 The three recent treatments ( Henderson and <unk> , <unk> , and <unk> and <unk> ) recognised a total of 73 species , but only 20 species are accepted by all of them . The remainder account for either nine species or more than 40 . For example , what Andrew Henderson considered a single species , <unk> <unk> , other authors have considered a species complex consisting of four or five species . <unk> doubted the validity of A. <unk> as a species , and described four new species from material that had previously been attributed to A. <unk> — A. <unk> , A. <unk> , A. <unk> and A. <unk> . <unk> and <unk> accepted both <unk> 's four species and A. <unk> . However , Jean @-@ Christophe <unk> was of the opinion that A. <unk> , A. <unk> and A. <unk> were all very similar , and thought it likely that they all represented the same species . 
