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The mountains of Glacier National Park began forming 170 million years ago when ancient rocks were forced eastward up and over much younger rock strata. Known as the Lewis Overthrust, these sedimentary rocks are considered to have some of the finest examples of early life fossils on Earth. The current shapes of the Lewis and Livingston mountain ranges and positioning and size of the lakes show the telltale evidence of massive glacial action, which carved U-shaped valleys and left behind moraines which impounded water, creating lakes. Of the estimated 150 glaciers which existed in the park in the mid-19th century, only 25 active glaciers remained by 2010. Scientists studying the glaciers in the park have estimated that all the active glaciers may disappear by 2030 if current climate patterns persist.Glacier National Park has almost all its original native plant and animal species. Large mammals such as grizzly bears, moose, and mountain goats, as well as rare or endangered species like wolverines and Canadian lynxes, inhabit the park. Hundreds of species of birds, more than a dozen fish species, and a few reptile and amphibian species have been documented. The park has numerous ecosystems ranging from prairie to tundra. The easternmost forests of western redcedar and hemlock grow in the southwest portion of the park. Large forest fires are unusual in the park; however, more than 13% of the park burned in 2003.Glacier National Park borders Waterton Lakes National Park in Canada—the two parks are known as the Waterton-Glacier International Peace Park and were designated as the world's first International Peace Park in 1932. Both parks were designated by the United Nations as Biosphere Reserves in 1976, and in 1995 as World Heritage sites.  In April 2017, the joint park received a provisional Gold Tier designation as Waterton-Glacier International Dark Sky Park through the International Dark Sky Association, the first transboundary dark sky park.

Waterton-Glacier International Peace Park