and provinces. Populations are more scattered outside the core area, and the edges of its known distribution range north to Ontario, south to northern Louisiana, west to Colorado, and east to New Brunswick, and in the Pacific Northwest in Oregon. In eastern Europe, a population was found in Moscow in 2003. From 2003 to 2016, this population has spread west towards the European Union at up to 40 km (25 mi) per year and is expected to reach central Europe between 2031 and 2036. Although not recorded from the European Union as of 2019, it has already spread to far eastern Ukraine from neighboring Russia.


In its native range, emerald ash borer is only a nuisance pest on native trees, as population densities typically do not reach levels lethal to healthy trees. In China, it infests native F. chinensis, F. mandshurica, and F. rhynchophylla; in Japan it also infests F. japonica and F. lanuginosa.
Emerald ash borer primarily infest and can cause significant damage to ash species including green ash (Fraxinus pennsylvanica), black ash (Fraxinus nigra), white ash (Fraxinus americana), and blue ash (Fraxinus quadrangulata) in North America. In Europe, Fraxinus excelsior is the main ash species colonized, which is moderately resistant to emerald ash borer infestation. Ash susceptibility can vary depending on the attractiveness of chemical volatiles to adults, or the ability of larvae to detoxify phenolic compounds. Emerald ash borer has also been found infesting white fringe tree in North America, which is a non-ash host, but it is unclear whether the trees were healthy when first infested, or were already in decline because of drought. Another non-ash host has also been discovered, Olea europaea, albeit in a lab setting.
Adults prefer to lay eggs on open grown or stressed ash but readily lay eggs on healthy trees amongst other tree species. Ashes that grow in pure stands, whether naturally occurring or in landscaping, are more prone to attack than isolated trees or ones located in mixed forest stands. Ashes used in landscaping also tend to be subjected to higher amounts of environmental stresses including compacted soil, lack of moisture, heating effects from urban islands, road salt, and pollution, which may also reduce their resistance to the borer. Furthermore, most ashes used in landscaping were produced from a handful of cultivars, resulting in low genetic diversity. Young trees with bark between 1.5 millimeters (0.059 in) to 5 millimeters (0.20 in) are preferred. Both males and females use leaf volatiles and sesquiterpenes in the bark to locate hosts. Damage occurs in infested trees by larval feeding. The serpentine feeding galleries of the larvae disrupt the flow of nutrients and water, effectively girdling, thus killing the tree, as it is no longer able to transport sufficient water and nutrients to the leaves to survive. Girdled ashes will often attempt to regenerate through stump sprouting, and there is evidence that stressed trees may also generate higher than normal seed crops as an emergency measure.


Outside its native range, emerald ash borer is an invasive species that is highly destructive to ash trees in its introduced range. Before emerald ash borer was found in North America, very little was known about the insect in its native range aside from a short description of life-history traits and taxonomic descriptions, which resulted in focused research on its biology in North America. The insect was first identified in Canton, Michigan (near Detroit), in 2002, but it may have been in the U.S. since the late 1980s. It is suspected that it was introduced from overseas in shipping materials such as packing crates.
Without factors that would normally suppress emerald ash borer populations in its native range (e.g., resistant trees, predators, and parasitoid wasps), populations can quickly rise to damaging levels. After initial infestation, all ash trees are expected to die in an area within 10 years without control measures. Every North American ash species has susceptibility to emerald ash borer, as North American species planted in China also have high mortality from infestations, but some Chinese ash species are resistant.
Green ash and black ash trees are preferred by emerald ash borer. White ash is also killed rapidly but usually only after all green and black ash trees are eliminated. Blue ash is known to exhibit a higher degree of resistance to emerald ash borer, which is believed to be caused by the high tannin content in the leaves making the foliage unpalatable to the insect. While most Asian ashes have evolved this defense, it is absent from American species other than blue ash. Researchers have examined populations of so-called "lingering ash", trees that survived ash borer attack with little or no damage, as a means of grafting or breeding new, resistant stock. Many of these lingering ashes were found to have unusual phenotypes that may result in increased resistance. Aside from their higher tannin content, Asian ashes also employ natural defenses to repel, trap, and kill emerald ash borer larvae. Although studies of American ashes have suggested that they are capable of mustering similar defensive mechanisms, the trees do not appear to recognize when they are under attack. Many of the specialized predators and parasitoids that suppressed emerald ash borer in Asia were not present in North America. Predators and parasitoids native to North America do not sufficiently suppress emerald ash borer, so populations continue to grow. Birds such as woodpeckers feed on emerald ash borer larva, although the adult beetles have not been used by any American fauna as food. Emerald ash borer populations can spread between 2.5 to 20 km (1.6 to 12.4 mi) per year. It primarily spreads through flight or by transportation of ash bark containing products such as firewood or nursery stock, which allows it to reach new areas and create satellite populations outside of the main infestation.
Other factors can limit spread. Winter temperatures of approximately −38 °C (−36 °F) limit range expansion., and overwintering emerald ash borer survive down to average temperatures of −30 °C (−22 °F) because of antifreeze chemicals in the body and insulation provided by tree bark. Larvae can also survive high heat up to 53 °C (127 °F). Conversely, much like ashes grown in the nursery trade, the population of emerald ash borer in North America is believed to have originated from a single group of insects from central China and also exhibits low genetic diversity.
North American predators and parasitoids can occasionally cause high emerald ash borer mortality, but generally offer only limited control. Mortality from native woodpeckers is variable. Parasitism by parasitoids such as Atanycolus cappaerti can be high, but overall such control is generally low.
The United States Department of Agriculture's Animal and Plant Health Inspection Service published a rule on December 14, 2020—to take effect one month later, January 14, 2021—ending all EAB quarantine activities in the United States due to ineffectiveness so far. Other means will be used instead, especially biological controls (see §Biological control below).


In areas where emerald ash borer has not yet been detected, surveys are used to monitor for new infestations. Visual surveys are used to find ash trees displaying emerald ash borer damage, and traps with colors attractive to emerald ash borer, such as purple or green, are hung in trees as part of a monitoring program. These traps can also have volatile pheromones applied to them that attract primarily males.
Sometimes trees are girdled to act as trap trees to monitor for emerald ash borer. The stressed tree attracts egg-laying females in the spring, and trees can be debarked in the fall to search for larvae. If detected, an area is often placed under a quarantine to prevent infested wood material from causing new infestations. Further control measures are then taken within the area to slow population growth by reducing beetle numbers, preventing them from reaching reproductive maturity and dispersing, and reducing the abundance of ash trees.
Government agencies in both the U.S. and Canada have utilized a native species of parasitoid wasp, Cerceris fumipennis, as a means of detecting areas to which emerald ash borer has spread. The females of these wasps hunt other jewel beetles and emerald ash borer if it is present. The wasps stun the beetles and carry them back to their burrows in the ground where they are stored until the wasps’ eggs hatch and the wasp larvae feed on the beetles. Volunteers catch the wasps as they return to their burrows carrying the beetles to determine whether emerald ash borer is present. This methodology is known as biological surveillance, as opposed to biological control, because it does not appear that the wasps have a significant negative impact on emerald ash borer populations.


Once an infestation is detected, quarantines are typically imposed by state or national government agencies disallowing transport of ash firewood or live plants outside of these areas without permits indicating the material has been inspected or treated (i.e., heat treatment or wood chipping) to ensure no live emerald ash borer are present in the bark and phloem. In urban areas, trees are often removed once an infestation is found to reduce emerald ash borer population densities and the likelihood of further spread. Urban ash are typically replaced with non-ash species such as maple, oak, or linden to limit food sources. In rural areas, trees can be harvested for lumber or firewood to reduce ash stand density, but quarantines may apply for this material, especially in areas where the material could be infested.
Kentucky Extension specialists suggest selecting uncommon species to replace removed ashes in the landscape. Previous generations created monocultures by planting ash trees in an overabundance, a factor in the extent of the devastation caused by the emerald ash borer. Favoring instead a diversity in species helps keep urban forests healthy. University of Kentucky scientists suggest choosing monotypic species such as the pawpaw, yellowwood, Franklin tree, Kentucky coffeetree, Osage orange, sourwood, and bald cypress.


Insecticides with active ingredients such as azadirachtin, imidacloprid,  emamectin benzoate, and dinotefuran are currently used. Dinotefuran and imidacloprid are systemic (i.e., incorporated into the tree) and remain effective for one to three years depending on the product. Insecticides are typically only considered a viable option in urban areas with high value trees near an infestation. Ash trees are primarily treated by direct injection into the tree or soil drench. Some insecticides cannot be applied by homeowners and must be applied by licensed applicators. Damage from emerald ash borer can continue to increase over time even with insecticide applications. Insecticide treatments are not feasible for large forested areas outside of urban areas.


Agrimonia pilosa is traditionally used in Korea for boils, eczema, and taeniasis (a tape worm condition). In Nepal and China, A. pilosa is traditionally used for abdominal pain, sore throat, headaches, and heat stroke.


Agriphila vulgivagellus, the vagabond crambus or vagabond sod webworm, is a moth of the family Crambidae. It is found from Quebec and New England to Florida, west to Texas and north to Alberta.
The wingspan is 20–39 mm. Adults are on wing from August to October in one generation per year.
The larvae feed on various grasses, as well as wheat, rye and other grains. Immature larvae overwinter.


Agropyron cristatum is one of several closely related grass species referred to as crested wheatgrass. It is unable to hybridize with its similar relatives, as it is a diploid species, whereas its closest relative, Agropyron desertorum, is a tetraploid species. It was introduced from Russia and Siberia to North America in the first half of the twentieth century, and widely used to reseed abandoned marginal cropland undergoing varying degrees of soil erosion and secondary succession. A. cristatum is very long lived, with stands often remaining productive for 30 years or more.