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DescriptionDioecious trees or shrubs. Trunk tall or very short, bearing spirally arranged leaf-bases and scale leaves. Leaves spirally arranged, pinnate; leaflets with longitudinal parallel veins and no midrib. Male cones consisting of leathery scales with numerous pollen-sacs on their abaxial surface. Female cones composed of thick and often woody, peltate scales bearing 2 (or more) ovules on their adaxial surface. Seeds angled, with a fleshy outer layer enclosing a hard inner shell.