Answer the following question given this paragraph: 

All that changed when plants moved from water to land. This may have happened by 500 million years ago or even earlier. On land, everything was wide open. There were no other living things. Without plants, there was nothing for other organisms to eat. Land could not be colonized by other organisms until land plants became established. The earliest land plants may have resembled the modern liverworts in Figure below .


Q: Land could not be colonized by other organisms until what became established?


A:
plants