 The most basal group of <unk> is the superfamily <unk> . <unk> have several primitive or <unk> features , including a single <unk> <unk> and a bone called the <unk> that is absent in other <unk> . <unk> include the Late <unk> genus <unk> and the family <unk> . <unk> has also been included in <unk> , and is the oldest known <unk> family . <unk> <unk> is the oldest species , having been present over 330 million years ago during the <unk> stage of the Early <unk> . Recent analyses place <unk> outside <unk> in a more derived position . Other primitive <unk> include <unk> and <unk> . <unk> and <unk> , both described in 2005 from <unk> , are also primitive yet come from the Late <unk> . They are almost 40 million years younger than other basal <unk> , implying a long ghost lineage of species that are not yet known in the fossil record . 
