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Sea otter. The sea otter (Enhydra lutris) is a marine mammal native to the coasts of the northern and eastern North Pacific Ocean. Adult sea otters typically weigh between 14 and 45 (kg), making them the heaviest members of the weasel family, but among the smallest marine mammals. Unlike most marine mammals, the sea otter's primary form of insulation is an exceptionally thick coat of fur, the densest in the animal kingdom. Although it can walk on land, the sea otter is capable of living exclusively in the ocean.The sea otter inhabits nearshore environments, where it dives to the sea floor to.

Sea otter. were once estimated at 150,000–300,000, were hunted extensively for their fur between 1741 and 1911, and the world population fell to 1,000–2,000 individuals living in a fraction of their historic range. A subsequent international ban on hunting, sea otter conservation efforts, and reintroduction programs into previously populated areas have contributed to numbers rebounding, and the species occupies about two-thirds of its former range. The recovery of the sea otter is considered an important success in marine conservation, although populations in the Aleutian Islands and California have recently declined or have plateaued at depressed levels. For these reasons, the sea otter.

Sea otter. remains classified as an endangered species.## Evolution.The sea otter is the heaviest (the giant otter is longer, but significantly slimmer) member of the family Mustelidae, a diverse group that includes the 13 otter species and terrestrial animals such as weasels, badgers, and minks. It is unique among the mustelids in not making or burrows, in having no functional anal scent glands, and in being able to live its entire life without leaving the water. The only living member of the genus "Enhydra", the sea otter is so different from other mustelid species that, as recently as 1982, some scientists believed.

Sea otter. spread east to the Aleutian Islands, mainland Alaska, and down the North American coast. In comparison to cetaceans, sirenians, and pinnipeds, which entered the water approximately 50, 40, and 20 million years ago, respectively, the sea otter is a relative newcomer to a marine existence. In some respects, though, the sea otter is more fully adapted to water than pinnipeds, which must haul out on land or ice to give birth. The full genome of the northern sea otter ("Enhydra lutris kenyoni") was sequenced in 2017 and may allow for examination of the sea otter's evolutionary divergence from terrestrial mustelids.##.

Sea otter. Taxonomy.The first scientific description of the sea otter is contained in the field notes of Georg Steller from 1751, and the species was described by Carl Linnaeus in his landmark 1758 10th edition of "Systema Naturae". Originally named "Lutra marina", it underwent numerous name changes before being accepted as "Enhydra lutris" in 1922. The generic name "Enhydra", derives from the Ancient Greek "en"/εν "in" and "hydra"/ύδρα "water", meaning "in the water", and the Latin word "lutris", meaning "otter". It was formerly sometimes referred to as the "sea beaver".## Subspecies.Three subspecies of the sea otter are recognized with distinct geographical distributions.

Sea otter. Description.The sea otter is one of the smallest marine mammal species, but it is the heaviest mustelid. Male sea otters usually weigh 22 to 45 (kg) and are 1.2 to 1.5 (m) in length, though specimens up to 54 (kg) have been recorded. Females are smaller, weighing 14 to 33 (kg) and measuring 1.0 to 1.4 (m) in length. For its size, the male otter's baculum is very large, massive and bent upwards, measuring 150 (mm) in length and 15 (mm) at the base.Unlike most other marine mammals, the sea otter has no blubber and relies on its exceptionally thick.

Sea otter. fur to keep warm. With up to 150000 (/cm2), its fur is the densest of any animal. The fur consists of long, waterproof guard hairs and short underfur; the guard hairs keep the dense underfur layer dry. There is an air compartment between the thick fur and the skin where air is trapped and heated by the body. Cold water is kept completely away from the skin and heat loss is limited. However, this air compartment can be disadvantageous because as the sea otter dives deeper into the water column, the air compartment compresses and heat needed to warm the.

Sea otter. the urchin's shell.The sea otter's use of rocks when hunting and feeding makes it one of the few mammal species to use tools. To open hard shells, it may pound its prey with both paws against a rock on its chest. To pry an abalone off its rock, it hammers the abalone shell using a large stone, with observed rates of 45 blows in 15 seconds. Releasing an abalone, which can cling to rock with a force equal to 4,000 times its own body weight, requires multiple dives.## Social structure.Although each adult and independent juvenile forages alone, sea otters tend.

Sea otter. to rest together in single-sex groups called "rafts". A raft typically contains 10 to 100 animals, with male rafts being larger than female ones. The largest raft ever seen contained over 2000 sea otters. To keep from drifting out to sea when resting and eating, sea otters may wrap themselves in kelp.A male sea otter is most likely to mate if he maintains a breeding territory in an area that is also favored by females. As autumn is the peak breeding season in most areas, males typically defend their territory only from spring to autumn. During this time, males patrol.

Sea otter. jaws during copulation. Visible scars are often present on females from this behavior.Births occur year-round, with peaks between May and June in northern populations and between January and March in southern populations. Gestation appears to vary from four to twelve months, as the species is capable of delayed implantation followed by four months of pregnancy. In California, sea otters usually breed every year, about twice as often as those in Alaska.Birth usually takes place in the water and typically produces a single pup weighing 1.4 to 2.3 (kg). Twins occur in 2% of births; however, usually only one pup survives.

Sea otter. At birth, the eyes are open, ten teeth are visible, and the pup has a thick coat of baby fur. Mothers have been observed to lick and fluff a newborn for hours; after grooming, the pup's fur retains so much air, the pup floats like a cork and cannot dive. The fluffy baby fur is replaced by adult fur after about 13 weeks.Nursing lasts six to eight months in Californian populations and four to twelve months in Alaska, with the mother beginning to offer bits of prey at one to two months. The milk from a sea otter's two abdominal.

Sea otter. water, sometimes wrapped in kelp to keep it from floating away; if the pup is not sleeping, it cries loudly until she returns. Mothers have been known to carry their pups for days after the pups' deaths.Females become sexually mature at around three or four years of age and males at around five; however, males often do not successfully breed until a few years later. A captive male sired offspring at age 19. In the wild, sea otters live to a maximum age of 23 years, with lifespans ranging from 10 to 15 years for males and 15–20 years for.

Sea otter. females. Several captive individuals have lived past 20 years, and a female at the Seattle Aquarium died at the age of 28 years. Sea otters in the wild often develop worn teeth, which may account for their apparently shorter lifespans.## Population and distribution.Sea otters live in coastal waters 15 to 23 (m) deep, and usually stay within a kilometre ( mi) of the shore. They are found most often in areas with protection from the most severe ocean winds, such as rocky coastlines, thick kelp forests, and barrier reefs. Although they are most strongly associated with rocky substrates, sea otters.

Sea otter. can also live in areas where the sea floor consists primarily of mud, sand, or silt. Their northern range is limited by ice, as sea otters can survive amidst drift ice but not land-fast ice. Individuals generally occupy a home range a few kilometres long, and remain there year-round.The sea otter population is thought to have once been 150,000 to 300,000, stretching in an arc across the North Pacific from northern Japan to the central Baja California Peninsula in Mexico. The fur trade that began in the 1740s reduced the sea otter's numbers to an estimated 1,000 to 2,000 members.

Sea otter. in 13 colonies. Hunting records researched by historian Adele Ogden place the westernmost limit of the hunting grounds off the northern Japanese island of Hokkaido and the easternmost limit off Punta Morro Hermosa about 21+1/2 (mi) south of Punta Eugenia, Baja California's westernmost headland in Mexico.In about two-thirds of its former range, the species is at varying levels of recovery, with high population densities in some areas and threatened populations in others. Sea otters currently have stable populations in parts of the Russian east coast, Alaska, British Columbia, Washington, and California, with reports of recolonizations in Mexico and Japan. Population.

Sea otter. estimates made between 2004 and 2007 give a worldwide total of approximately 107,000 sea otters.## Japan.Adele Ogden wrote in "The California Sea Otter Trade" that sea otter were hunted "from Yezo northeastward past the Kuril Group and Kamchatka to the Aleutian Chain". "Yezo" refers to the island of Hokkaido in northern Japan; the only confirmed sea otter population in Japanese territory is on the coast surrounding the town of Erimo, Hokkaido.## Russia.Currently, the most stable and secure part of the sea otter's range is Russia. Before the 19th century, around 20,000 to 25,000 sea otters lived near the Kuril Islands,.

Sea otter. with more near Kamchatka and the Commander Islands. After the years of the Great Hunt, the population in these areas, currently part of Russia, was only 750. By 2004, sea otters had repopulated all of their former habitat in these areas, with an estimated total population of about 27,000. Of these, about 19,000 are at the Kurils, 2,000 to 3,500 at Kamchatka and another 5,000 to 5,500 at the Commander Islands. Growth has slowed slightly, suggesting the numbers are reaching carrying capacity.## British Columbia.Along the North American coast south of Alaska, the sea otter's range is discontinuous. A remnant population.

Sea otter. survived off Vancouver Island into the 20th century, but it died out despite the 1911 international protection treaty, with the last sea otter taken near Kyuquot in 1929. From 1969 to 1972, 89 sea otters were flown or shipped from Alaska to the west coast of Vancouver Island. This population increased to over 5,600 in 2013 with an estimated annual growth rate of 7.2%, and their range on the island's west coast extended north to Cape Scott and across the Queen Charlotte Strait to the Broughton Archipelago and south to Clayoquot Sound and Tofino. In 1989, a separate colony was.

Sea otter. discovered in the central British Columbia coast. It is not known if this colony, which numbered about 300 animals in 2004, was founded by transplanted otters or was a remnant population that had gone undetected. By 2013, this population exceeded 1,100 individuals, was increasing at an estimated 12.6% annual rate, and its range included Aristazabal Island, and Milbanke Sound south to Calvert Island. In 2008, Canada determined the status of sea otters to be "special concern".## United States.## Alaska.Alaska is the central area of the sea otter's range. In 1973, the population in Alaska was estimated at between 100,000 and.

Sea otter. 125,000 animals. By 2006, though, the Alaska population had fallen to an estimated 73,000 animals. A massive decline in sea otter populations in the Aleutian Islands accounts for most of the change; the cause of this decline is not known, although orca predation is suspected. The sea otter population in Prince William Sound was also hit hard by the Exxon Valdez oil spill, which killed thousands of sea otters in 1989.## Washington.In 1969 and 1970, 59 sea otters were translocated from Amchitka Island to Washington, and released near La Push and Point Grenville. The translocated population is estimated to have.

Sea otter. declined to between 10 and 43 individuals before increasing, reaching 208 individuals in 1989. As of 2017, the population was estimated at over 2,000 individuals, and their range extends from Point Grenville in the south to Cape Flattery in the north and east to Pillar Point along the Strait of Juan de Fuca.In Washington, sea otters are found almost exclusively on the outer coasts. They can swim as close as six feet off shore along the Olympic coast. Reported sightings of sea otters in the San Juan Islands and Puget Sound almost always turn out to be North American river.

Sea otter. otters, which are commonly seen along the seashore. However, biologists have confirmed isolated sightings of sea otters in these areas since the mid-1990s.## Oregon.The last native sea otter in Oregon was probably shot and killed in 1906. In 1970 and 1971, a total of 95 sea otters were transplanted from Amchitka Island, Alaska to the Southern Oregon coast. However, this translocation effort failed and otters soon again disappeared from the state. In 2004, a male sea otter took up residence at Simpson Reef off of Cape Arago for six months. This male is thought to have originated from a colony.

Sea otter. in Washington, but disappeared after a coastal storm. On 18 February 2009, a male sea otter was spotted in Depoe Bay off the Oregon Coast. It could have traveled to the state from either California or Washington.## California.The historic population of California sea otters was estimated at 16,000 before the fur trade decimated the population, leading to their assumed extinction. Today's population of California sea otters are the descendants of a single colony of about 50 sea otters located near Bixby Creek Bridge in March 1938 by Howard G. Sharpe, owner of the nearby Rainbow Lodge on Bixby Bridge in.

Sea otter. Big Sur. Their principal range has gradually expanded and extends from Pigeon Point in San Mateo County to Santa Barbara County.Sea otters were once numerous in San Francisco Bay. Historical records revealed the Russian-American Company sneaked Aleuts into San Francisco Bay multiple times, despite the Spanish capturing or shooting them while hunting sea otters in the estuaries of San Jose, San Mateo, San Bruno and around Angel Island. The founder of Fort Ross, Ivan Kuskov, finding otters scarce on his second voyage to Bodega Bay in 1812, sent a party of Aleuts to San Francisco Bay, where they met another.

Sea otter. Russian party and an American party, and caught 1,160 sea otters in three months. By 1817, sea otters in the area were practically eliminated and the Russians sought permission from the Spanish and the Mexican governments to hunt further and further south of San Francisco. Remnant sea otter populations may have survived in the bay until 1840, when the Rancho Punta de Quentin was granted to Captain John B. R. Cooper, a sea captain from Boston, by Mexican Governor Juan Bautista Alvarado along with a license to hunt sea otters, reportedly then prevalent at the mouth of Corte Madera Creek.In.

Sea otter. the late 1980s, the USFWS relocated about 140 southern sea otters to San Nicolas Island in southern California, in the hope of establishing a reserve population should the mainland be struck by an oil spill. To the surprise of biologists, the majority of the San Nicolas sea otters swam back to the mainland. Another group of twenty swam 74 (mi) north to San Miguel Island, where they were captured and removed. By 2005, only 30 sea otters remained at San Nicolas, although they were slowly increasing as they thrived on the abundant prey around the island. The plan that authorized.

Sea otter. the translocation program had predicted the carrying capacity would be reached within five to 10 years. The spring 2016 count at San Nicolas Island was 104 sea otters, continuing a 5-year positive trend of over 12% per year. Sea otters were observed twice in Southern California in 2011, once near Laguna Beach and once at Zuniga Point Jetty, near San Diego. These are the first documented sightings of otters this far south in 30 years.When the USFWS implemented the translocation program, it also attempted, in 1986, to implement "zonal management" of the Californian population. To manage the competition between sea.

Sea otter. response to lawsuits filed by the Santa Barbara-based Environmental Defense Center and the Otter Project, on 19 December 2012 the USFWS declared that the "no otter zone" experiment was a failure, and will protect the otters re-colonizing the coast south of Point Conception as threatened species. Although abalone fisherman blamed the incursions of sea otters for the decline of abalone, commercial abalone fishing in southern California came to an end from overfishing in 1997, years before significant otter moved south of Point Conception. In addition, white abalone ("Haliotis sorenseni"), a species never overlapping with sea otter, had declined in numbers.

Sea otter. 99% by 1996, and became the first marine invertebrate to be federally listed as endangered.Although the southern sea otter's range has continuously expanded from the remnant population of about 50 individuals in Big Sur since protection in 1911, from 2007 to 2010, the otter population and its range contracted and since 2010 has made little progress. As of spring 2010, the northern boundary had moved from about Tunitas Creek to a point 2 (km) southeast of Pigeon Point, and the southern boundary has moved along the Gaviota Coast from approximately Coal Oil Point to Gaviota State Park. A toxin called.

Sea otter. microcystin, produced by a type of cyanobacteria ("Microcystis"), seems to be concentrated in the shellfish the otters eat, poisoning them. Cyanobacteria are found in stagnant water enriched with nitrogen and phosphorus from septic tank and agricultural fertilizer runoff, and may be flushed into the ocean when streamflows are high in the rainy season. A record number of sea otter carcasses were found on California's coastline in 2010, with increased shark attacks an increasing component of the mortality. Great white sharks do not consume relatively fat-poor sea otters but shark-bitten carcasses have increased from 8% in the 1980s to 15% in.

Sea otter. the 1990s and to 30% in 2010 and 2011.For southern sea otters to be considered for removal from threatened species listing, the U.S. Fish and Wildlife Service (USFWS) determined that the population should exceed 3,090 for three consecutive years. In response to recovery efforts, the population climbed steadily from the mid-20th century through the early 2000s, then remained relatively flat from 2005 to 2014 at just under 3,000. There was some contraction from the northern (now Pigeon Point) and southern limits of the sea otter's range during the end of this period, circumstantially related to an increase in lethal shark.

Sea otter. bites, raising concerns that the population had reached a plateau. However, the population increased markedly from 2015 to 2016, with the United States Geological Survey (USGS) California sea otter survey 3-year average reaching 3,272 in 2016, the first time it exceeded the threshold for delisting from the Endangered Species Act (ESA). If populations continued to grow and ESA delisting occurred, southern sea otters would still be fully protected by state regulations and the Marine Mammal Protection Act, which set higher thresholds for protection, at approximately 8,400 individuals. However, ESA delisting seems unlikely due to a precipitous population decline recorded in.

Sea otter. the spring 2017 USGS sea otter survey count, from the 2016 high of 3,615 individuals to 2,688, a loss of 25% of the California sea otter population.## Mexico.Historian Adele Ogden described sea otters are particularly abundant in "Lower California", now the Baja California Peninsula, where "seven bays...were main centers". The southernmost limit was Punta Morro Hermoso about 21+1/2 (mi) south of Punta Eugenia, in turn a headland at the southwestern end of Sebastián Vizcaíno Bay, on the west coast of the Baja Peninsula. Otter were also taken from San Benito Island, Cedros Island, and Isla Natividad in the Bay. By.

Sea otter. the early 1900s, Baja's sea otters were extirpated by hunting. In a 1997 survey, small numbers of sea otters, including pups, were reported by local fishermen, but scientists could not confirm these accounts. However, male and female otters have been confirmed by scientists off shores of the Baja Peninsula in a 2014 study, who hypothesize that otter dispersed there beginning in 2005. These sea otters may have dispersed from San Nicolas Island, which is 300 (km) away, as individuals have been recorded traversing distances of over 800 (km). Genetic analysis of most of these animals were consistent with California, i.e.

Sea otter. United States, otter origins, however one otter had a haplotype not previously reported, and could represent a remnant of the original native Mexican otter population.## Ecology.## Diet.High energetic requirements of sea otter metabolism require them to consume at least 20% of their body weight a day. Surface swimming and foraging are major factors in their high energy expenditure due to drag on the surface of the water when swimming and the thermal heat loss from the body during deep dives when foraging. Sea otter muscles are specially adapted to generate heat without physical activity.Sea otters consume over 100 prey species.

Sea otter. also eaten. In studies performed at Amchitka Island in the 1960s, where the sea otter population was at carrying capacity, 50% of food found in sea otter stomachs was fish. The fish species were usually bottom-dwelling and sedentary or sluggish forms, such as "Hemilepidotus hemilepidotus" and family Tetraodontidae. However, south of Alaska on the North American coast, fish are a negligible or extremely minor part of the sea otter's diet. Contrary to popular depictions, sea otters rarely eat starfish, and any kelp that is consumed apparently passes through the sea otter's system undigested.The individuals within a particular area often differ.

Sea otter. and not eat it. On land, young sea otters may face attack from bears and coyotes. In California, great white sharks are their primary predator but there is no evidence that the sharks eat them.Urban runoff transporting cat feces into the ocean brings "Toxoplasma gondii", an obligate parasite of felids, which has killed sea otters. Parasitic infections of "Sarcocystis neurona" are also associated with human activity. According to the U.S. Geological Survey and the CDC, northern sea otters off Washington have been infected with the H1N1 flu virus and "may be a newly identified animal host of influenza viruses".## Relationship.

Sea otter. with humans.## Fur trade.Sea otters have the thickest fur of any mammal, which makes them a common target for many hunters. Archaeological evidence indicates that for thousands of years, indigenous peoples have hunted sea otters for food and fur. Large-scale hunting, part of the Maritime Fur Trade, which would eventually kill approximately one million sea otters, began in the 18th century when hunters and traders began to arrive from all over the world to meet foreign demand for otter pelts, which were one of the world's most valuable types of fur.In the early 18th century, Russians began to hunt sea.

Sea otter. otters in the Kuril Islands and sold them to the Chinese at Kyakhta. Russia was also exploring the far northern Pacific at this time, and sent Vitus Bering to map the Arctic coast and find routes from Siberia to North America. In 1741, on his second North Pacific voyage, Bering was shipwrecked off Bering Island in the Commander Islands, where he and many of his crew died. The surviving crew members, which included naturalist Georg Steller, discovered sea otters on the beaches of the island and spent the winter hunting sea otters and gambling with otter pelts. They returned to.

Sea otter. Siberia, having killed nearly 1,000 sea otters, and were able to command high prices for the pelts. Thus began what is sometimes called the "Great Hunt", which would continue for another hundred years. The Russians found the sea otter far more valuable than the sable skins that had driven and paid for most of their expansion across Siberia. If the sea otter pelts brought back by Bering's survivors had been sold at Kyakhta prices they would have paid for one tenth the cost of Bering's expedition.Russian fur-hunting expeditions soon depleted the sea otter populations in the Commander Islands, and by.

Sea otter. 1745, they began to move on to the Aleutian Islands. The Russians initially traded with the Aleuts inhabitants of these islands for otter pelts, but later enslaved the Aleuts, taking women and children hostage and torturing and killing Aleut men to force them to hunt. Many Aleuts were either murdered by the Russians or died from diseases the hunters had introduced. The Aleut population was reduced, by the Russians' own estimate, from 20,000 to 2,000. By the 1760s, the Russians had reached Alaska. In 1799, Tsar Paul I consolidated the rival fur-hunting companies into the Russian-American Company, granting it an.

Sea otter. imperial charter and protection, and a monopoly over trade rights and territorial acquisition.Under Aleksander I, the administration of the merchant-controlled company was transferred to the Imperial Navy, largely due to the alarming reports by naval officers of native abuse; in 1818, the indigenous peoples of Alaska were granted civil rights equivalent to a townsman status in the Russian Empire.Other nations joined in the hunt in the south. Along the coasts of what is now Mexico and California, Spanish explorers bought sea otter pelts from Native Americans and sold them in Asia. In 1778, British explorer Captain James Cook reached Vancouver.

Sea otter. Island and bought sea otter furs from the First Nations people. When Cook's ship later stopped at a Chinese port, the pelts rapidly sold at high prices, and were soon known as "soft gold". As word spread, people from all over Europe and North America began to arrive in the Pacific Northwest to trade for sea otter furs.Russian hunting expanded to the south, initiated by American ship captains, who subcontracted Russian supervisors and Aleut hunters in what are now Washington, Oregon, and California. Between 1803 and 1846, 72 American ships were involved in the otter hunt in California, harvesting an.

Sea otter. estimated 40,000 skins and tails, compared to only 13 ships of the Russian-American Company, which reported 5,696 otter skins taken between 1806 and 1846. In 1812, the Russians founded an agricultural settlement at what is now Fort Ross in northern California, as their southern headquarters.Eventually, sea otter populations became so depleted, commercial hunting was no longer viable. It had stopped in the Aleutian Islands, by 1808, as a conservation measure imposed by the Russian-American Company. Further restrictions were ordered by the company in 1834. When Russia sold Alaska to the United States in 1867, the Alaska population had recovered to.

Sea otter. over 100,000, but Americans resumed hunting and quickly extirpated the sea otter again. Prices rose as the species became rare. During the 1880s, a pelt brought $105 to $165 in the London market, but by 1903, a pelt could be worth as much as $1,125. In 1911, Russia, Japan, Great Britain (for Canada) and the United States signed the Treaty for the Preservation and Protection of Fur Seals, imposing a moratorium on the harvesting of sea otters. So few remained, perhaps only 1,000–2,000 individuals in the wild, that many believed the species would become extinct.## Recovery and conservation.During the 20th.

Sea otter. Soviet Union in 1991; however, it has declined significantly with stricter law enforcement and better economic conditions.The most significant threat to sea otters is oil spills, to which they are particularly vulnerable, since they rely on their fur to keep warm. When their fur is soaked with oil, it loses its ability to retain air, and the animals can quickly die from hypothermia. The liver, kidneys, and lungs of sea otters also become damaged after they inhale oil or ingest it when grooming. The Exxon Valdez oil spill of 24 March 1989 killed thousands of sea otters in Prince William.

Sea otter. an oil spill wiping out a population. However, because of the species' reputation for depleting shellfish resources, advocates for commercial, recreational, and subsistence shellfish harvesting have often opposed allowing the sea otter's range to increase, and there have even been instances of fishermen and others illegally killing them.In the Aleutian Islands, a massive and unexpected disappearance of sea otters has occurred in recent decades. In the 1980s, the area was home to an estimated 55,000 to 100,000 sea otters, but the population fell to around 6,000 animals by 2000. The most widely accepted, but still controversial, hypothesis is that killer.

Sea otter. whales have been eating the otters. The pattern of disappearances is consistent with a rise in predation, but there has been no direct evidence of orcas preying on sea otters to any significant extent.Another area of concern is California, where recovery began to fluctuate or decline in the late 1990s. Unusually high mortality rates amongst adult and subadult otters, particularly females, have been reported. In 2017 the US Geological Survey found a 3% drop in the sea otter population of the California coast. This number still keeps them on track for removal from the endangered species list, although just barely.

Sea otter. States, Russia and Canada. In marine protected areas, polluting activities such as dumping of waste and oil drilling are typically prohibited. An estimated 1,200 sea otters live within the Monterey Bay National Marine Sanctuary, and more than 500 live within the Olympic Coast National Marine Sanctuary.## Economic impact.Some of the sea otter's preferred prey species, particularly abalone, clams, and crabs, are also food sources for humans. In some areas, massive declines in shellfish harvests have been blamed on the sea otter, and intense public debate has taken place over how to manage the competition between sea otters and humans for.

Sea otter. greatly depleted, no longer rely as intimately on sea mammals for survival.Since the mid-1970s, the beauty and charisma of the species have gained wide appreciation, and the sea otter has become an icon of environmental conservation. The round, expressive face and soft, furry body of the sea otter are depicted in a wide variety of souvenirs, postcards, clothing, and stuffed toys.## Aquariums and zoos.Sea otters can do well in captivity, and are featured in over 40 public aquariums and zoos. The Seattle Aquarium became the first institution to raise sea otters from conception to adulthood with the birth of Tichuk.

Sea otter. in 1979, followed by three more pups in the early 1980s. In 2007, a YouTube video of two sea otters holding paws drew 1.5 million viewers in two weeks, and had over 22 million views . Filmed five years previously at the Vancouver Aquarium, it was YouTube's most popular animal video at the time, although it has since been surpassed. The lighter-colored otter in the video is Nyac, a survivor of the 1989 "Exxon Valdez" oil spill. Nyac died in September 2008, at the age of 20. Milo, the darker one, died of lymphoma in January 2012. California Fur Rush.

Sea otter. "Enhydra lutris" (Linnaeus, 1758) at the Integrated Taxonomic Information System "Enhydra lutris" (Linnaeus, 1758) at the World Register of Marine Species "De Bestiis Marinis, or, The Beasts of the Sea" (1751) (PDF), pp. 68–82, transcribed field notes from 18th-century German zoologist Georg Wilhelm Steller Precipice of Survival: The Southern Sea Otter (Adobe Flash), a 48-minute program on the southern sea otter's history by the United States Geological Survey.


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Römerberg. Römerberg is a municipality in the Rhein-Pfalz-Kreis, in Rhineland-Palatinate, Germany. It is situated on the left bank of the Rhine, approximately 5 (km) southwest of Speyer.## History.The municipality of Römerberg was established by the fusion of the municipalities of Berghausen, Heiligenstein and Mechtersheim in 1969. The seat of the municipality is in the village Heiligenstein.


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Bedside. Bedside is a 1934 American pre-Code drama film starring Warren William, Jean Muir and Allen Jenkins. A man passes himself off as a doctor. Warren William as Bob Brown Jean Muir as Caroline Grant Allen Jenkins as Sam Sparks David Landau as Smith Katharine Sergava (Kathryn Sergava) as Mimi Maritza Henry O'Neill as Dr. William Chester Donald Meek as Dr. George Wiley Renee Whitney as Mme. Varsova Walter Walker as Dr. Michaels Marjorie Lytell as Patient with Sprained Ankle Frederick Burton as Hospital Superintendent Philip Faversham as Intern Attending Caroline (as Phillip Faversham) Louise Beavers as Pansy Earle Foxe as.


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Airplane. North American X-15 broke many speed and altitude records in the 1960s and pioneered engineering concepts for later aircraft and spacecraft. Military transport aircraft may employ rocket-assisted take offs for short-field situations. Otherwise, rocket aircraft include spaceplanes, like SpaceShipTwo, for travel beyond the Earth's atmosphere and sport aircraft developed for the short-lived Rocket Racing League.## Design and manufacture.Most airplanes are constructed by companies with the objective of producing them in quantity for customers. The design and planning process, including safety tests, can last up to four years for small turboprops or longer for larger planes.During this process, the objectives and.

Airplane. resulted in reduced noise from aircraft engines in response to increased noise pollution from growth in air traffic over urban areas near airports.Small planes can be designed and constructed by amateurs as homebuilts. Other homebuilt aircraft can be assembled using pre-manufactured kits of parts that can be assembled into a basic plane and must then be completed by the builder.Few companies produce planes on a large scale. However, the production of a plane for one company is a process that actually involves dozens, or even hundreds, of other companies and plants, that produce the parts that go into the plane.

Airplane. to stabilize the plane's pitch (tilt up or down) and mounts the elevators, which provide pitch control. "Landing gear", a set of wheels, skids, or floats that support the plane while it is on the surface. On seaplanes, the bottom of the fuselage or floats (pontoons) support it while on the water. On some planes the landing gear retracts during flight to reduce drag.## Wings.The wings of a fixed-wing aircraft are static planes extending either side of the aircraft. When the aircraft travels forwards, air flows over the wings, which are shaped to create lift. This shape is called an.

Airplane. instruments.Airplanes have complex flight control systems. The main controls allow the pilot to direct the aircraft in the air by controlling the attitude (roll, pitch and yaw) and engine thrust.On manned aircraft, cockpit instruments provide information to the pilots, including flight data, engine output, navigation, communications and other aircraft systems that may be installed.## Safety.When risk is measured by deaths per passenger kilometer, air travel is approximately 10 times safer than travel by bus or rail. However, when using the deaths per journey statistic, air travel is significantly more dangerous than car, rail, or bus travel. Air travel insurance is.

Airplane. relatively expensive for this reason—insurers generally use the deaths per journey statistic. There is a significant difference between the safety of airliners and that of smaller private planes, with the per-mile statistic indicating that airliners are 8.3 times safer than smaller planes.## Environmental impact.Like all activities involving combustion, fossil-fuel-powered aircraft release soot and other pollutants into the atmosphere. Greenhouse gases such as carbon dioxide (CO2) are also produced. In addition, there are environmental impacts specific to airplanes: for instance, Airplanes operating at high altitudes near the tropopause (mainly large jet airliners) emit aerosols and leave contrails, both of which can.

Airplane. increase cirrus cloud formationcloud cover may have increased by up to 0.2% since the birth of aviation. Airplanes operating at high altitudes near the tropopause can also release chemicals that interact with greenhouse gases at those altitudes, particularly nitrogen compounds, which interact with ozone, increasing ozone concentrations. Most light piston aircraft burn avgas, which contains tetraethyllead (TEL). Some lower-compression piston engines can operate on unleaded mogas and turbine engines and diesel enginesneither of which require leadare used on some newer light aircraft. Some non-polluting light electric aircraft are already in production.Another environmental impact of airplanes is noise pollution, mainly caused.


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Torres de Serranos. The Serrans Gate or Serranos Gate (Valencian: Porta dels Serrans, Valencian: [ˈpɔɾta ðels seˈrans]; Spanish: Puerta de Serranos, [ˈpweɾta ðe seˈranos]), also known as Serrans Towers or Serranos Towers (Valencian: Torres dels Serrans, Valencian: [ˈtorez ðels seˈrans]; Spanish: Torres de Serranos, [ˈtorez ðe seˈranos]) is one of the twelve gates that formed part of the ancient city wall, the Christian Wall (Muralla cristiana), of the city of Valencia, Spain. It was built in Valencian Gothic style at the end of the 14th century (between 1392 and 1398). Its name is probably due to its location in the northeast of the.


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Acer macrophyllum. gold and yellow, contrasting against backdrops of evergreen conifers.In spring, the tree produces flowers in pendulous racemes 10 - 15 (cm) long, greenish-yellow with inconspicuous petals. It is hermaphroditic, bearing both male and female flowers in each raceme. The flowers appear in early spring, before the leaves.The fruit is a paired winged samara, each seed 1 - 1.5 (cm) in diameter with a 4 - 5 (cm) wing. Bigleaf maple begins bearing seed at about ten years of age.In May 2018 the oldest two Oregon maples in Europe, 175 years old, were removed from Trinity College Dublin (TCD), Ireland. The.


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Wolf. competition for food within the pack. Wolves are also territorial and fights over territory are among the principal causes of wolf mortality. The wolf is mainly a carnivore and feeds on large wild hooved mammals as well as smaller animals, livestock, carrion, and garbage. Single wolves or mated pairs typically have higher success rates in hunting than do large packs. Pathogens and parasites, notably rabies virus, may infect wolves.The global wild wolf population was estimated to be 300,000 in 2003 and is considered to be of Least Concern by the International Union for Conservation of Nature (IUCN). Wolves have a.

Wolf. long history of interactions with humans, having been despised and hunted in most pastoral communities because of their attacks on livestock, while conversely being respected in some agrarian and hunter-gatherer societies. The wolf is also considered the ancestor of the domestic dog. Although the fear of wolves exists in many human societies, the majority of recorded attacks on people have been attributed to animals suffering from rabies. Wolf attacks on humans are rare because wolves are relatively few, live away from people, and have developed a fear of humans because of their experiences with hunters, ranchers, and shepherds.## Etymology.The English.

Wolf. and Wolfdregil ("Wolf Runner").## Taxonomy.In 1758, the Swedish botanist and zoologist Carl Linnaeus published in his "Systema Naturae" the binomial nomenclature. "Canis" is the Latin word meaning "dog", and under this genus he listed the doglike carnivores including domestic dogs, wolves, and jackals. He classified the domestic dog as "Canis familiaris", and the wolf as "Canis lupus". Linnaeus considered the dog to be a separate species from the wolf because of its "cauda recurvata" (upturning tail) which is not found in any other canid.## Subspecies.In the third edition of "Mammal Species of the World" published in 2005, the mammalogist W.

Wolf. Christopher Wozencraft listed under "C. lupus" 36 wild subspecies, and proposed two additional subspecies: "familiaris" (Linnaeus, 1758) and "dingo" (Meyer, 1793). Wozencraft included "hallstromi"—the New Guinea singing dog—as a taxonomic synonym for the dingo. Wozencraft referred to a 1999 mitochondrial DNA (mtDNA) study as one of the guides in forming his decision, and listed the 38 subspecies of "C. lupus" under the biological common name of "wolf", the nominate subspecies being the Eurasian wolf ("C. l. lupus") based on the type specimen that Linnaeus studied in Sweden. Studies using paleogenomic techniques reveal that the modern wolf and the dog are.

Wolf. sister taxa, as modern wolves are not closely related to the population of wolves that was first domesticated. In 2019, a workshop hosted by the IUCN/Species Survival Commission's Canid Specialist Group considered the New Guinea singing dog and the dingo to be feral "Canis familiaris", and therefore should not be assessed for the IUCN Red List.## Evolution.The phylogenetic descent of the extant wolf "C. lupus" from "C. etruscus" through "C. mosbachensis" is widely accepted. The earliest fossils of "C. lupus" were found in what was once eastern Beringia at Old Crow, Yukon, Canada, and at Cripple Creek Sump, Fairbanks, Alaska.

Wolf. often processed carcasses, or that they competed with other carnivores and needed to consume their prey quickly. Compared with those found in the modern spotted hyena, the frequency and location of tooth fractures in these wolves indicates they were habitual bone crackers.Genomic studies suggest modern wolves and dogs descend from a common ancestral wolf population that existed 20,000 years ago. A 2021 study found that the Himalayan wolf and the Indian plains wolf are part of a lineage that is basal to other wolves and split from them 200,000 years ago. Other wolves appear to have originated in Beringia in.

Wolf. an expansion that was driven by the huge ecological changes during the close of the Late Pleistocene. A study in 2016 indicates that a population bottleneck was followed by a rapid radiation from an ancestral population at a time during, or just after, the Last Glacial Maximum. This implies the original morphologically diverse wolf populations were out-competed and replaced by more modern wolves.A 2016 genomic study suggests that Old World and New World wolves split around 12,500 years ago followed by the divergence of the lineage that led to dogs from other Old World wolves around 11,100–12,300 years ago. An.

Wolf. extinct Late Pleistocene wolf may have been the ancestor of the dog, with the dog's similarity to the extant wolf being the result of genetic admixture between the two. The dingo, Basenji, Tibetan Mastiff and Chinese indigenous breeds are basal members of the domestic dog clade. The divergence time for wolves in Europe, the Middle East, and Asia is estimated to be fairly recent at around 1,600 years ago. Among New World wolves, the Mexican wolf diverged around 5,400 years ago.## Admixture with other canids.In the distant past, there has been gene flow between African wolves, golden jackals, and gray.

Wolf. wolves. The African wolf is a descendant of a genetically admixed canid of 72% wolf and 28% Ethiopian wolf ancestry. One African wolf from the Egyptian Sinai Peninsula shows admixture with Middle Eastern gray wolves and dogs. There is evidence of gene flow between golden jackals and Middle Eastern wolves, less so with European and Asian wolves, and least with North American wolves. This indicates the golden jackal ancestry found in North American wolves may have occurred before the divergence of the Eurasian and North American wolves.The common ancestor of the coyote and the wolf has admixed with a ghost.

Wolf. population of an extinct unidentified canid. This canid is genetically close to the dhole and evolved after the divergence of the African hunting dog from the other canid species. The basal position of the coyote compared to the wolf is proposed to be due to the coyote retaining more of the mitochondrial genome of this unidentified canid. Similarly, a museum specimen of a wolf from southern China collected in 1963 showed a genome that was 12–14% admixed from this unknown canid. In North America, most coyotes and wolves show varying degrees of past genetic admixture. The red wolf of the.

Wolf. southeastern United States is a hybrid animal with 40%:60% wolf to coyote ancestry. In addition, there was found to be 60%:40% wolf to coyote genetics in Eastern timber wolves, and 75%:25% in the Great Lakes region wolves.In more recent times, some male Italian wolves originated from dog ancestry, which indicates female wolves will breed with male dogs in the wild. In the Caucasus Mountains, ten percent of dogs including livestock guardian dogs, are first generation hybrids. Although mating between golden jackals and wolves has never been observed, evidence of jackal-wolf hybridization was discovered through mitochondrial DNA analysis of jackals living.

Wolf. in the Caucasus Mountains and in Bulgaria.In 2021, a genetic study found that the dog's similarity to the extant gray wolf was the result of substantial dog-into-wolf gene flow, with almost negligible wolf-into-dog gene flow since the dog's domestication. Some gray wolves were related to all ancient and modern dogs.## Description.The wolf is the largest extant member of the Canidae family, and is further distinguished from coyotes and jackals by a broader snout, shorter ears, a shorter torso and a longer tail. It is slender and powerfully built, with a large, deeply descending rib cage, a sloping back, and a.

Wolf. the calcaneal tendons. The winter fur is highly resistant to the cold. Wolves in northern climates can rest comfortably in open areas at -40 (C) by placing their muzzles between the rear legs and covering their faces with their tail. Wolf fur provides better insulation than dog fur and does not collect ice when warm breath is condensed against it.In cold climates, the wolf can reduce the flow of blood near its skin to conserve body heat. The warmth of the foot pads is regulated independently from the rest of the body and is maintained at just above tissue-freezing point.

Wolf. southern Canada, the U.S., and Mexico being predominantly gray. In some areas of the Rocky Mountains of Alberta and British Columbia, the coat colour is predominantly black, some being blue-gray and some with silver and black. Differences in coat colour between sexes is absent in Eurasia; females tend to have redder tones in North America. Black-coloured wolves in North America acquired their colour from wolf-dog admixture after the first arrival of dogs across the Bering Strait 12,000 to 14,000 years ago. Research into the inheritance of white colour from dogs into wolves has yet to be undertaken.## Ecology.## Distribution and.

Wolf. habitat.Wolves occur across Eurasia and North America. However, deliberate human persecution because of livestock predation and fear of attacks on humans has reduced the wolf's range to about one-third of its historic range; the wolf is now extirpated (locally extinct) from much of its range in Western Europe, the United States and Mexico, and completely in Ireland, the United Kingdom, and Japan. In modern times, the wolf occurs mostly in wilderness and remote areas. The wolf can be found between sea level and 3000 (m). Wolves live in forests, inland wetlands, shrublands, grasslands (including Arctic tundra), pastures, deserts, and rocky.

Wolf. in diet between wolves living on different continents is based on the variety of hoofed mammals and of available smaller and domesticated prey.In North America, the wolf's diet is dominated by wild large hoofed mammals (ungulates) and medium-sized mammals. In Asia and Europe, their diet is dominated by wild medium-sized hoofed mammals and domestic species. The wolf depends on wild species, and if these are not readily available, as in Asia, the wolf is more reliant on domestic species. Across Eurasia, wolves prey mostly on moose, red deer, roe deer and wild boar. In North America, important range-wide prey are.

Wolf. elk, moose, caribou, white-tailed deer and mule deer. They also prey on bison and muskoxen. Wolves can digest their meal in a few hours and can feed several times in one day, making quick use of large quantities of meat. A well-fed wolf stores fat under the skin, around the heart, intestines, kidneys, and bone marrow, particularly during the autumn and winter.Nonetheless, wolves are not fussy eaters. Smaller-sized animals that may supplement their diet include rodents, rabbits, hares, insectivores and smaller carnivores. They frequently eat waterfowl and their eggs. When such foods are insufficient, they prey on lizards, snakes, frogs,.

Wolf. and large insects when available. Wolves in northern Minnesota prey on northern pike in freshwater streams. The diet of coastal wolves in Alaska includes 20% salmon, while those of coastal wolves in British Columbia includes 25% marine sources, and those on the nearby islands 75%.In Europe, wolves eat apples, pears, figs, melons, berries and cherries. In North America, wolves eat blueberries and raspberries. Wolves also eat grass, which may provide some vitamins, but is most likely used mainly to induce vomiting to rid themselves of intestinal parasites or long guard hairs. They are known to eat the berries of mountain-ash,.

Wolf. lily of the valley, bilberries, cowberries, European black nightshade, grain crops, and the shoots of reeds.In times of scarcity, wolves will readily eat carrion. In Eurasian areas with dense human activity, many wolf populations are forced to subsist largely on livestock and garbage. Prey in North America continue to occupy suitable habitats with low human density, the wolves eating livestock and garbage only in dire circumstances. Cannibalism is not uncommon in wolves during harsh winters, when packs often attack weak or injured wolves and may eat the bodies of dead pack members.## Interactions with other predators.Wolves typically dominate other canid.

Wolf. adjacent mountain ranges. Wolves and cougars typically avoid encountering each other by hunting at different elevations for different prey (niche partitioning). In winter, when snow accumulation forces their prey into valleys, interactions between the two species become more likely. Wolves in packs usually dominate cougars and can steal their kills or even kill them, while one-to-one encounters tend to be dominated by the cat. There are several documented cases of cougars killing wolves. Wolves more broadly affect cougar population dynamics and distribution by dominating territory and prey opportunities and disrupting the feline's behaviour. Wolf and Siberian tiger interactions are well-documented.

Wolf. off single or outnumbered hyenas. There is at least one case in Israel of a hyena associating and cooperating with a wolf pack. It is proposed that the hyena could benefit from the wolves' superior ability to hunt large, agile prey. The wolves could benefit from the hyena's superior sense of smell, to locate and dig out tortoises, to crack open large bones, and to tear open discarded food containers like tin cans.## Behaviour.## Social structure.The wolf is a social animal. Its populations consist of packs and lone wolves, most lone wolves being temporarily alone while they disperse from packs.

Wolf. to form their own or join another one. The wolf's basic social unit is the nuclear family consisting of a mated pair accompanied by their offspring. The average pack size in North America is eight wolves and in Europe 5.5 wolves. The average pack across Eurasia consists of a family of eight wolves (two adults, juveniles, and yearlings), or sometimes two or three such families, with examples of exceptionally large packs consisting of up to 42 wolves being known. Cortisol levels in wolves rise significantly when a pack member dies, indicating the presence of stress. During times of prey abundance.

Wolf. and the Denali National Park and Preserve were due to other wolves.## Communication.Wolves communicate using vocalizations, body postures, scent, touch, and taste. The phases of the moon have no effect on wolf vocalization, and despite popular belief, wolves do not howl at the moon. Wolves howl to assemble the pack usually before and after hunts, to pass on an alarm particularly at a den site, to locate each other during a storm, while crossing unfamiliar territory, and to communicate across great distances. Wolf howls can under certain conditions be heard over areas of up to 130 (km2). Other vocalizations include.

Wolf. growls, barks and whines. Wolves do not bark as loudly or continuously as dogs do in confrontations, rather barking a few times and then retreating from a perceived danger. Aggressive or self-assertive wolves are characterized by their slow and deliberate movements, high body posture and raised hackles, while submissive ones carry their bodies low, flatten their fur, and lower their ears and tail.Scent marking involves urine, feces, and anal gland scents. This is more effective at advertising territory than howling and is often used in combination with scratch marks. Wolves increase their rate of scent marking when they encounter the.

Wolf. marks of wolves from other packs. Lone wolves will rarely mark, but newly bonded pairs will scent mark the most. These marks are generally left every 240 (m) throughout the territory on regular travelways and junctions. Such markers can last for two to three weeks, and are typically placed near rocks, boulders, trees, or the skeletons of large animals. Raised leg urination is considered to be one of the most important forms of scent communication in the wolf, making up 60–80% of all scent marks observed.## Reproduction.Wolves are monogamous, mated pairs usually remaining together for life. Should one of the.

Wolf. better warmed by sunlight exposure, and the snow can thaw more quickly. Resting places, play areas for the pups, and food remains are commonly found around wolf dens. The odor of urine and rotting food emanating from the denning area often attracts scavenging birds like magpies and ravens. Though they mostly avoid areas within human sight, wolves have been known to nest near domiciles, paved roads and railways. During pregnancy, female wolves remain in a den located away from the peripheral zone of their territories, where violent encounters with other packs are less likely to occur.The gestation period lasts 62–75.

Wolf. blood loss, thereby lessening the risk of injury to themselves. With medium-sized prey, such as roe deer or sheep, wolves kill by biting the throat, severing nerve tracks and the carotid artery, thus causing the animal to die within a few seconds to a minute. With small, mouselike prey, wolves leap in a high arc and immobilize it with their forepaws.Once prey is brought down, wolves begin to feed excitedly, ripping and tugging at the carcass in all directions, and bolting down large chunks of it. The breeding pair typically monopolizes food to continue producing pups. When food is scarce,.

Wolf. wolf attacks on people being attributed to rabid animals. Although canine distemper is lethal in dogs, it has not been recorded to kill wolves, except in Canada and Alaska. The canine parvovirus, which causes death by dehydration, electrolyte imbalance, and endotoxic shock or sepsis, is largely survivable in wolves, but can be lethal to pups. Wolves may catch infectious canine hepatitis from dogs, though there are no records of wolves dying from it. Papillomatosis has been recorded only once in wolves, and likely does not cause serious illness or death, though it may alter feeding behaviours. The canine coronavirus has.

Wolf. been recorded in Alaskan wolves, infections being most prevalent in winter months.Bacterial diseases carried by wolves include: brucellosis, Lyme disease, leptospirosis, tularemia, bovine tuberculosis, listeriosis and anthrax. Wolves can catch "Brucella suis" from wild and domestic reindeer. While adult wolves tend not to show any clinical signs, it can severely weaken the pups of infected females. Although lyme disease can debilitate individual wolves, it does not appear to significantly affect wolf populations. Leptospirosis can be contracted through contact with infected prey or urine, and can cause fever, anorexia, vomiting, anemia, hematuria, icterus, and death. Wolves living near farms are more.

Wolf. vulnerable to the disease than those living in the wilderness, probably because of prolonged contact with infected domestic animal waste. Wolves may catch tularemia from lagomorph prey, though its effect on wolves is unknown. Although bovine tuberculosis is not considered a major threat to wolves, it has been recorded to have killed two wolf pups in Canada.## Parasitic.Wolves carry ectoparasites and endoparasites; those in the former Soviet Union have been recorded to carry at least 50 species. Most of these parasites infect wolves without adverse effects, though the effects may become more serious in sick or malnourished specimens. Parasitic infection.

Wolf. in wolves is of particular concern to people. Wolves can spread them to dogs, which in turn can carry the parasites to humans. In areas where wolves inhabit pastoral areas, the parasites can be spread to livestock.Wolves are often infested with a variety of arthropod exoparasites, including fleas, ticks, lice, and mites. The most harmful to wolves, particularly pups, is the mange mite ("Sarcoptes scabiei"), though they rarely develop full-blown mange, unlike foxes. Lice, such as "Trichodectes canis", may cause sickness in wolves, but rarely death. Ticks of the genus "Ixodes" can infect wolves with Lyme disease and Rocky Mountain.

Wolf. spotted fever. The tick "Dermacentor pictus" also infests wolves. Other ectoparasites include chewing lice, sucking lice and the fleas "Pulex irritans" and "Ctenocephalides canis".Endoparasites known to infect wolves include: protozoans and helminths (flukes, tapeworms, roundworms and thorny-headed worms). Of 30,000 protozoan species, only a few have been recorded to infect wolves: "Isospora", "Toxoplasma", "Sarcocystis", "Babesia", and "Giardia". Some wolves carry "Neospora caninum", which can be spread to cattle and is correlated with bovine miscarriages. Among flukes, the most common in North American wolves is "Alaria", which infects small rodents and amphibians which are eaten by wolves. Upon reaching maturity, "Alaria.

Wolf. migrates to the wolf's intestine, but does little harm. "Metorchis conjunctus", which enters wolves through eating fish, infects the wolf's liver or gall bladder, causing liver disease, inflammation of the pancreas, and emaciation. Most other fluke species reside in the wolf's intestine, though "Paragonimus westermani" lives in the lungs. Tapeworms are commonly found in wolves, as their primary hosts are ungulates, small mammals, and fish, which wolves feed upon. Tapeworms generally cause little harm in wolves, though this depends on the number and size of the parasites, and the sensitivity of the host. Symptoms often include constipation, toxic and allergic.

Wolf. intestinal irritation, bloating, vomiting, and diarrhea. Wolves may catch "Dioctophyma renale" from minks, which infects the kidneys, and can grow to lengths of 100 (cm). "D. renale" causes the complete destruction of the kidney's functional tissue and can be fatal if both kidneys are infected. Wolves can tolerate low levels of "Dirofilaria immitis" for many years without showing any ill effects, though high levels can kill wolves through cardiac enlargement and congestive hepatopathy. Wolves probably become infected with "Trichinella spiralis" by eating infected ungulates. Although "T. spiralis" is not known to produce clinical signs in wolves, it can cause emaciation,.

Wolf. salivation, and crippling muscle pains in dogs. Thorny-headed worms rarely infect wolves, though three species have been identified in Russian wolves: "Nicolla skrjabini", "Macracanthorhynchus catulinus", and "Moniliformis moniliformis".## Status and conservation.The global wild wolf population in 2003 was estimated at 300,000. Wolf population declines have been arrested since the 1970s. This has fostered recolonization and reintroduction in parts of its former range as a result of legal protection, changes in land use, and rural human population shifts to cities. Competition with humans for livestock and game species, concerns over the danger posed by wolves to people, and habitat fragmentation pose.

Wolf. world. His most famous is the fable of "The Boy Who Cried Wolf", which is directed at those who knowingly raise false alarms, and from which the idiomatic phrase "to cry wolf" is derived. Some of his other fables concentrate on maintaining the trust between shepherds and guard dogs in their vigilance against wolves, as well as anxieties over the close relationship between wolves and dogs. Although Aesop used wolves to warn, criticize and moralize about human behaviour, his portrayals added to the wolf's image as a deceitful and dangerous animal. The Bible uses an image of a wolf lying.

Wolf. Ivan Bunin, Leonid Pavlovich Sabaneyev, and others. Tolstoy's "War and Peace" and Chekhov's "Peasants" both feature scenes in which wolves are hunted with hounds and Borzois. The musical "Peter and the Wolf" involves a wolf being captured for eating a duck, but is spared and sent to a zoo.Wolves are among the central characters of Rudyard Kipling's "The Jungle Book". His portrayal of wolves has been praised posthumously by wolf biologists for his depiction of them: rather than being villainous or gluttonous, as was common in wolf portrayals at the time of the book's publication, they are shown as living.

Wolf. in amiable family groups and drawing on the experience of infirm but experienced elder pack members. Farley Mowat's largely fictional 1963 memoir "Never Cry Wolf" is widely considered to be the most popular book on wolves, having been adapted into a Hollywood film and taught in several schools decades after its publication. Although credited with having changed popular perceptions on wolves by portraying them as loving, cooperative and noble, it has been criticized for its idealization of wolves and its factual inaccuracies.## Conflicts.Human presence appears to stress wolves, as seen by increased cortisol levels in instances such as snowmobiling near.

Wolf. their territory.## Predation on livestock.Livestock depredation has been one of the primary reasons for hunting wolves and can pose a severe problem for wolf conservation. As well as causing economic losses, the threat of wolf predation causes great stress on livestock producers, and no foolproof solution of preventing such attacks short of exterminating wolves has been found. Some nations help offset economic losses to wolves through compensation programs or state insurance. Domesticated animals are easy prey for wolves, as they have been bred under constant human protection, and are thus unable to defend themselves very well. Wolves typically resort to.

Wolf. attacking livestock when wild prey is depleted. In Eurasia, a large part of the diet of some wolf populations consists of livestock, while such incidents are rare in North America, where healthy populations of wild prey have been largely restored.The majority of losses occur during the summer grazing period, untended livestock in remote pastures being the most vulnerable to wolf predation. The most frequently targeted livestock species are sheep (Europe), domestic reindeer (northern Scandinavia), goats (India), horses (Mongolia), cattle and turkeys (North America). The number of animals killed in single attacks varies according to species: most attacks on cattle and.

Wolf. horses result in one death, while turkeys, sheep and domestic reindeer may be killed in surplus. Wolves mainly attack livestock when the animals are grazing, though they occasionally break into fenced enclosures.## Competition with dogs.A review of the studies on the competitive effects of dogs on sympatric carnivores did not mention any research on competition between dogs and wolves. Competition would favour the wolf, which is known to kill dogs; however, wolves usually live in pairs or in small packs in areas with high human persecution, giving them a disadvantage when facing large groups of dogs.Wolves kill dogs on occasion,.

Wolf. and some wolf populations rely on dogs as an important food source. In Croatia, wolves kill more dogs than sheep, and wolves in Russia appear to limit stray dog populations. Wolves may display unusually bold behaviour when attacking dogs accompanied by people, sometimes ignoring nearby humans. Wolf attacks on dogs may occur both in house yards and in forests. Wolf attacks on hunting dogs are considered a major problem in Scandinavia and Wisconsin. The most frequently killed hunting breeds in Scandinavia are Harriers, older animals being most at risk, likely because they are less timid than younger animals, and react.

Wolf. to the presence of wolves differently. Large hunting dogs such as Swedish Elkhounds are more likely to survive wolf attacks because of their better ability to defend themselves.Although the number of dogs killed each year by wolves is relatively low, it induces a fear of wolves entering villages and farmyards to prey on them. In many cultures, dogs are seen as family members, or at least working team members, and losing one can lead to strong emotional responses such as demanding more liberal hunting regulations.Dogs that are employed to guard sheep help to mitigate human–wolf conflicts, and are often proposed.

Wolf. as one of the non-lethal tools in the conservation of wolves. Shepherd dogs are not particularly aggressive, but they can disrupt potential wolf predation by displaying what is to the wolf ambiguous behaviours, such as barking, social greeting, invitation to play or aggression. The historical use of shepherd dogs across Eurasia has been effective against wolf predation, especially when confining sheep in the presence of several livestock guardian dogs. Shepherd dogs are sometimes killed by wolves.## Attacks on humans.The fear of wolves has been pervasive in many societies, though humans are not part of the wolf's natural prey. How wolves.

Wolf. almost always women. Indian wolves have a history of preying on children, a phenomenon called "child-lifting". They may be taken primarily in the spring and summer periods during the evening hours, and often within human settlements.Cases of rabid wolves are low when compared to other species, as wolves do not serve as primary reservoirs of the disease, but can be infected by animals such as dogs, jackals and foxes. Incidents of rabies in wolves are very rare in North America, though numerous in the eastern Mediterranean, the Middle East and Central Asia. Wolves apparently develop the "furious" phase of rabies.

Wolf. to a very high degree. This, coupled with their size and strength, makes rabid wolves perhaps the most dangerous of rabid animals. Bites from rabid wolves are 15 times more dangerous than those of rabid dogs. Rabid wolves usually act alone, travelling large distances and often biting large numbers of people and domestic animals. Most rabid wolf attacks occur in the spring and autumn periods. Unlike with predatory attacks, the victims of rabid wolves are not eaten, and the attacks generally occur only on a single day. The victims are chosen at random, though most cases involve adult men. During.

Wolf. the fifty years up to 2002, there were eight fatal attacks in Europe and Russia, and more than two hundred in southern Asia.## Human hunting of wolves.Theodore Roosevelt said wolves are difficult to hunt because of their elusiveness, sharp senses, high endurance, and ability to quickly incapacitate and kill a dog. Historic methods included killing of spring-born litters in their dens, coursing with dogs (usually combinations of sighthounds, Bloodhounds and Fox Terriers), poisoning with strychnine, and trapping.A popular method of wolf hunting in Russia involves trapping a pack within a small area by encircling it with fladry poles carrying a.

Wolf. human scent. This method relies heavily on the wolf's fear of human scents, though it can lose its effectiveness when wolves become accustomed to the odor. Some hunters can lure wolves by imitating their calls. In Kazakhstan and Mongolia, wolves are traditionally hunted with eagles and falcons, though this practice is declining, as experienced falconers are becoming few in number. Shooting wolves from aircraft is highly effective, due to increased visibility and direct lines of fire. Several types of dog, including the Borzoi and Kyrgyz Tajgan, have been specifically bred for wolf hunting.## As pets and working animals.Wolves and wolf-dog.

Wolf. hybrids are sometimes kept as exotic pets. Although closely related to domestic dogs, wolves do not show the same tractability as dogs in living alongside humans, being generally less responsive to human commands and more likely to act aggressively. A person is more likely to be fatally mauled by a pet wolf or wolf-dog hybrid than by a dog.The International Wolf CenterStaying Safe in Wolf Country, ADFG (January 2009)Watch "Death of a Legend" and "Cry of the Wild" by Bill Mason.


Question: What is the height of this building in metre?

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Sighișoara Clock Tower. In every fortification system there is one fortress that dominates the others: the master-tower. The Clock Tower of Sighișoara (, ) is the main entry point to the citadel, opposite guarded by Tailors' Tower. With its 64 meters of height, the tower is visible from almost every corner of the city from Mures County, its purpose was to defend the main gate of the citadel and also served as the town hall until 1556. Now It's considered one of the most expressive clock tower in the whole Transylvania.## History.The tower began to be built at the end of 13th century,.

Sighișoara Clock Tower. to dig for 35m to find a source of water.## Restorations.The tower was modernized in 1964 with an electric motor, without diminishing the archaic effect of the clock. Since 1898, the tower serves as a museum in Sighisoara, a title that gives to it the status of symbol of the Citadel, and the main attraction for visitors.## Architectural Features.The tower is a rectangular prism with massive walls, with a rectangular base measuring 12x8,66 meters, with four floors and an observation gallery. The current distance between the base of the tower and the base of the roof is 30 meters, and.


Question: What was the building or structure prior to the existence of this building?

Instruction: Determine the Wikipedia snippet that matches the question of this image.

30 St Mary Axe. 30 St Mary Axe (previously known as the Swiss Re Building and informally known as the Gherkin) is a commercial skyscraper in London's primary financial district, the City of London. It was completed in December 2003 and opened in April 2004. With 41 floors, it is 180 (m) tall and stands on the sites of the former Baltic Exchange and Chamber of Shipping, which were extensively damaged in 1992 in the Baltic Exchange bombing by a device placed by the Provisional IRA in St Mary Axe, a narrow street leading north from Leadenhall Street.After plans to build the 92-storey Millennium.

30 St Mary Axe. Tower were dropped, 30 St Mary Axe was designed by Norman Foster and the Arup Group. It was erected by Skanska; construction started in 2001.The building has become a recognisable landmark of London, and it is one of the city's most widely recognised examples of contemporary architecture. It won the 2003 Emporis Skyscraper Award.## Site and early plans.The building stands on the site of the former Baltic Exchange (24–28 St Mary Axe), which was the headquarters of a global marketplace for shipping freight contracts and soft commodities, and the Chamber of Shipping (30–32 St Mary Axe). The tower's topmost panoramic.

30 St Mary Axe. dome, known as the "lens", recalls the iconic glass dome that covered part of the ground floor of the Baltic Exchange and much of which is now displayed at the National Maritime Museum. "The Gherkin" nickname was applied to the current building at least as early as 1999, referring to the plan's highly unorthodox layout and appearance.On 10 April 1992, the Provisional IRA detonated a bomb close to the Baltic Exchange, causing extensive damage to the historic building and neighbouring structures. The United Kingdom government's statutory adviser on the historic environment, English Heritage, and the City of London's governing body,.

30 St Mary Axe. the City of London Corporation, were keen that any redevelopment must restore the Baltic Exchange's old façade onto St Mary Axe. The Exchange Hall was a celebrated fixture of the shipping market.English Heritage then discovered that the damage was far more severe than initially thought, and they stopped insisting on full restoration, albeit over the objections of architectural conservationists. The Baltic Exchange and the Chamber of Shipping sold the land to Trafalgar House in 1995. Most of the remaining structures on the Baltic Exchange site were then carefully dismantled, and the interior of Exchange Hall and the façade were preserved,.

30 St Mary Axe. accepted.## Planning process.John Prescott, the Deputy Prime Minister, granted planning permission on 23 August 2000 to construct a building on the site, which would be much larger than the old Exchange. The site needed development, was not on any of the "sight lines" (planning guidance requires that new buildings do not obstruct or detract from the view of St Paul's Cathedral dome when viewed from a number of locations around London), and had housed the Baltic Exchange.The plan for the site was to reconstruct the Baltic Exchange. GMW Architects proposed a new rectangular building surrounding a restored exchange: it would.