Peramelemorphia (bandicoots and bilbies) consists of 22 species that are divided among 8 genera and 3 families: Peramelidae (bandicoots and echymiperas), Chaeropodidae (pig-footed bandicoot), and Thylacomyidae (bilbies). They have a rodent-like appearance with short legs, a stocky body, a short neck, and a long, pointy nose. They are largely nocturnal, and possess a well-developed sense of smell and eyes that are well adapted for night vision. Most peramelemorphs have brownish-red or tan fur and are sometimes marked with stripes. Long, rabbit-like ears also characterize some species. They range in size from less than 100 grams to over 5 kilograms, though most are about the size of a rabbit or smaller. Peramelemorphs are omnivores that eat mainly insects, but also consume a variety of vegetable material and some vertebrates as well. They occupy a wide range of habitats throughout Australia, New Guinea, Tasmania and the surrounding islands. ("Bandicoots", 2006; Cavendish, 2007; IUCN, 2008; Vaughan, et al., 2000)
Bandicoots and bilbies are mainly found throughout Australia, New Guinea, and the surrounding islands, with the eastern barred bandicoot and southern brown bandicoot also being found in Tasmania. It is believed that members of the subfamily Peroryctinae most likely originated and radiated in New Guinea. However, their origin is speculative due to a lack of fossil evidence. Only 2 out of the 11 species of peroryctine bandicoots, the rufous spiny bandicoot and the Seram bandicoot, are currently found outside of New Guinea having ranges that extend to the tip of Cape York and Seram Island, respectively. The northern brown bandicoot is the only perameline bandicoot that is found outside of Australia in Southern New Guinea. ("Bandicoots", 2006; "Tasmanian Parks and Wildlife Service", 2008; Vaughan, et al., 2000)
- Other Geographic Terms
- island endemic
Peramelemorphs occupy a wide range of habitats, with altitude and climatic differences heavily influencing the distribution of species. Members of the family Peramelidae inhabit a variety of ecosystems, ranging from deserts to subalpine grasslands to tropical lowland rainforests, while thylacomyids primarily live in arid areas. Eastern barred bandicoots and the now extinct pig-footed bandicoot prefer grassland habitats, golden bandicoots inhabit the Top End and Kimberly tropics of Australia, brown bandicoots live in more secluded forests and the only living species of bilby, the greater bilby, is a desert-dweller. By occupying a wide variety of habitats and vegetation types, bandicoots and bilbies largely avoid competition. In New Guinea, peramelemorphs (Peroryctinae) are distributed throughout a wide range of altitudes. However, several species may occur sympatrically at moderate altitudes. The northern brown bandicoot, giant bandicoot and most species of spiny bandicoots prefer lowland areas, though some may live as high as 2000 m. Mouse bandicoots, striped bandicoots and Raffray’s bandicoots are upland species and typically live at elevations above 1000 m. There is one known high altitude species, Seram bandicoots, that are only found at altitudes of around 1800 m. ("Bandicoots", 2006; Cavendish, 2007; Jackson, 2003; Vaughan, et al., 2000)
- Terrestrial Biomes
- desert or dune
- savanna or grassland
- scrub forest
Members of the order Peramelemorphia are terrestrial, ground-dwelling mammals. They range from 15 cm in length (excluding tail) and 100 g in weight to 60 cm in length and 5 kg in weight. Their bodies are compact in size with relatively short tails compared to the length of their bodies, except in the case of the greater bilby, which possesses a long, brush-like tail. Peramelemorphs have short necks, elongate skulls, and long, tapered snouts. Their ears are upright and can range from being small and rounded to fairly large and pointy. Males are usually larger than females and are socially dominant. ("Bandicoots and Bilbies: Peramelemorphia", 2009; Department of Primary Industries and Water, 2008; Gordon and Seeback, 2001; Myers, 2001; "Peramelemorphia", 2005)
The hind limbs of peramelemorphs are relatively long and exceptionally powerful. On the hind feet, the forth toe is the largest, while the bones of the second and third toes are fused, but still maintain separate claws (i.e., syndactyly). The front limbs are very short and well-adapted for ground foraging and digging. The first and fifth toes on the forefeet are either absent or lack claws if present. The second, third, and fourth toes have strong, flat claws for digging. They typically use their strong hind limbs to leap and hop through brushy habitats; however, when escaping danger they are able to run at a fast gallop. Their front and back legs work alternately. Characteristically, they land on hind and forefeet, and then take off with a push of their large hindfeet. ("Bandicoots and Bilbies: Peramelemorphia", 2009; Department of Primary Industries and Water, 2008; Myers, 2001; "Peramelemorphia", 2005)
Members of the order Peramelemorphia can be most noticeably recognized by their unique marsupium, the pouch located on the venter used to carry immature young. Unlike teh marsupium of kangaroos and wallabies, the marsupium of peramelemorphs opens to the rear. Although this condition is present in some diprotodonts (e.g., wombats), it is probably uniquely derived in each lineage. ("Bandicoots and Bilbies: Peramelemorphia", 2009; Department of Primary Industries and Water, 2008; Myers, 2001; "Peramelemorphia", 2005)
Peramelemorphs are omnivorous and their dentition is well-suited to a diet consisting of plants and insects. Unlike diprotodonts, which have only 2 lower incisors, peramelemorphs are polyprotodonts, having multiple lower incisors and anywhere from 4 to 5 upper incisors. Their incisors are flattened at the tips with the crown of the last lower incisor having two lobes. The canines are present and well-developed and they also have 3 premolars, which are narrow and pointed (plagialacoid) and 4 molars, which are tribosphenic or quadrate, in the upper and lower sets. This gives them the dental formula of 4-5/3, 1/1, 3/3, 4/4 = 46 or 48.
- Sexual Dimorphism
- male larger
Direct observations of mating in peramelemorphs are rare, however, based on behavioral data they are probably either polygynous or promiscuous, and females are polyestrous. Although peramelemorphs are solitary, male territories overlap with those of several females, and during mating season males spend a majority of their time searching for receptive females. Once they find an estrus female, they follow the potential mate until she is ready to be mounted. Females may mate with more than one male if the opportunity presents itself. ("Bandicoots and Bilbies: Peramelemorphia", 2009; "Bandicoots", 2006; Anderson and Knox Jones, 1984; Jackson, 2003; Seebeck, 1990; "Peramelemorphia", 2005)
Peramelemorphs are known for their accelerated breeding process, which enables a single female to give birth to as many as 16 young per year. Unlike all other marsupials, members of Peramelemorphia have a chorioallantoic placenta, which replaces the more typical yolk sac placenta a few days into gestation. Unlike the placenta found in 'true mammals', the placenta of peramelemorphs lacks villi, resulting in relatively shorter gestation when compared to 'true mammals', which developed the chorioallantoic placenta independently. Breeding can take place year-round for some genera, while others breed in the spring only. Day length, food availability, and weather conditions appear to have a significant impact on the timing of breeding in seasonal breeders. Year-round breeders occasionally show a decline in birthrate during times of food scarcity or drought. Gestation time is variable, from as little as 12.5 days in long-nosed bandicoots (among the shortest in any mammal) to about 14 days in several other species. Litters range in size from 2 to 5 offspring, but usually no more than 4 survive. Like other marsupials, young are altricial, weighing about 0.2 grams at birth. Immediately after birth, they crawl into their mother’s pouch and attach to a nipple. They leave the pouch after about 60 days, and are weaned in about 70 days. Females generally mate at about the time their previous litter leaves the pouch, so the weaning of one litter coincides with the birth of the next. ("Bandicoots and Bilbies: Peramelemorphia", 2009; "Bandicoots", 2006; Anderson and Knox Jones, 1984; Feldhammer, et al., 2007; Jackson, 2003; Seebeck, 1990; Wilson and Reeder, 2005)
While the ranges of male and female peramelemorphs extensively overlap, females likely dictate distribution as they select and defend high-quality habitats for nesting and foraging. Many species have scent glands just posterior to the ears. Present in both genders, it is thought that these glands are used to mark territorial boundaries or during male-male competition for mates. While some species, such as northern brown bandicoots create terrestrial nests with an internal chamber, others, such as eastern barred bandicoots make several different kinds of nests, including subterranean chambers that are used during parturition. (Gordon and Seeback, 2001; "Peramelemorphia", 2005)
- Key Reproductive Features
- seasonal breeding
- year-round breeding
- gonochoric/gonochoristic/dioecious (sexes separate)
- post-partum estrous
The accelerated reproductive cycle of Peramelemorphia results in minimal parental care to young. The unique placenta of peramelemorphs lacks villi, which reduces direct contact between mother and fetus. However, the umbilical cord remains attached for a few hours afterbirth to serve as a safety rope while young leave the uterus and crawl into the rear-opening marsupium. Juveniles may continue to live in the mother’s nest for some time after leaving the pouch, but it is not known if they remain in their mother's nest after weaning. There is no contact between mother and offspring after young leave the nest. Young peramelemorphs can reach reproductive maturity in as little as four months, however, only 11.5% of young survive to adulthood. ("Bandicoots and Bilbies: Peramelemorphia", 2009; "Bandicoots", 2006; Anderson and Knox Jones, 1984; Feldhammer, et al., 2007; Seebeck, 1990)
Bandicoots and bilbies live, on average, 1 to 2 years in the wild. While only 1 in 10 offspring usually survive, once they reach maturity life expectancy ranges from 2.5 to 3 years. In captivity, mean longevity for peramelemorphs is 2 to 4 years. ("Bandicoots and Bilbies: Peramelemorphia", 2009; "Bandicoots", 2006; Jackson, 2003; "Peramelemorphia", 2005)
All members of Peramelemorphia are solitary, coming together only to breed. Both males and females select territories, although male territories are larger and generally overlap with those of several different females. Most bandicoots are hostile toward one another, defending their territory with fighting, chasing, and scratching. Many species possess a scent gland just posterior to the ear, which is present in both genders in some species (e.g., northern brown bandicoot) and only present in males in others. These glands are used for marking territorial boundaries, and during male-male competition for mates or territory. Males are extremely territorial and during an encounter, they often mark the ground and surrounding plants with scents from the posterior ear gland. Males often warn each other with puffing sounds and may attempt to chase each other. Smaller males usually do not defend themselves against larger individuals when attacked. The only time peramelemorphs do not exhibit intraspecific aggression is when an estrus female encounters a male. All extant members of Peramelemorphia are nocturnal. ("Bandicoots and Bilbies: Peramelemorphia", 2009; "Bandicoots", 2006; Anderson and Knox Jones, 1984; Gordon and Seeback, 2001; Jackson, 2003; Seebeck, 1990; Vaughan, et al., 2000; Wilson and Reeder, 2005)
While bilbies are somewhat less aggressive than their close relatives, like the rest of Peramelemorphia, they are solitary and defend their territory when necessary. Bilbies are fossorial and are the only peramelemorphs to construct their own burrows; however, some species of bandicoot are known to burrow into the sand to escape hot weather. While most bandicoots live in burrows that are constructed from piles of vegetation covering small ground depressions, some species are known to occupy tree hollows or abandoned rabbit burrows. All extant peramelomorphs are nocturnal or crepuscular, although the recently-extinct pig-footed bandicoot was diurnal. While adapted for insect-eating, bandicoots and bilbies are often omnivorous, eating insects, grubs, plant material, and sometimes small vertebrates. Food is obtained by digging or rooting through plant litter on the ground. ("Bandicoots and Bilbies: Peramelemorphia", 2009; "Bandicoots", 2006; Anderson and Knox Jones, 1984; Gordon and Seeback, 2001; Jackson, 2003; Seebeck, 1990; Vaughan, et al., 2000; Wilson and Reeder, 2005)
Communication and Perception
Like other nocturnal mammals, peramelemorphs depend greatly on their senses of touch, smell, and hearing while hunting. Little is known about communication in northern brown bandicoot) and only present in the males of others. These glands are used for marking territorial boundaries and during male-male competition for mates or territory. Males warn potential rivals with by making puffing sounds and exhibit aggression with open-mouthed fighting and chasing. Captive peramelemorphs have been observed to make “soft spitting noises” when threatened. A few species have calls, which can ranged from shrill alarm calls to low, huffing noises accompanied by barred teeth. ("Bandicoots and Bilbies: Peramelemorphia", 2009; "Bandicoots", 2006; Anderson and Knox Jones, 1984; Jackson, 2003; Seebeck, 1990; "Peramelemorphia", 2005). Many species possess a scent glands just posterior to the ears, which are present in both genders of some species (e.g.,
- Other Communication Modes
Peramelemorphs are omnivorous and eat a wide variety of invertebrates including ants, termites, insect larvae, earthworms, spiders and centipedes as well as plant matter such as bulbs, grasses and seeds. Some species supplement their diet with fungi, bird eggs and small vertebrates such as lizards and mice. Peramelemorphs forage by digging with their strong front claws and then using their long snout and tongue grab ahold of food items. While they can eat many different foods, each colony tends to show preference for one or two particular food types. This is most likely due to regional availability of each food type and helps reduce intraspecific competition for resources. Many members of the family Peramelidae are not obligate drinkers, as they acquire much of their hydration needs through their diet. Their front limbs are short and well-adapted for ground foraging and digging, and their dentition is ideally suited to a diet of plants and insects. ("Bandicoots and Bilbies: Peramelemorphia", 2009; Jackson, 2003)
Peramelemorphs have few native predators. The only significant natural predators to bandicoots and bilbies are owls, quolls, and dingos. However, feral and domestic cats, dogs, foxes and other introduced animals have come to pose a considerable threat to the persistence of many local populations. In the past, bandicoots could often be found in Australian suburbs, however, domestic animals have significantly reduced their population. To protect themselves from predators, bandicoots and bilbies make nests in shallow holes in the ground, which they line with leaf litter. Leaf litter helps hide them from predators and protects them from inclement weather. A few species have calls, which can ranged from shrill alarm calls to low, huffing noises accompanied by barred teeth. ("Bandicoots", 2006; "New South Wales Government Department of Environment and Climate Change", 2008)
- Anti-predator Adaptations
Although peramelemorphs may help control insect pest populations, none are considered keystone species. As omnivores that also eat plants, they may play an important role as seed dispersers. Bilbies are semifossorial and may help aerate soil by burrowing. (Department of Primary Industries and Water, 2008; IUCN, 2008)
Peramelemorphs are host to a number of endoparasites including an array of protists and roundworms. For example, renal coccidiosis and toxoplasmosis, which is transmitted by cats and is fatal to peramelemorphs, are both caused by parasitic protozoa, and toxoplasmosis is considered a major threat to the persistence of eastern barred bandicoots by the International Union for Conservation of Nature. Various members of Peramelemorphia are also vulnerable to a number of ectoparasitic arthropods including mites, ticks, and fleas. Their solitary nature, however, may help reduce intraspecific transmission of parasites. (Bennett, et al., 2006; Bennett, et al., 2007; Bennett, 2008; Breed, et al., 2009; CDC, 2008; Department of Primary Industries and Water, 2008; IUCN, 2008)
- Ecosystem Impact
- soil aeration
Economic Importance for Humans: Positive
Members of Peramelemorphia have little direct economic importance, however, the giant bandicoot is still hunted by natives for its high market value in the bushmeat trade and for its fur. It is estimated that the species sells for the equivalent of 6 U.S. dollars. ("Bandicoots and Bilbies: Peramelemorphia", 2009; IUCN, 2008; "Peramelemorphia", 2005)
- Positive Impacts
- body parts are source of valuable material
Economic Importance for Humans: Negative
Due to their rapid decline, peramelemorphs have little to no effect on humans. Bandicoots and bilbies have been found to venture into suburban areas digging up lawns and gardens in search of food. As a result, they are sometimes viewed as pests. Peramelemorphs carry ticks, mites, and fleas, which can be transmitted to domestic animals and humans. Some species carry diseases, such as coccidiosis and toxoplasmosis, that can also be transmitted to domestic animals and humans. ("Bandicoots and Bilbies: Peramelemorphia", 2009; Bennett, 2008; Department of Primary Industries and Water, 2008; IUCN, 2008; "Peramelemorphia", 2005)
- Negative Impacts
- injures humans
- crop pest
There are a total of 22 recognized species within Peramelemorphia. According to the IUCN’s Red List of Threatened Species, 3 have recently gone extinct, 4 are classified as endangered, 2 are vulnerable, 1 is near threatened, 9 are of least concern, and the remaining 3 are data deficient. Species adapted to arid and semi-arid habitats have experienced significant declines since European settlement. A major threat to peramelemorphs is the changing of fire regimens for agriculture and grazing animals across Australia and New Guinea. As fire regimens change and grazing increases, ground cover becomes reduced and predation increases. In addition, the introduction of non-native species into Australia and New Guinea (e.g., cats, red foxes, dogs and dingos) has resulted in increased predation and resource competition. By occupying a wide variety of habitats and vegetation types, bandicoots and bilbies largely avoid competition. However, the introduction of sheep, cattle, and European rabbits has caused many species to become threatened, and in some cases extinct, due to increased competition for resources. Some introduced species carry diseases that can be transmitted peramelemorphs. For example, toxoplasmosis, which is fatal to many species of bandicoot, was first introduced by cats. The giant bandicoot, which is endangered, is hunted and sold as food by local people. ("Bandicoots and Bilbies: Peramelemorphia", 2009; CDC, 2008; Department of Primary Industries and Water, 2008; IUCN, 2008; "Peramelemorphia", 2005)
Recovery plans for some species within Peramelemorphia have been implemented. These plans include protecting wild populations and their habitats, maintaining captive populations, intensive control of predators, and enhancing community education and participation in local conservation efforts. In some cases, the reintroduction of of locally extinct species has been successful (e.g., eastern barred bandicoot). However, many species and the habitats they depend upon remain vulnerable and unprotected. Some species have insufficient habitat for population growth due to intrafamilial range overlap and inbreeding. Small population sizes increase a species risk of extinction caused by predation and hunting. ("Bandicoots and Bilbies: Peramelemorphia", 2009; CDC, 2008; Department of Primary Industries and Water, 2008; IUCN, 2008; "Peramelemorphia", 2005)
- IUCN Red List [Link]
- Not Evaluated
Kathryn Frens (author), University of Michigan-Ann Arbor, Phil Myers (editor), University of Michigan-Ann Arbor, John Berini (editor), Animal Diversity Web Staff.
Living in Australia, New Zealand, Tasmania, New Guinea and associated islands.
uses sound to communicate
living in landscapes dominated by human agriculture.
young are born in a relatively underdeveloped state; they are unable to feed or care for themselves or locomote independently for a period of time after birth/hatching. In birds, naked and helpless after hatching.
- bilateral symmetry
having body symmetry such that the animal can be divided in one plane into two mirror-image halves. Animals with bilateral symmetry have dorsal and ventral sides, as well as anterior and posterior ends. Synapomorphy of the Bilateria.
an animal that mainly eats meat
- causes disease in humans
an animal which directly causes disease in humans. For example, diseases caused by infection of filarial nematodes (elephantiasis and river blindness).
Found in coastal areas between 30 and 40 degrees latitude, in areas with a Mediterranean climate. Vegetation is dominated by stands of dense, spiny shrubs with tough (hard or waxy) evergreen leaves. May be maintained by periodic fire. In South America it includes the scrub ecotone between forest and paramo.
uses smells or other chemicals to communicate
active at dawn and dusk
having markings, coloration, shapes, or other features that cause an animal to be camouflaged in its natural environment; being difficult to see or otherwise detect.
- desert or dunes
in deserts low (less than 30 cm per year) and unpredictable rainfall results in landscapes dominated by plants and animals adapted to aridity. Vegetation is typically sparse, though spectacular blooms may occur following rain. Deserts can be cold or warm and daily temperates typically fluctuate. In dune areas vegetation is also sparse and conditions are dry. This is because sand does not hold water well so little is available to plants. In dunes near seas and oceans this is compounded by the influence of salt in the air and soil. Salt limits the ability of plants to take up water through their roots.
animals that use metabolically generated heat to regulate body temperature independently of ambient temperature. Endothermy is a synapomorphy of the Mammalia, although it may have arisen in a (now extinct) synapsid ancestor; the fossil record does not distinguish these possibilities. Convergent in birds.
- female parental care
parental care is carried out by females
an animal that mainly eats leaves.
A substance that provides both nutrients and energy to a living thing.
forest biomes are dominated by trees, otherwise forest biomes can vary widely in amount of precipitation and seasonality.
Referring to a burrowing life-style or behavior, specialized for digging or burrowing.
An animal that eats mainly plants or parts of plants.
An animal that eats mainly insects or spiders.
- island endemic
animals that live only on an island or set of islands.
offspring are produced in more than one group (litters, clutches, etc.) and across multiple seasons (or other periods hospitable to reproduction). Iteroparous animals must, by definition, survive over multiple seasons (or periodic condition changes).
having the capacity to move from one place to another.
This terrestrial biome includes summits of high mountains, either without vegetation or covered by low, tundra-like vegetation.
- native range
the area in which the animal is naturally found, the region in which it is endemic.
active during the night
an animal that mainly eats all kinds of things, including plants and animals
chemicals released into air or water that are detected by and responded to by other animals of the same species
the kind of polygamy in which a female pairs with several males, each of which also pairs with several different females.
having more than one female as a mate at one time
rainforests, both temperate and tropical, are dominated by trees often forming a closed canopy with little light reaching the ground. Epiphytes and climbing plants are also abundant. Precipitation is typically not limiting, but may be somewhat seasonal.
- scrub forest
scrub forests develop in areas that experience dry seasons.
- seasonal breeding
breeding is confined to a particular season
reproduction that includes combining the genetic contribution of two individuals, a male and a female
- soil aeration
digs and breaks up soil so air and water can get in
living in residential areas on the outskirts of large cities or towns.
uses touch to communicate
that region of the Earth between 23.5 degrees North and 60 degrees North (between the Tropic of Cancer and the Arctic Circle) and between 23.5 degrees South and 60 degrees South (between the Tropic of Capricorn and the Antarctic Circle).
Living on the ground.
defends an area within the home range, occupied by a single animals or group of animals of the same species and held through overt defense, display, or advertisement
the region of the earth that surrounds the equator, from 23.5 degrees north to 23.5 degrees south.
- tropical savanna and grassland
A terrestrial biome. Savannas are grasslands with scattered individual trees that do not form a closed canopy. Extensive savannas are found in parts of subtropical and tropical Africa and South America, and in Australia.
A grassland with scattered trees or scattered clumps of trees, a type of community intermediate between grassland and forest. See also Tropical savanna and grassland biome.
- temperate grassland
A terrestrial biome found in temperate latitudes (>23.5° N or S latitude). Vegetation is made up mostly of grasses, the height and species diversity of which depend largely on the amount of moisture available. Fire and grazing are important in the long-term maintenance of grasslands.
uses sight to communicate
reproduction in which fertilization and development take place within the female body and the developing embryo derives nourishment from the female.
- year-round breeding
breeding takes place throughout the year
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